Syncarida

Allobathynella japonica, from Morimoto & Miura (1957).


Belongs within: Eumalacostraca.
Contains: Bathynella, Hexabathynella, Iberobathynella.

The Syncarida are a group of malacostracan crustaceans found primarily in fresh water that lack a carapace. Living examples are divided between the Anaspidacea, in which the first thoracic segment is fused to the head and a tail fan is present, and the minute Bathynellacea that lack a tail fan.

<==Syncarida S86
    |  i. s.: Allanaspides S86
    |--Palaeocaridacea MD01
    |    |--Clarkecaris [Clarkecarididae] G88
    |    |    `--C. brazilicus (Clarke 1920) S77
    |    |--Minicarididae BWW93
    |    |    |--Minicaris brandi Schram 1979 BWW93
    |    |    `--Erythrogaulos carrizoensis Schram 1984 BWW93
    |    |--Squillitidae BWW93
    |    |    |--Squillites spinosa Scott 1938 BWW93
    |    |    `--Nectotelson krejcii (Fritsch 1875) BWW93
    |    |--Palaeocaris [Palaeocarididae] G88
    |    |    |--P. canadensis G88
    |    |    |--P. retractata Calman 1932 BWW93
    |    |    |--P. secretanae Schram 1984 BWW93
    |    |    `--P. typus G88
    |    `--Acanthotelsonidae BWW93
    |         |--Palaeosyncaris dakotensis Brooks 1962 BWW93
    |         |--Acanthotelson [incl. Eileticus] WA04
    |         |    |--‘Eileticus’ anthracinus Scudder 1882 WA04
    |         |    `--A. stimpsoni G88
    |         `--Uronectes BWW93
    |              |--U. fimbriatus (Jordan 1947) BWW93
    |              `--U. kinniensis G88
    |--Anaspidacea [Anaspidinea] LSE13
    |    |--Anaspididae BWW93
    |    |    |--Anaspidites antiquus (Chilton 1929) BWW93
    |    |    |--Anaspides Thomson 1894
    |    |    |    `--A. tasmaniae (Thomson 1893)
    |    |    `--Koonaspides indistinctus G88
    |    |--Koonungidae S86
    |    |    |--Koonunga N31
    |    |    `--Micraspides Nicholls 1931 N31
    |    |         `--*M. calmani Nicholls 1931 N31
    |    |--Psammaspididae S86
    |    |    |--Psammaspides Schminke 1974 S86
    |    |    |    `--P. williamsi S86
    |    |    `--Eucrenonaspides Knott & Lake 1980 S86
    |    |         `--E. oinotheke Knott & Lake 1980 S86
    |    `--Stygocarididae [Stygocaridacea] S86
    |         |--Oncostygocaris Schminke 1980 S86
    |         |    `--O. patagonica (Noodt 1963) S86
    |         |--Stygocarella Schminke 1980 S86
    |         |    `--S. pleotelson Schminke 1980 S86
    |         |--Parastygocaris Noodt 1963 S86
    |         |    |--P. andina Noodt 1963 S86
    |         |    `--P. goerssi Noodt 1963 S86
    |         `--Stygocaris Noodt 1963 S86
    |              |--S. giselae Schminke 1980 S86
    |              |--S. gomezmillasi Noodt 1963 S86
    |              `--S. townsendi Morimoto 1977 S86
    `--Bathynellacea LSE13
         |--Bathynellidae S86
         |    |--Bathynella S86
         |    |--Sardobathynella Serban 1973 S86
         |    |    `--S. cottarelli Serban 1973 S86
         |    |--Vandelibathynella Serban, Coineau & Delamare Deboutteville 1971 S86
         |    |    `--V. vandeli (Delamare Deboutteville & Chappuis 1954) S86
         |    |--Agnathobathynella Schminke 1980 S86
         |    |    `--A. ecclesi Schminke 1980 S86
         |    |--Austrobathynella Delamare Deboutteville 1960 S86
         |    |    `--A. patagonica Delamare Deboutteville 1960 S86
         |    |--Transvaalthynella Serban & Coineau 1975 S86
         |    |    `--T. coineaui Serban & Coineau 1975 S86
         |    |--Transkeithynella Serban & Coineau 1975 S86
         |    |    `--T. paradoxa Serban & Coineau 1975 S86
         |    |--Parameridiobathynella Serban & Leclerc 1984 S86
         |    |    `--P. gardensis Serban & Leclerc 1984 S86
         |    |--Pseudobathynella Serban, Coineau & Delamare Deboutteville 1971 S86
