Pteromalinae

Pachyneuron formosum, photographed by J. K. Lindsey.


Belongs within: Chalcidoidea.
Contains: Isoplatoides, Callitula, Mesopolobus, Miscogasterini, Sycoscapter, Philotrypesis, Otitesellinae, Pteromalus.

The Pteromalinae is a diverse (albeit somewhat poorly defined) group of chalcidoid wasps parasitising a wide range of host insects. The taxa listed below were treated as separate subfamilies by Bouček (1988), but these were not resolved as reciprocally monophyletic by the molecular analysis of Munro et al. (2011).

<==Pteromalinae (see below for synonymy) B88
    |--+--Pachyneuron Walker 1833 MH11, B88 (see below for synonymy)
    |  |    |--*P. formosum B88
    |  |    |--P. aphidis (Bouché 1834) [=Diplolepis aphidis, P. (Serimus) aphidis; incl. P. siphonophorae] B88
    |  |    |--P. emersoni Girault 1916 B88
    |  |    |--P. eros (Girault 1917) [=*Nepachyneuron eros] B88
    |  |    |--P. kingsleyi Girault 1916 B88
    |  |    |--P. mucronatum Girault 1917 GM79
    |  |    `--P. nelsoni Girault 1928 [incl. P. aeneus Masi 1929, Atrichoptilus aeneus] B88
    |  `--Coruna clavata MH11
    `--+--+--Cratomus [Cratominae] MH11
       |  |    `--C. megacephalus MH11
       |  `--Habritys Thomson 1878 MH11, GM79
       |       `--H. brevicornis MH11
       `--+--Miscogasterini MH11
          `--+--+--+--Sycoscapter MH11
             |  |  `--Plutothrix MH11
             |  `--+--Pegopus MH11
             |     `--Pseudocatolaccus MH11
             `--+--+--Philotrypesis B88
                |  `--Otitesellinae MH11
                |--+--Stenoselma MH11
                |  `--+--Grasseiana Abdurahiman & Joseph 1967 MH11, B88
                |     |    |--*G. callosa B88
                |     |    `--G. boschmai (Wiebes 1964) [=Sycobiella boschmai] B88
                |     `--+--Platygerrhus MH11
                |        |    |--P. affinis MH11
                |        |    `--P. columbianus [=Panstenon columbianum] B88
                |        `--Cecidostiba MH11
                |             |--C. burkei Crawford 1912 GM79
                |             `--C. fungosa MH11
                `--+--Merisus MH11
                   |    |--M. febriculosus Girault 1917 G17b
                   |    |--M. flaviventris Girault 1915 GM79
                   |    |--M. mordellistenae Crawford 1910 GM79
                   |    |--M. splendidus MH11
                   |    `--M. texensis Girault 1917 G17b
                   `--+--+--Sycoecinae MH11
                      |  |    |--Crossogaster B88
                      |  |    |--Sycoecus B88
                      |  |    |--Seres B88
                      |  |    |--Diaziella Grandi 1928 B88
                      |  |    |    |--*D. bicolour B88
                      |  |    |    `--D. falcate B88
                      |  |    `--Robertsia Bouček 1988 B88
                      |  |         |--*R. mandibularis Bouček 1988 B88
                      |  |         `--R. xylosyciae Bouček 1988 B88
                      |  `--Cheiropachys Haliday 1833 MH11, GM79
                      |       `--C. quadrum MH11
                      |--+--Nasonia Ashmead 1904 MH11, B88 [incl. Mormoniella Ashmead 1904 B88]
                      |  |    |--N. vitripennis (Walker 1836) B88 (see below for synonymy)
                      |  |    |--*Mormoniella brevicornis’ non Nasonia brevicornis Ashmead 1904 B88
