Scutellidae

Scutella subrotunda, copyright Parent Géry.

Belongs within: Clypeasteroida.

The Scutellidae are a group of sand dollars known from the Oligocene and Miocene (Durham 1966).

Characters (from Durham 1966): Medium-sized to large, flattened, internal supports well developed; petals closed, outer member of pore pair subdivided; interambulacra continuous, usually as wide as ambulacra at ambitus; primordial ambulacral and interambulacral plates about equal; four genital pores; periproct on oral surface; food grooves bifurcating close to peristome.

<==Scutellidae D66
    |--Parascutella Durham 1953 D66
    |    |--*P. leognanensis (Lambert 1903) [=Scutella leognanensis] M89
    |    `--P. bonali M89
    |--Parmulechinus Lambert 1910 [incl. Stenaster Lambert 1905 non Billings 1858] D66
    |    |--*P. agassizi (Oppenheim 1902) M89 (see below for synonymy)
    |    `--P. subtetragona M89
    `--Scutella Lamarck 1816 M89 [incl. Lambertiella Checchia-Rispoli 1917 D66]
         |--*S. subrotunda (Leske 1778) [=Echinodiscus subrotundus] M89
         `--S. vindobonensis D66
              |--S. v. vindobonensis D66
              `--S. v. secunda D66

*Parmulechinus agassizi (Oppenheim 1902) M89 [=Scutella agassizi M89; incl. *Stenaster labrei Lambert 1905 D66, Sc. striatula Agassiz 1841 non de Serres 1829 D66]

*Type species of generic name indicated

REFERENCES

[D66] Durham, J. D. 1966. Clypeasteroids. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt U. Echinodermata 3 vol. 2 pp. U450–U491. The Geological Society of America, Inc., and The University of Kansas Press.

[M89] Mooi, R. 1989. Living and fossil genera of the Clypeasteroida (Echinoidea: Echinodermata): an illustrated key and annotated checklist. Smithsonian Contributions to Zoology 488: 1–51.