Ceratomorpha

Lowland tapir Tapirus terrestris, copyright Geoff Gallice.

Belongs within: Perissodactyla.
Contains: Rhinocerotidae.

The Ceratomorpha include the modern tapirs and rhinoceroses. Compared to the other living perissodactyls, the horses, ceratomorphs are graviportal, heavily built animals.

Synapomorphies (from Hooker & Dashzeveg 2004): M₃ hypolophid complete; P₃ paraconid strong and approaching height of paraconid; upper molars with metaconal fold consistently joined to metaconule forming complete metaloph; lower preultimate molar with hypolophid complete, comprising equal buccal and lingual segments joined into long unnotched loph, with hypoconulid median; post P₁ diastema absent.

<==Ceratomorpha
    |--Pachynolophus HD04
    |    |--P. hookeri Godinot in Godinot et al. 1987 HD04
    |    |--P. lavocati F02
    |    `--P. livinerensis F02
    `--+--Karagalax mamikhelensis HD04
       `--+--Rhinocerotidae J84
          `--Tapiroidea D07
               |--Miotapirus Schlaikjer 1937 D07
               |    |--M. harrisonensis D07
               |    `--M. marslandensis D07
               |--Hyrachyus Leidy 1870 D07
               |    |--H. asiaticus D07
               |    |--H. eximius D07
               |    |--H. modestus D07
               |    `--H. singularis Cope 1875 C77
               |--Heptodon Cope 1882 D07
               |    |--H. calciculus D07
               |    |--H. minimus D07
               |    |--H. niushanensis D07
               |    |--H. posticus D07
               |    `--H. tienshanensis MHL03
               `--Tapirus [Tapiridae] J84
                    |--T. (Tapirella) bairdii G69
                    |--T. haysii R76
                    |    |--T. h. haysii R76
                    |    `--T. h. californicus R76
                    |--T. indicus J84
                    |--T. pinchaque J84
                    `--T. terrestris J84

Ceratomorpha incertae sedis:
  Protapirus F02
  Triplopus F02
  Alicornops complanatum CS04
  Didermoceros sumatrensis A71
  Amynodentopsis JJ84
  Indricotherium Forster Cooper 1911 D07
    |--I. bughtiense D07
    |--I. orgosensis D07
    |--I. prohorovi D07
    `--I. transauralicum D07
  Coelodonta Blumenbach 1807 D07
    |--C. antiquitatis FP64
    `--C. nihowanensis DW04
  Elasmotherium Fischer 1808 D07
    |--E. caucasicum D07
    |--E. inexpectatum D07
    `--E. sibiricum D07
  Parelasmotherium DW04
    |--P. linxiaense DW04
    `--P. simplum DW04
  Sinotherium DW04
  Shansirhinus Kretzoi 1942 DW04
    |--*S. brancoi (Schlosser 1903) [=Rhinoceros brancoi] DW04
    `--S. ringstromi Kretzoi 1942 DW04
  Hyracodon Leidy 1850 D07
    |--H. browni D07
    |--H. leidyanus D07
    |--H. medius D07
    |--H. nebraskensis D07
    |--H. petersodi D07
    `--H. priscidens D07
  Metamynodon Scott & Osborn 1887 D07
    |--M. chadronensis D07
    |--M. mckinneyi D07
    `--M. planifrons D07
  Cadurcodon Kretzoi 1942 D07
    |--C. ardynense D07
    `--C. saisanensis D07
  Diceratherium Marsh 1875 D07
    |--D. annectens D07
    |--D. armatum D07
    |--D. gregorii D07
    |--D. niobrarense D07
    `--D. tridactylum D07
  Menoceras Troxell 1921 D07
    |--M. arikarensis D07
    `--M. barbouri D07

*Type species of generic name indicated

REFERENCES

[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[CS04] Chaimanee, Y., V. Suteethorn, P. Jintasakul, C. Vidthayanon, B. Murandat & J.-J. Jaeger. 2004. A new orang-utan relative from the Late Miocene of Thailand. Nature 427: 439–441.

[C77] Cope, E. D. 1877. Report upon the extinct Vertebrata obtained in New Mexico by parties of the expedition of 1874. Geographical Surveys West of the One Hundredth Meridian 4 (2): i–iv, 1–370.

[DW04] Deng T., Wang X., Ni X. & Liu L. 2004. Sequence of the Cenozoic mammalian faunas of the Linxia Basin in Gansu, China. Acta Geologica Sinica (English Edition) 78 (1): 8–14.

[D07] Dixon, D. 2007. The Complete Illustrated Encyclopedia of Dinosaurs & Prehistoric Creatures. Hermes House: London.

[FP64] Fisher, J., & R. T. Peterson. 1964. The World of Birds: A comprehensive guide to general ornithology. Macdonald: London.

[F02] Froehlich, D. J. 2002. Quo vadis eohippus? The systematics and taxonomy of the early Eocene equids (Perissodactyla). Zoological Journal of the Linnean Society 134: 141–256.

[G69] Goodwin, G. G. 1969. Mammals from the State of Oaxaca, Mexico, in the American Museum of Natural History. Bulletin of the American Museum of Natural History 141 (1): 1–269, 40 pls.

[HD04] Hooker, J. J. & D. Dashzeveg. 2004. The origin of chalicotheres (Perissodactyla, Mammalia). Palaeontology 47 (6): 1363–1386.

[J84] Janis, C. 1984. Odd-toed ungulates. In: Macdonald, D. (ed.) All the World’s Animals: Hoofed Mammals pp. 40–41. Torstar Books: New York.

[JJ84] Janis, C., & P. J. Jarman. 1984. The hoofed mammals. In: Macdonald, D. (ed.) All the World’s Animals: Hoofed Mammals pp. 28–39. Torstar Books: New York.

[MHL03] Meng, J., Y. Hu & C. Li. 2003. The osteology of Rhombomylus (Mammalia, Glires): Implications for phylogeny and evolution of Glires. Bulletin of the American Museum of Natural History 275: 1–247.

[R76] Ray, C. E. 1976. Fossil marine mammals of Oregon. Systematic Zoology 25 (4): 420–436.