                      |  |    |--N. giraulti GJR97
                      |  |    |--N. longicornis GJR97
                      |  |    `--N. peratripes Girault 1917 G17c
                      |  |--Homoporus Thomson 1878 MH11, B88 (see below for synonymy)
                      |  |    |--*H. fulvicornis [=Pteromalus fulvicornis, Merisus (*Homoporus) fulvicornis] B88
                      |  |    |--H. chalcidiphagus (Walsh & Riley 1869) (see below for synonymy) B88
                      |  |    `--H. hallami (Girault 1924) [=Arthrolytus hallami] B88
                      |  `--Pteromalus MH11
                      |--+--Coelopisthia Forster 1856 MH11, GM79
                      |  |    `--C. extenta MH11
                      |  `--Meraporus graminicola MH11
                      `--Panstenon Walker 1846 MH11, B88 [incl. Caudonia Walker 1850 B88; Panstenoninae]
                           |--*P. oxylus [=Miscogaster oxylus] B88
                           |--*Caudonia’ agylla B88
                           |--P. australe Girault 1913 [=P. australis] B88
                           |--P. australiense Girault 1913 [=P. australiensis] B88
                           |--P. bellicosum Girault 1913 [=P. bellicosus] B88
                           |--P. clarum Girault 1915 [=P. clarus] B88
                           `--P. gracile Girault 1913 [=P. gracilis] B88

Pteromalinae incertae sedis:
  Isoplatoides B88
  Callitula B88
  Dinotiscus B88
  Manineura B88
  Mesopolobus B88
  Eurydinota Förster 1878 B88
  Goidanichium Bouček 1970 B88
  Euneura B88
  Platecrizotes soudanensis B88
  Asoka B88
  Merismus Walker 1833 G26, GM79
  Pycnetron Gahan 1925 B88
    `--*P. curculionidis Gahan 1925 B88
  Apycnetron Bouček 1988 B88
    `--*A. brevithorax Bouček 1988 B88
  Frena Bouček 1988 B88
    `--*F. oculata Bouček 1988 B88
  Halticopterella Girault & Dodd in Girault 1915 B88
    `--*H. nigriflagellum Girault 1915 B88
  Mokrzeckia Mokrzecki 1933 [incl. Beierina Deluchhi 1958] B88
    `--*M. pini [=Pteromalus pini] B88
  Canberrana Bouček 1988 B88
    `--*C. hypera Bouček 1988 B88
  Nuchata Bouček 1988 B88
    `--*N. conica Bouček 1988 B88
  Fijita Bouček 1988 B88
    `--*F. leveri Bouček 1988 B88
  Pestra Bouček 1988 B88
    `--*P. splendida Bouček 1988 B88
  Neocylus Bouček 1988 B88
    `--*N. nigraeneus (Girault 1913) (see below for synonymy) B88
  Delislea Girault 1936 B88
    `--*D. pattersoni Girault 1936 B88
  Habritella Girault & Dodd in Girault 1915 B88
    `--*H. graciliventris Girault & Dodd in Girault 1915 B88
  Cairnsia Bouček 1988 B88
    `--*C. stylifera Bouček 1988 B88
  Trinotiscus Bouček 1988 B88
    `--*T. susuanus Bouček 1988 B88
  Briania Bouček 1988 B88
    `--*B. kukensis Bouček 1988 B88
  Buloloa Bouček 1988 B88
    `--*B. bidens Bouček 1988 B88
  Kratinka Bouček 1988 B88
    `--*K. brevis Bouček 1988 B88
  Chlorocytus Graham 1956 B88
    `--*C. pulchripes [=Pteromalus pulchripes] B88
  Mirekia Bouček 1988 B88
    `--*M. planiventris Bouček 1988 B88
  Inkaka Girault 1939 [incl. Leptospermophila Girault 1939] B88
    `--*I. quadridentata Girault 1939 [incl. *Leptospermophila fauna Girault 1939] B88
  Pachycrepoideus Ashmead 1904 [incl. Anisopteromalia Bouček 1954, Toxeumopsis Girault 1915] B88
    `--P. vindemmiae (Rondani 1875) (see below for synonymy) B88
  Laticlypa Bouček 1988 B88
