Geomyidae

Variable pocket gopher Orthogeomys heterodus, copyright INBio.

Belongs within: Castorimorpha.
Contains: Thomomys, Geomys.

The Geomyidae, pocket gophers, are a group of burrowing rodents found in North and Central America. They share the presence of externally opening, fur-lined cheek pouches with the related kangaroo rats and pocket mice of the Heteromyidae, but are distinguished by their adaptions for a fossorial lifestyle (Hafner & Hafner 1983).

<==Geomyidae
    |  i. s.: ‘Sorex’ mexicanus Kerr 1792 HH83
    |         Dikkomys B74
    |         Heterogeomys hispidus G69
    |           |--H. h. hispidus G69
    |           |--H. h. isthmicus G69
    |           |--H. h. tehuantepecus G69
    |           `--H. h. torridus G69
    |         Gregorymys MJ11
    |--Entoptychus MJ11 [Entoptychinae HH83]
    |--Pleurolicus MJ11 [Pleurolicinae HH83]
    `--Geomyinae HH83
         |--Parapliosaccomys HH83
         `--+--Thomomys GE05
            `--+--Geomys GE05
               `--+--Zygogeomys trichopus GE05
                  |--Orthogeomys GE05
                  |    |  i. s.: O. cuniculus G69
                  |    |         O. dariensis IT07
                  |    |         O. (Orthogeomys) grandis HH83
                  |    |           |--O. g. grandis G69
                  |    |           |--O. g. alleni G69
                  |    |           |--O. g. alvarezi Schaldach 1966 G69
                  |    |           |--O. g. carbo G69
                  |    |           |--O. g. felipensis G69
                  |    |           |--O. g. nelsoni G69
                  |    |           `--O. g. scalops G69
                  |    |         O. lanius IT07
                  |    |         O. matagalpae IT07
                  |    |         O. thaeleri IT07
                  |    |--O. hispidus GE05
                  |    `--+--+--O. cavator GE05
                  |       |  `--O. underwoodi GE05
                  |       `--+--O. cherriei GE05
                  |          `--O. (Macrogeomys) heterodus GE05, HH83
                  `--+--Cratogeomys GE05
                     |    |--C. castanops GE05
                     |    `--C. merriami GE05
                     `--Pappogeomys GE05
                          |--P. alcorni IT07
                          |--P. bulleri GE05
                          |--P. castanops MB86
                          |    |--P. c. castanops MB86
                          |    |--P. c. goldmani (Merriam 1895) MB86
                          |    |--P. c. rubellus (Nelson & Goldman 1934) MB86
                          |    |--P. c. subnubilus (Nelson & Goldman 1934) MB86
                          |    `--P. c. surculus Russell 1968 MB86
                          |--P. fumosus IT07
                          |--P. gymnurus HH83
                          |--P. merriami IT07
                          |--P. neglectus IT07
                          |--P. tylorhinus IT07
                          `--P. zinseri IT07

*Type species of generic name indicated

REFERENCES

[B74] Bugge, J. 1974. The cephalic arterial system in insectivores, primates, rodents and lagomorphs, with special reference to the systematic classification. Acta Anatomica 87 (Suppl 62): 1–160.

[G69] Goodwin, G. G. 1969. Mammals from the State of Oaxaca, Mexico, in the American Museum of Natural History. Bulletin of the American Museum of Natural History 141 (1): 1–269, 40 pls.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[HH83] Hafner, J. C., & M. S. Hafner. 1983. Evolutionary relationships of heteromyid rodents. In Biology of Desert Rodents. Great Basin Naturalist Memoirs 7: 3–29.

[IT07] Isaac, N. J. B., S. T. Turvey, B. Collen, C. Waterman & J. E. M. Baillie. 2007. Mammals on the EDGE: conservation priorities based on threat and phylogeny. PloS One 2 (3): e296.

[MB86] Matson, J. O. & R. H. Baker. 1986. Mammals of Zacatecas. Special Publications, Museum of Texas Tech University 24: 1–88.

[MJ11] Meredith, R. W., J. E. Janečka, J. Gatesy, O. A. Ryder, C. A. Fisher, E. C. Teeling, A. Goodbla, E. Eizirik, T. L. L. Simão, T. Stadler, D. L. Rabosky, R. L. Honeycutt, J. J. Flynn, C. M. Ingram, C. Steiner, T. L. Williams, T. J. Robinson, A. Burk-Herrick, M. Westerman, N. A. Ayoub, M. S. Springer & W. J. Murphy. 2011. Impacts of the Cretaceous terrestrial revolution and KPg extinction on mammal diversification. Science 334: 521–524.