         |    |    |--P. helvetica Serban & Delamare Deboutteville 1974 S86
         |    |    `--P. magniezi Serban, Coineau & Delamare Deboutteville 1971 S86
         |    |--Meridiobathynella Serban, Coineau & Delamare Deboutteville 1971 S86
         |    |    |--M. catalanensis Serban, Coineau & Delamare Deboutteville 1971 S86
         |    |    `--M. rouchi Serban, Coineau & Delamare Deboutteville 1971 S86
         |    |--Vejdovskybathynella Serban & Leclerc 1984 S86
         |    |    |--V. balazuci Serban & Leclerc 1984 S86
         |    |    `--V. espattyensis Serban & Leclerc 1984 S86
         |    |--Antrobathynella Serban 1966 S86
         |    |    `--A. stammeri (Jakobi 1954) S86
         |    |         |--A. s. stammeri S86
         |    |         `--A. s. ciscaucasia (Birstein & Ljovuschkin 1964) S86
         |    |--Nannobathynella Noodt 1969 S86
         |    |    |--N. africana Schminke & Wells 1974 S86
         |    |    |--N. eburnea Schminke 1979 S86
         |    |    `--N. marcusi Noodt 1969 S86
         |    `--Gallobathynella Serban, Coineau & Delamare Deboutteville 1971 S86
         |         |--G. boui Serban, Coineau & Delamare Deboutteville 1971 S86
         |         |--G. coiffaiti (Delamare Deboutteville 1961) S86
         |         |--G. delayi Serban, Coineau & Delamare Deboutteville 1971 S86
         |         |--G. juberthiae Serban, Coineau & Delamare Deboutteville 1971 S86
         |         `--G. tarissei Serban, Coineau & Delamare Deboutteville 1971 S86
         `--Parabathynellidae S86
              |--Lamtobathynella pentodonta S86
              |--Hexabathynella S86
              |--Iberobathynella S86
              |--Batubathynella Schminke 1973 S86
              |    `--B. malaya (Sars 1929) S86
              |--Nipponbathynella Schminke 1973 S86
              |    `--N. miurai (Ueno 1952) S86
              |--Ctenophallonella Coineau & Serban 1978 S86
              |    `--C. mutlumuviensis Coineau & Serban 1978 S86
              |--Haplophallonella Serban & Coineau 1975 S86
              |    `--H. heterodonta Serban & Coineau 1975 S86
              |--Heterodontobathynella Schminke 1973 S86
              |    `--H. ninianae (Green 1964) S86
              |--Brasilibathynella Jakobi 1958 S86
              |    `--B. florianopolis Jakobi 1958 S86
              |--Odontobathynella Delamare Deboutteville & Serban 1979 S86
              |    `--O. amazonica Delamare Deboutteville & Serban 1979 S86
              |--Acanthobathynella Coineau 1967 S86
              |    `--A. knoepffleri Coineau 1967 S86
              |--Afrobathynella Schminke 1976 S86
              |    `--A. trimera Schminke 1976 S86
              |--Nunubathynella Schminke 1976 S86
              |    `--N. dimera Schminke 1976 S86
              |--Psalidobathynella Schminke 1979 S86
              |    `--P. stocki Schminke 1979 S86
              |--Nilobathynella Dumont 1984 S86
              |    `--N. predynastica Dumont 1984 S86
              |--Thermobathynella Capart 1951 S86
              |    |--T. adami Capart 1951 S86
              |    `--T. amyxi Siewing 1956 S86
              |--Habrobathynella Schminke 1973 S86
              |    |--H. jeanneli (Delamare Deboutteville & Paulian 1954) S86
              |    `--H. milloti (Delamare Deboutteville & Paulian 1954) S86
              |--Chilibathynella Noodt 1963 S86
              |    |--C. australiensis Schminke 1973 S86
              |    `--C. clandestina Noodt 1963 S86
              |--Noodtibathynella Schminke 1973 S86
              |    |--N. jumboli (Siewing 1958) S86
              |    `--N. neotropica (Noodt 1965) S86
              |--Parabathynella Chappuis 1926 S86
              |    |--P. malaya N31
