Brown rat Rattus norvegicus, photographed by Warwick Sloss.

Belongs within: Muridae.

Rattus, the true rats, is a genus of medium-sized rodents native to southern and eastern Asia and Australasia. Two species, the black rat Rattus rattus and the brown rat Rattus norvegicus, have dispersed worldwide in association with humans.

    |--R. adustus IT07
    |--R. annandalei (Bonhote 1903) K92
    |--R. argentiventer (Robinson & Kloss 1916) K92
    |--R. baluensis IT07
    |--R. bontanus IT07
    |--R. burrus IT07
    |--R. callitrichus BP87
    |--R. colletti IT07
    |--R. culionensis BP87
    |--R. elaphinus AC98
    |--R. enganus IT07
    |--R. everetti [incl. R. albigularis, R. gala, R. tyrannus] M82
    |--R. exulans (Peale 1848) K92 [=Mus exulans R64, Epimys exulans R14]
    |    |--R. e. exulans R14
    |    `--‘Mus’ e. maorium R14
    |--R. feliceus IT07
    |--R. foramineus IT07
    |--R. fuscipes S83
    |    |--R. f. fuscipes S83
    |    |--R. f. coracius S83
    |    |--R. f. greyii S83
    |    |--R. ‘greyi’ peccatus Troughton 1937 [=R. g. ravus Brazenor 1936 non Robison & Kloss 1916] R64
    |    `--R. ‘greyi’ pelori Finlayson 1960 R64
    |--R. giluwensis IT07
    |--R. hainaldi IT07
    |--R. hoffmanni (Matchie 1901) K92
    |--R. hoogerwerfi IT07
    |--R. jobiensis IT07
    |--R. koopmani IT07
    |--R. korinchi IT07
    |--R. lacus Tate 1951 R64
    |--R. latidens BP87
    |--R. leucopus R64
    |    |--R. l. leucopus R64
    |    |--R. l. cooktownensis Tate 1951 R64
    |    `--R. l. mcilwraithi Tate 1951 R64
    |--R. losea IT07
    |--R. lugens IT07
    |--R. lutreolus R64
    |    |--R. l. lutreolus R64
    |    |--R. l. cambricus Troughton 1937 R64
    |    `--R. l. imbil Troughton 1937 R64
    |--R. macleari BP87
    |--R. marmosurus IT07
    |--R. mindorensis IT07
    |--R. mollicomulus IT07
    |--R. montanus IT07
    |--R. mordax IT07
    |--R. morotaiensis IT07
    |--R. nativitatis BP87
    |--R. niobe (Thomas 1906) K92
    |--R. nitidus IT07
    |--R. norvegicus (Berkenhout 1769) [=Mus norvegicus] B75
    |--R. novaeguineae IT07
    |--R. omichlodes BP87
    |--R. osgoodi Musser & Newcomb 1985 FN97
    |--R. owiensis BP87
    |--R. palmarum IT07
    |--R. pelurus IT07
    |--R. praetor IT07
    |--R. pulliventer BP87
    |--R. ranjiniae IT07
    |--R. rattus (Linnaeus 1758) K92
    |    |--R. r. rattus B75
    |    |--R. r. alexandrinus (Geoffroy St.-Hilaire 1803) B75
    |    `--R. r. frugivorus (Rafinesque-Schmaltz 1814) KK54
    |--R. remotus BP87
    |--R. rogersi BP87
    |--R. sanila IT07
    |--R. sikkimensis IT07
    |--R. simalurensis IT07
    |--R. sordidus IT07
    |--R. steini IT07
    |--R. stoicus IT07
    |--R. tanezumi IT07
    |--R. tawitawiensis IT07
    |--R. timorensis IT07
    |--R. tiomanicus (Miller 1900) K92
    |--R. tunneyi Thomas 1904 [incl. R. culmorum] K92
    |    |--R. t. tunneyi R64
    |    |--R. ‘culmorum’ apex Troughton 1939 R64
    |    `--R. t. dispar Brazenor 1936 R64
    |--R. turkestanicus IT07
    |--R. villosissimus IT07
    `--R. xanthurus IT07

*Type species of generic name indicated


[AC98] Alcover, J. A., X. Campillo, M. Macias & A. Sans. 1998. Mammal species of the world: additional data on insular mammals. American Museum Novitates 3248: 1-29.

[B75] Bowles, J. B. 1975. Distribution and biogeography of mammals of Iowa. Special Publications, The Museum, Texas Tech University 9: 1-184.

[BP87] Burton, J. A., & B. Pearson. 1987. Collins Guide to the Rare Mammals of the World. Collins: London.

[FN97] Feiler, A., & T. Nadler. 1997. Recently discovered mammals in Vietnam—present results on taxonomy, zoogeography, status and on the protection of the animals (Mammalia). Zoologische Abhandlungen 49 (21): 331-335.

[IT07] Isaac, N. J. B., S. T. Turvey, B. Collen, C. Waterman & J. E. M. Baillie. 2007. Mammals on the EDGE: conservation priorities based on threat and phylogeny. PloS One 2 (3): e296.

[KK54] Klemmer, K., & H. E. Krampitz. 1954. Zur Kenntnis der Säugetierfauna Siziliens. Senckenbergiana Biologica 35 (3-4): 121-135.

[K92] Klompen, J. S. H. 1992. Phylogenetic relationships in the mite family Sarcoptidae (Acari: Astigmata). Miscellaneous Publications, Museum of Zoology, University of Michigan 180: i-vi, 1-154.

[M82] Musser, G. G. 1982. Results of the Archbold Expeditions. No. 110. Crunomys and the small-bodied shrew rats native to the Philippine Islands and Sulawesi (Celebes). Bulletin of the American Museum of Natural History 174 (1): 1-95.

[R14] Revilliod, P. 1914. Les mammifères de la Nouvell-Calédonie et des îles Loyalty. In Nova Caledonia: Forschungen in Neu-Caledonian und auf den Loyalty-Inseln. A. Zoologie vol. 1, Heft IV (F. Sarasin & J. Roux, eds) pp. 341-365, pl. 10. C. W. Kreidels Verlag: Wiesbaden.

[R64] Ride, W. D. L. 1964. A list of mammals described from Australia between the years 1933 and 1963 (comprising newly proposed names and additions to the Australian faunal list). Australian Mammal Society Bulletin 7 (Suppl.): 1-15.

[S83] Smith, A. M. A. 1983. The subspecific biochemical taxonomy of Antechinus minimus, A. swainsonii and Sminthopsis leucopus (Marsupialia: Dasyuridae). Australian Journal of Zoology 32: 753-762.

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