    `--*L. sedlaceki Bouček 1988 B88
  Yanchepia Bouček 1988 B88
    `--*Y. rugosa Bouček 1988 B88
  Genangula Bouček 1988 B88
    `--*G. silvestris Bouček 1988 B88
  Sphegipterosema Girault 1913 B88
    `--*S. ferale Girault 1913 [=S. feralis] B88
  Acroclypa Bouček 1988 B88
    `--*A. pallipes Bouček 1988 B88
  Sphegipterosemella Girault 1913 B88
    `--*S. unicolor Girault 1913 [=Pterosema unicolor Girault 1914] B88
  Megadicylus Girault 1929 B88
    `--M. dubius (Girault 1917) [=Sphegipterosema dubia; incl. *M. montanus Girault 1929] B88
  Longinucha Bouček 1988 B88
    `--*L. petiolaris Bouček 1988 B88
  Paracarotomus Ashmead 1894 [=Paracaratomus (l. c.)] B88
    `--*P. cephalotes Ashmead 1894 B88
  Tsela Bouček 1988 B88
    `--*T. robusta Bouček 1988 B88
  Acanthometopon Ashmead 1904 [=Acanthometapon] B88
    `--*A. clavicorne Ashmead 1904 [=A. clavicornis] B88
  Acroclisella Girault 1915 [incl. Asyntomopus Girault & Dodd in Girault 1915] B88
    `--*A. perplexa Girault 1915 [incl. *Asyntomopus flaviscapus Girault & Dodd in Girault 1915] B88
  Hillerita Bouček 1988 B88
    `--*H. buccata Bouček 1988 B88
  Acroclisissa Girault 1933 B88
    `--*A. clypeata (Girault 1926) [=Acroclisella clypeata] B88
  Lomonosoffiella Girault 1913 B88
    `--*L. albipes Girault 1913 B88
  Maorita Bouček 1988 B88
    `--*M. reticulata Bouček 1988 B88
  Paracroclisis Girault 1913 B88
    `--*P. washingtoni Girault 1913 B88
  Notoglyptus Masi 1917 B88
    `--N. scutellaris (Dodd & Girault in Girault 1915) (see below for synonymy) B88
  Muscidifurax Girault & Sanders 1910 B88
    |--*M. raptor Girault & Sanders 1910 B88
    `--M. uniraptor GJR97
  Oxysychus Delucchi 1956 B88
    |--*O. silvestrii [=Dinarmus silvestrii] B88
    `--O. ferus (Girault 1927) [=Metastenoides ferus] B88
  Uniclypea Bouček 1976 B88
    |--*U. conica B88
    `--U. deporai Bouček 1988 B88
  Propicroscytus Szelényi 1941 [incl. Obtusiclava Subba Rao 1973] B88
    |--P. mirificus (Girault 1915) (see below for synonymy) B88
    `--P. oryzae (Subba Rao 1973) [=*Obtusiclava oryzae] B88
  Anisopteromalus Ruschka 1912 [incl. Aplastomorpha Crawford 1913] B88
    |--*A. mollis B88
    `--A. calandrae (Howard 1881) (see below for synonymy) B88
  Lariophagus Crawford 1909 [incl. Uriellomyia Girault 1915] B88
    |--*L. texanus B88
    `--L. distinguendus (Förster 1841) (see below for synonymy) B88
  Pterosemigastra Girault & Dodd in Girault 1915 B88
    |--*P. oenone Girault & Dodd in Girault 1915 B88
    `--P. nigriflagellum Girault & Dodd in Girault 1915 B88
  Roptrocerus Ratzeburg 1848 [=Pachyceras Ratzeburg 1844 (nom. rej.); incl. Roptroceroidea Ishii 1939] B88
    |--*R. xylophagorum (Ratzeburg 1844) [=*Pachyceras xylophagorum] B88
    `--*Roptroceroidea’ karafutoensis B88
  Rhaphitelus Walker 1834 (see below for synonymy) B88
    |--*R. maculatus Walker 1834 [incl. Eucerchysius scolytii] B88
    `--R. ladenbergi B88
  Stinoplus Thomson 1878 B88
    |--*S. militaris [=Etroxys (*Stinoplus) militaris] B88
    `--S. etearchus (Walker 1848) (see below for synonymy) B88
  Lampoterma Graham 1956 B88
    |--*L. viridis [=Metastenus viridis] B88
    `--L. gympicum Bouček 1988 B88
  Cyclogastrella Bukowski 1938 B88
    |--C. deplanata [incl. *C. quercina] B88
    `--C. shorti Bouček 1988 B88
  Toxeumorpha Girault 1915 [incl. Nigricolana Bouček 1976] B88
