Phyllodoceae

Pink mountain-heath Phyllodoce empetriformis, copyright Walter Siegmund.

Belongs within: Ericaceae.

The Phyllodoceae are a clade of boreal shrubs with flowers on articulated pedicels.

Characters (from Kron et al. 2002): Shrubs. Leaves evergreen, alternate and spiral, decussate, or whorled, ericoid or non-ericoid, entire to serrulate, convolute or revolute in bud. Leaf epidermal cells not lignified. Indumentum of glandular hairs and elongate non-glandular hairs, and sometimes papillae. Inflorescence terminal, rarely axillary, spicate, racemose, or corymbose, sometimes reduced; bracts paired and more or less basal, bracteoles usually present; calyx articulated with pedicel. Flowers 4–6-merous, actinomorphic. Calyx lobes variable in size, withering persistent or persistent; corolla sympetalous or choripetalous, rotate to urceolate, campanulate, the lobes small to large. Stamens 5–10, included or exserted, the filaments straight to curved, unicellular-pubescent or smooth; anthers smooth or roughened, lacking appendages, dehiscing by slits or terminal pores. Pollen usually with viscin threads. Ovary 2–6-locular, with axile placentation; style impressed or not; stigma truncate, sometimes slightly expanded. Fruit a septicidal capsule, sometimes partly loculicidal as well; seeds with testa cells more or less isodiametric to elongated; embryo with two cotyledons.

<==Phyllodoceae [Cladothamneae, Epigaeae, Leiophylleae, Phyllodocoideae]
    |--Elliottia KJ02
    |    |--E. bracteata KJ02
    |    `--E. racemosa KJ02
    `--+--Kalmia [incl. Leiophyllum, Loiseleuria] KJ02
       |    |--K. angustifolia KJ02
       |    |--K. polifolia KJ02
       |    `--+--K. latifolia KJ02
       |       `--+--K. buxifolia (Bergius) Gift, Kron & Stevens in Kron, Judd et al. 2002 KJ02
       |          `--K. procumbens (Linnaeus) Gift, Kron & Stevens in Kron, Judd et al. 2002 (see below for synonymy) KJ02
       `--+--+--Epigaea repens KJ02
          |  `--Rhodothamnus chamaecistus KJ02
          `--+--Kalmiopsis leachiana KJ02
             `--Phyllodoce KJ02
                  |--P. breweri W81
                  |--P. caerulea KJ02
                  |--P. empetriformis KJ02
                  |--P. nipponica KJ02
                  `--P. taxifolia [incl. Menziesia coerulea] C55

Kalmia procumbens (Linnaeus) Gift, Kron & Stevens in Kron, Judd et al. 2002 [=Azalea procumbens, Loiseleuria procumbens] KJ02

*Type species of generic name indicated

REFERENCES

[C55] Candolle, A. de. 1855. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 2. Librairie de Victor Masson: Paris.

[KJ02] Kron, K. A., W. S. Judd, P. F. Stevens, D. M. Crayn, A. A. Anderberg, P. A. Gadek, C. J. Quinn & J. L. Luteyn. 2002. Phylogenetic classification of Ericaceae: molecular and morphological evidence. Botanical Review 68: 335–423.

[W81] West, E. W. 1981. Adaptive patterns in behavior of the Sierran pika, Ochotona princeps. Dissertation Abstracts International B 41: 4374.