Male great grig Cyphoderris monstrosa, photographed by James Miskelly.

Belongs within: Ensifera.
Contains: Apohaglida.

The Haglida are a clade within the Orthoptera ancestrally characterised by the presence in males of a 'fan' in the forewings, an expansion between the branches of the posterior cubital vein. With the exception of the basal Hagloptera intermedia, most have veins RP and MA1 distinct in the fore wings (Béthoux & Nel 2002).

Synapomorphies (from Béthoux & Nel 2002): Forewings with RP posteriorly ramified, with its branches distally ramified or RP simple; handle (pseudo-vein between origin of anastomosis CuA+CuPaα and CuPaβ) present; fan present (reversed in some derived taxa); branches of CuA + CuPaα strongly approximate (reversed in some derived taxa).

    |--Hagloptera intermedia Gorokhov 1986 BN02
    `--Metahaglida BN02
         |--Apohaglida BN02
         `--+--Isfaroptera [Isfaropterinae] BN02
            |    `--I. grylliformis Martynov 1937 BN02
            `--Haglidae GR02
                 |--Cyphoderris monstrosa TW05
                 |--Prohagla superba Riek 1954 K-P91, F71
                 |--Neohagla K-P91
                 |--Hagla BNL02
                 |    |--*H. gracilis BN02
                 |    `--H. contorta Riek 1974 BNL02
                 `--Lyrohagla BN02
                      |--L. pravdini Gorokhov 1986 BN02
                      `--L. uvarovi GR02

*Type species of generic name indicated


[BN02] Béthoux, O., & A. Nel. 2002. Venation pattern and revision of Orthoptera sensu nov. and sister groups. Phylogeny of Palaeozoic and Mesozoic Orthoptera sensu nov. Zootaxa 96: 1–88.

[BNL02] Béthoux, O., A. Nel, J. Lapeyrie, G. Gand & J. Galtier. 2002. Raphogla rubra gen. n., sp. n., the oldest representative of the clade of modern Ensifera (Orthoptera: Tettigoniidea, Gryllidea). European Journal of Entomology 99: 111–116.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[GR02] Gorochov, A. V., & A. P. Rasnitsyn. 2002. Superorder Gryllidea Laicharting, 1781 (=Orthopteroidea Handlirsch, 1903). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 293–303. Kluwer Academic Publishers: Dordrecht.

[K-P91] Kukalová-Peck, J. 1991. Fossil history and the evolution of hexapod structures. In: CSIRO. The Insects of Australia: A textbook for students and research workers vol. 1 pp. 141–179. Melbourne University Press: Carlton (Victoria).

[TW05] Terry, M. D., & M. F. Whiting. 2005. Mantophasmatodea and phylogeny of the lower neopterous insects. Cladistics 21: 240–257.

Last updated: 25 May 2022.

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