    |--*T. nigra Girault 1915 B88
    `--T. nigricola [=Trigonogastra nigricola, *Nigricolana nigricola] B88
  Pterosemopsis Girault 1917 B88
    |--*P. javensis Girault 1917 G17a
    `--P. eilogoa Bouček 1988 B88
  Neopolycystus Girault 1915 B88
    |--*N. insectifurax Girault 1915 B88
    `--N. abdominalis (Girault & Dodd in Girault 1915) [=Polycystella abdominalis] B88
  Oxyharma Bouček 1988 B88
    |--*O. subaenea (Dodd 1917) [=Pterosema subaenea] B88
    `--O. pulchra (Girault & Dodd in Girault 1915) [=Eurydinota pulchra, Trichomalopsis pulchra] B88
  Trigonogastrella Girault 1915 [incl. Cryptoprymnoides Girault 1915] B88
    |--*T. parasitica Girault 1915 B88
    `--T. rabiosa (Girault 1915) [=*Cryptoprymnoides rabiosus] B88
  Syntomopus Walker 1833 [incl. Merismorella Girault 1926] B88
    |--*S. thoracicus B88
    `--S. shakespearei (Girault 1926) [=*Merismorella shakespearei] B88
  Neocatolaccus Ashmead 1904 [incl. Paradicylus Dodd in Girault 1915] B88
    |--*N. tylodermae [=Catolaccus tylodermae] B88
    |--N. moneilemae B88
    `--N. varicornis (Dodd in Girault 1915) [=*Paradicylus varicornis] B88
  Eurydinoteloides Girault 1913 [incl. Aeronea Cameron 1913, Protolaccus Burks 1954] B88
    |--*E. americanus B88
    |--E. albiclavus Girault 1917 G17e
    `--E. longiventris B88
  Pterisemoppa Girault 1933 [incl. Lincolnanna Girault 1939] B88
    |--*P. australiensis (Girault & Dodd in Girault 1915) [=Sphegipterosema australiensis] B88
    |--P. malpighii (Girault 1939) [=*Lincolnanna malpighii] B88
    `--P. poeta Girault 1933 B88
  Epanogmus Girault 1920 B88
    |--*E. breviventris Girault 1920 B88
    |--E. bebius (Walker 1839) [=Pteromalus bebius] B88
    `--E. sanguiniventris Girault 1938 B88
  Apsilocera Bouček 1956 B88
    |--*A. verticillata B88
    |--A. australis Bouček 1988 B88
    `--A. bramleyi B88
  Klabonosa Bouček 1976 B88
    |--*K. ferox B88
    |--K. austini Bouček 1988 B88
    `--K. cabra Bouček 1988 B88
  Pseudanogmus Dodd & Girault in Girault 1915 [incl. Pseudomicromelus Gahan & Fagan 1923] B88
    |--*P. fasciipennis B88
    |--P. australia (Girault 1917) [=Dibrachys australia, Pseudomicromelus australia] B88
    `--P. silanus (Walker 1843) (see below for synonymy) B88
  Metastenus Walker 1834 [incl. Scymnophagus Ashmead 1904, Tripolycystus Dodd in Girault 1915] B88
    |--*M. concinnus B88
    |--M. sulcatus (Dodd in Girault 1915) [=*Tripolycystus sulcatus] B88
    `--*Scymnophagus’ townsendi B88
  Aiemea Bouček 1988 B88
    |--*A. albicrus Bouček 1988 B88
    |--A. flavipes Bouček 1988 B88
    `--A. monticola Bouček 1988 B88
  Psilocera Walker 1833 (see below for synonymy) B88
    |--*P. obscura B88
    |--P. pulchricornis (Girault 1915) [=*Parapolycystus pulchricornis] B88
    `--P. tennysoni (Girault 1913) (see below for synonymy) B88
  Acroclisoides Girault & Dodd in Girault 1915 B88
    |--*A. megacephalus Girault & Dodd in Girault 1915 B88
    |--A. laticeps Dodd & Girault in Girault 1915 B88
    |--A. major Dodd & Girault in Girault 1915 B88
    `--A. tectacorisi (Girault 1924) [=Pachycrepis tectacorisi] B88
  Agiommatus Crawford 1911 [incl. Polycystella Girault 1913, Pterosemella Girault 1913] B88
    |--*A. sumatraensis B88
    |--A. javensis (Girault 1917) [=Cyrtogaster javensis] B88
    |--A. paria B88
    `--A. viridis (Girault 1913) (see below for synonymy) B88
