Kalotermes rufinotum, from CSIRO Entomology.

Belongs within: Isoptera.

The Kalotermitidae includes the drywood termites, a pantropical termite family lacking a division between the working and reproductive castes (Grimaldi & Engel 2005).

Characters (from Hill 1942): Tarsi four-segmented; fontanelle absent. Winged adult with ocelli; mandibles of Heterotermes type; pronotum usually wider than head; anterior wing stumps usually large. Soldier with rudimentary eyes; mandibles very strong; sutures of head usually present.

<==Kalotermitidae GE05
    |--Incisitermes minor TW05
    |--Paraneotermes simplicicornis N90
    |--Neotermes Holmgren 1911 H42
    |    |--N. castaneus Burmeister 1839 BP01
    |    |--N. connexus [=Calotermes connexus] H42
    |    |--N. ferrugineus (Holmgren 1911) [=Calotermes (Neotermes) ferrugineus] H42
    |    |--N. insularis (White in Walker 1853) (see below for synonymy) H42
    |    |--N. papua (Desneux 1905) [=Calotermes (Neotermes) papua, Kalotermes (N.) papua] H42
    |    |--N. rainbowi (Hill 1926) [=Calotermes (Neotermes) rainbowi, Halotermes (N.) rainbowi] H42
    |    |--N. rouxi (Holmgren 1915) [=Calotermes (Neotermes) rouxi] H42
    |    |--N. samoanus (Holmgren 1912) (see below for synonymy) H42
    |    |--N. sarasini (Holmgren 1915) [=Calotermes (Neotermes) sarasini] H42
    |    |--N. schultzei (Holmgren 1911) [=Calotermes (Neotermes) schultzei] H42
    |    |--N. semilunaris (Holmgren 1915) [=Calotermes (Neotermes) semilunaris] H42
    |    |--N. sjoestedti (Desneux 1907) [=Calotermes sjoestedti, C. (Neotermes) sjoestedti] H42
    |    `--N. wagneri BB03
    |--Cryptotermes Banks 1906 TW05, H42 (see below for synonymy)
    |    |--C. albipes (Holmgren 1915) [=Calotermes (Cryptotermes) albipes] H42
    |    |--C. arcanus (Hill 1925) [=Calotermes (Cryptotermes) arcanus] H42
    |    |--C. atratus (Hill 1933) [=Calotermes (Cryptotermes) atratus] H42
    |    |--C. brevis [=Kalotermes (Cryptotermes) brevis] H42
    |    |--‘Calotermes’ campbelli Light 1924 H42
    |    |--C. canalensis (Holmgren 1915) [=Calotermes (Procryptotermes) canalensis, Ca. (Cryptotermes) canalensis] H42
    |    |--C. cynocephalus Light 1921 [=Calotermes (Cryptotermes) cynocephalus] H42
    |    |--‘Calotermes’ domesticus Haviland 1898 H42
    |    |--C. gearyi (Hill 1942) [=Calotermes (Cryptotermes) gearyi] H42
    |    |--C. hermsi Kirby 1925 (see below for synonymy) H42
    |    |--‘Calotermes’ kotoensis Oshima 1912 H42
    |    |--C. nocens (Light 1921) (see below for synonymy) H42
    |    |--C. primus (Hill 1921) [=Calotermes (Cryptotermes) primus] H42
    |    |--C. queenslandis (Hill 1933) [=Calotermes (Cryptotermes) queenslandis] H42
    |    `--C. speiseri (Holmgren 1915) [=Calotermes (Procryptotermes) speiseri] H42
    `--Kalotermes Hagen 1853 GE05, H42 [=Calotermes Hagen 1858 H42]
         |  i. s.: K. banksiae Hill 1942 H74, H42 [=Calotermes banksiae H42]
         |         K. brouni Froggatt 1896 M59, H42 [=Calotermes brouni H42]
         |         ‘Termes’ convexus Walker 1853 [=Calotermes convexus] H42
         |         K. flavicollis Fabricius 1793 BP01
         |         ‘Calotermes’ improbus Hagen 1858 H42
         |         K. minor M59
         |         ‘Termes’ obscurus Walker 1853 [=Calotermes (Glyptotermes) obscurus] H42
         |         ‘Calotermes’ pallidinotum Hill 1942 H42
         |         K. sinaica BB03
         |         ‘Calotermes’ spoliator Hill 1932 H42
         |         K. swinhoei GE05
         |         ‘Calotermes’ tillyardi Hill 1932 H42
         |--K. (Kalotermes) H42
         |    |--‘Calotermes (Calotermes)’ condonensis Hill 1922 (see below for synonymy) H42
         |    |--‘Calotermes (Calotermes)’ immigrans H42
         |    |--‘Calotermes (Calotermes)’ repandus Hill 1926 H42
         |    `--‘Calotermes (Calotermes)’ rufinotum Hill 1925 H42
         `--K. (Glyptotermes Froggatt 1896) H42
              |--‘Calotermes’ (G.) brevicornis (Froggatt 1896) (see below for synonymy) H42
              |--‘Calotermes’ (G.) eucalypti (Froggatt 1896) [=Glyptotermes eucalypti] H42
              |--‘Calotermes’ (G.) iridipennis (Froggatt 1896) [=Glyptotermes iridipennis] H42
              |--‘Calotermes’ (G.) neotuberculatus Hill 1933 H42
              |--‘Calotermes’ (G.) taveuniensis Hill 1926 H42
              |--‘Calotermes’ (G.) tuberculatus (Froggatt 1896) [=Glyptotermes tuberculatus] H42
              `--K. (G.) xantholabrum Hill 1926 [=Calotermes (G.) xantholabrum; incl. K. (G.) juddi Light 1932] H42