              |    |--P. motasi Dancau & Serban 1963 S86
              |    `--P. stygia Chappuis 1926 [incl. P. phreatica Chappuis 1939] S86
              |--Parvulobathynella Schminke 1973 S86
              |    |--P. camposicola (Jakobi 1961) S86
              |    |--P. riegelorum (Noodt 1965) S86
              |    `--P. ypacaraiensis (Noodt 1963) S86
              |--Leptobathynella Noodt 1965 [Leptobathynellidae] S86
              |    |--L. camposicola (Jakobi 1961) S86
              |    `--L. richerti (Noodt 1963) S86
              |         |--L. r. richerti S86
              |         `--L. r. brasiliensis Noodt 1972 S86
              |--Atopobathynella Schminke 1973 S86
              |    |--A. chelifera Schminke 1973 S86
              |    |--A. compagana Schminke 1973 S86
              |    |--A. hospitalis Schminke 1973 S86
              |    `--A. valdiviana (Noodt 1965) S86
              |--Eobathynella Birstein & Ljovuschkin 1964 S86
              |    |--E. gracillima (Ueno 1956) S86
              |    |--E. matuta (Morimoto 1970) S86
              |    |--E. mesasiatica (Birstein & Ljovuschkin 1964) S86
              |    |--E. minima Jankowskaja 1972 S86
              |    `--E. tianshanica (Jankowskaja 1964) S86
              |--Notobathynella Schminke 1973 S86
              |    |--N. chiltoni Schminke 1973 S86
              |    |--N. hineoneae Schminke 1973 S86
              |    |--N. longipes Schminke 1978 S86
              |    |--N. remota Schminke 1973 S86
              |    `--N. williamsi Schminke 1973 S86
              |--Cteniobathynella Schminke 1973 S86
              |    |--C. bakeri (Green 1964) S86
              |    |--C. calmani (Por 1968) S86
              |    |--C. caparti (Fryer 1957) S86
              |    |--C. essameuri Dumont 1981 S86
              |    |--C. leleupi (Delamare Deboutteville & Chappuis 1955) S86
              |    |--C. noodti Schminke 1973 S86
              |    `--C. teocchii (Coineau & Knoepffler 1971) S86
              `--Allobathynella Morimoto & Miura 1957 S86
                   |--A. carinata (Ueno 1952) S86
                   |--A. coreana Morimoto 1970 S86
                   |--A. gigantea (Morimoto 1959) S86
                   |    |--A. g. gigantea S86
                   |    `--A. g. pluto (Morimoto 1963) S86
                   |--A. japonica Morimoto & Miura 1957 S86
                   |--A. kuma (Ueno 1956) S86
                   |--A. mirabilis Ueno 1961 S86
                   `--A. yaye (Ueno 1956) S86

*Type species of generic name indicated

REFERENCES

[BWW93] Briggs, D. E. G., M. J. Weedon & M. A. Whyte. 1993. Arthropoda (Crustacea excluding Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 321–342. Chapman & Hall: London.

[G88] Gray, J. 1988. Evolution of the freshwater ecosystem: the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology 62: 1–214.

[LSE13] Legg, D. A., M. D. Sutton & G. D. Edgecombe. 2013. Arthropod fossil data increase congruence of morphological and molecular phylogenies. Nature Communications 4: 2485.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1–124.

[N31] Nicholls, G. E. 1931. Micraspides calmani, a new syncaridan from the west coast of Tasmania. Linnean Society's Journal—Zoology 37 (254): 473–488.

[S86] Schminke, H. K. 1986. Syncarida. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 389–404. E. J. Brill/Dr W. Backhuys: Leiden.

[S77] Schram, F. R. 1977. Paleozoogeography of Late Paleozoic and Triassic Malacostraca. Systematic Zoology 26 (4): 367–379.

[WA04] Wilson, H. M., & L. I. Anderson. 2004. Morphology and taxonomy of Paleozoic millipedes (Diplopoda: Chilognatha: Archipolypoda) from Scotland. Journal of Paleontology 78 (1): 169–184.

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