Probolomyrmex filiformis, copyright April Nobile.

Belongs within: Formicidae.

The Proceratiinae are a group of small ants characterised by fully or nearly fully exposed antennal sockets close to the anterior margin of the head, together with a single petiolar segment and a fused promesonotal suture (Heterick 2009).

    |--Probolomyrmex Mayr 1901 TB85 [Probolomyrmecini H09]
    |    |  i. s.: *P. filiformis Mayr 1901 TB85
    |    |         P. bidens SGH12
    |    |         P. maryatiae SGH12
    |    |         P. okinawensis Terayama & Ogata 1988 I92
    |    |         P. tani RBV08
    |    |         P. vieti SGH12
    |    |--P. greavesi group SGH12
    |    |    |--P. greavesi Taylor 1965 TB85
    |    |    |--P. latalongus Shattuck, Gunawardene & Heterick 2012 SGH12
    |    |    `--P. salomonis SGH12
    |    `--P. longinodus group SGH12
    |         |--P. aliundus Shattuck, Gunawardene & Heterick 2012 SGH12
    |         |--P. longinodus Terayama & Ogata 1988 I92
    |         |--P. newguinensis Shattuck, Gunawardene & Heterick 2012 SGH12
    |         `--P. simplex Shattuck, Gunawardene & Heterick 2012 SGH12
    `--Proceratiini H09
         |--Proceratium Roger 1863 TB85
         |    |--*P. silaceum Roger 1863 TB85
         |    |--P. californicum WW90
         |    |--P. papuanum Emery 1897 TB85
         |    `--P. stictum Brown 1958 TB85
         `--Discothyrea Roger 1863 H09, KT99 (see below for synonymy)
              |--*D. testacea Roger 1863 KT99
              |--D. bidens Clark 1928 TB85
              |--D. crassicornis Clark 1926 TB85
              |--D. kamiteta Kubota & Terayama 1999 KT99
              |--D. leae Clark 1934 TB85
              |--*Pseudosysphincta’ poweri Arnold 1916 KT99
              |--D. sauteri KT99
              |--D. turtoni Clark 1934 TB85
              `--D. velutina (Wheeler 1916) TB85 [=*Prodiscothyrea velutina KT99]

Discothyrea Roger 1863 H09, KT99 [incl. Prodiscothyrea Wheeler 1916 KT99, Pseudosysphincta Arnold 1916 KT99, Pseudosphincta (l. c.) KT99]

*Type species of generic name indicated


[H09] Heterick, B. E. 2009. A guide to the ants of south-western Australia. Records of the Western Australian Museum Supplement 76: 1–206.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[KT99] Kubota, M. & M. Terayama. 1999. A description of a new species of the genus Discothyrea Roger from the Ryukyus, Japan (Hymenoptera: Formicidae). Memoirs of the Myrmecological Society of Japan 1: 1–5.

[RBV08] Rabeling, C., J. M. Brown & M. Verhaagh. 2008. Newly discovered sister lineage sheds light on early ant evolution. Proceedings of the National Academy of Sciences of the USA 105 (39): 14913–14917.

[SGH12] Shattuck, S. O., N. R. Gunawardene & B. Heterick. 2012. A revision of the ant genus Probolomyrmex (Hymenoptera: Formicidae: Proceratiinae) in Australia and Melanesia. Zootaxa 3444: 40–50.

[TB85] Taylor, R. W., & D. R. Brown. 1985. Formicoidea. In: Walton, D. W. (ed.) Zoological Catalogue of Australia vol. 2. Hymenoptera: Formicoidea, Vespoidea and Sphecoidea pp. 1–149. Australian Government Publishing Service: Canberra.

[WW90] Wheeler, G. C., & J. Wheeler. 1990. Insecta: Hymenoptera Formicidae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1277–1294. John Wiley & Sones: New York.

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