‘Calotermes (Calotermes)’ condonensis Hill 1922 [incl. C. (C.) oldfieldi Hill 1925, C. oldfieldii var. chryseus Hill 1926] H42

‘Calotermes’ (Glyptotermes) brevicornis (Froggatt 1896) [=Glyptotermes brevicornis; incl. C. (G.) affinis Mjöberg 1920, C. (G.) claripennis Hill 1925, C. (G.) dubius Mjöberg 1920, C. (G.) nigrolabrum Hill 1921, C. (G.) perangustus Hill 1926, C. (G.) trilineatus Mjöberg 1920] H42

Cryptotermes Banks 1906 TW05, H42 [incl. Allotermes Wasmann 1910 H42, Planocryptotermes Light 1921, Procryptotermes Holmgren 1910 H42]

Cryptotermes hermsi Kirby 1925 [=Kalotermes (Cryptotermes) hermsi; incl. Calotermes (Cr.) buxtoni Hill 1926, Ca. (Cr.) gulosus Hill 1927, Ca. (Cr.) repentinus Hill 1927] H42

Cryptotermes nocens (Light 1921) [=Planocryptotermes nocens, Calotermes (Cryptotermes) nocens; incl. Ca. (Cr.) secundus Hill 1925] H42

Neotermes insularis (White in Walker 1853) [=Termes insularis, Calotermes (Neotermes) insularis; incl. C. (Neotermes) deuqueti Hill 1926, C. irregularis Froggatt 1896, C. (Neotermes) irregularis, C. longiceps Froggatt 1896, C. (Neotermes) longiceps, C. malandensis Mjöberg 1920, C. (Neotermes) oculifer Mjöberg 1920, C. (Neotermes) paralleliceps Mjöberg 1920, C. robustus Froggatt 1896, C. (Neotermes) robustus] H42

Neotermes samoanus (Holmgren 1912) [=Calotermes (Neotermes) samoanus, Kalotermes (N.) samoanus; incl. N. sanctaecrucis Snyder 1925] H42

*Type species of generic name indicated


[BB03] Brugerolle, G., & C. Bordereau. 2003. Ultrastructure of Joenoides intermedia (Grassé 1952), a symbiotic parabasalid flagellate of Hodotermes mossambicus, and its comparison with other joeniid genera. European Journal of Protistology 39: 1-10.

[BP01] Brugerolle, G., & D. J. Patterson. 2001. Ultrastructure of Joenina pulchella Grassi, 1917 (Protista, Parabasalia), a reassessment of evolutionary trends in the parabasalids, and a new order Cristamonadida for devescovinid, calonymphid and lophomonad flagellates. Organisms Diversity and Evolution 1: 147-160.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[H74] Helson, G. A. H. 1974. Insect Pests: Identification, life history, and control of pests of farms, horticulture, gardens, and public health. A. R. Shearer, Government Printer: Wellington (New Zealand).

[H42] Hill, G. F. 1942. Termites (Isoptera) from the Australian Region (including Australia, New Guinea and islands south of the Equator between 140°E. longitude and 170°W. longitude). Commonwealth of Australia Council for Scientific and Industrial Research: Melbourne.

[M59] Morgan, F. D. 1959. The ecology and external morphology of Stolotermes ruficeps Brauer (Isoptera: Hodotermitidae). Transactions of the Royal Society of New Zealand 86 (1): 155-195.

[N90] Nutting, W. L. 1990. Insecta: Isoptera. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 997-1032. John Wiley & Sones: New York.

[TW05] Terry, M. D., & M. F. Whiting. 2005. Mantophasmatodea and phylogeny of the lower neopterous insects. Cladistics 21: 240-257.

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