  Cryptoprymna Förster 1856 (see below for synonymy) B88
    |--*C. atra [=*Prosodes ater] B88
    |--C. africana B88
    |--C. australiensis (Girault 1913) (see below for synonymy) B88
    `--C. crucigera Bouček 1988 B88
  Dinarmus Thomson 1878 (see below for synonymy) B88
    |--*D. acutus [=Dimachus (*Dinarmus) acutus] B88
    |--D. colemani B88
    |--*Bruchobius’ laticeps B88
    |--D. simus (Girault 1915) [=*Metastenoides simus; incl. Toxeumella pulchripes Girault & Dodd 1915] B88
    `--*Oedaule’ stringifrons B88
  Eurydinotomorpha Girault 1913 B88
    |--*E. pax Girault 1913 B88
    |--E. basalis Girault 1915 B88
    |--E. fusciventris Girault 1913 B88
    |--E. grandis Girault 1915 B88
    |--E. incerta Girault 1915 B88
    `--E. monteithi Bouček 1988 B88
  Trichomalopsis Crawford 1913 (see below for synonymy) B88
    |--*T. shirakii B88
    |--T. australiensis (Girault 1926) [=Metadicylus australiensis] B88
    |--T. braconis (Dodd 1917) [=Eurydinota braconis] B88
    |--T. braconophaga (Cameron 1912) [=Trichoglenes braconophagus] B88
    |--T. iambe (Walker 1839) [=Pteromalus iambe] B88
    |--‘Pteromalus’ nidulans [=*Eupteromalus nidulans] B88
    `--‘Merisus’ subapterus Riley 1885 G17d, GM79 (see below for synonymy)
  Norbanus Walker 1843 (see below for synonymy) B88
    |--*N. dysaulus B88
    |--N. aureicornis (Dodd & Girault in Girault 1915) [=Neocaudonia aureicornis] B88
    |--N. cyaneus (Girault 1913) [=*Amicromelus cyaneus] B88
    |--N. grandaevus (Girault 1925) [=Amicromelus grandaevus] B88
    |--N. longifasciatus (Girault 1914) [=*Apirene longifasciata] B88
    |--N. scelestus (Girault 1914) [=*Epicaudonia scelestus] B88
    |--N. spinosiclava (Girault & Dodd in Girault 1915) [=*Neocaudonia spinosiclava] B88
    `--N. tenuicornis B88
  Merismomorpha Girault 1913 (see below for synonymy) B88
    |--*M. acutiventris Girault 1913 B88
    |--M. asilus (Girault 1915) [=Epipolycystus asilus] B88
    |--M. faunus Girault 1933 B88
    |--M. flavipetiole (Girault 1933) [=*Giorgionia flavipetiole] B88
    |--M. fulvicoxa Girault 1913 B88
    |--M. nigra Girault 1913 B88
    |--M. petiolata (Girault & Dodd in Girault 1915) [=Polycystella petiolata] B88
    `--M. sicarius (Girault 1915) [=Neopolycystella sicarius] B88
  Dibrachys Förster 1856 B88
    |  i. s.: D. apatelae (Ashmead 1893) GM79
    |         D. cavus B88
    |         D. daphne Girault 1917 G17c
    |         ‘Coelopisthia’ diacrisiae Crawford 1910 G17f, GM79
    |         ‘Tritneptis’ hemerocampae Girault 1908 G17f, GM79
    |--D. (Dibrachys) [incl. Coelopisthoidea Gahan 1913] B88
    |    |--*D. (D.) boucheanus [=Pteromalus boucheanus] B88
    |    `--D. (D.) boarmiae (Walker in Newman 1863) [=Pteromalus boarmiae] B88
    `--D. (Allodibrachys Bouček 1965) B88
  Trigonoderus irvingi Girault 1917 G17d
  Trigonoderoides Kamijo 1963 B88
    |--*T. nigricans [=Glyptosticha nigricans] B88
    `--T. niger Bouček 1988 B88
  Miscogasteriella Girault 1915 [incl. Glyptosticha Masi 1926] B88
    |--*M. longiventris Girault 1915 B88
    |--M. flavipes [=*Glyptosticha flavipes] B88
    `--M. sulcata (Kamijo 1963) [=Glyptosticha sulcata] B88
  Apocrypta Coquerel 1855 (see below for synonymy) B88
    |--*A. perplexa B88
    |--A. acaeta Ulenberg in Ulenberg & Pelt 1985 B88
    |--A. caudata (Girault 1915) [=*Goniogastrella caudatus] B88
    |--A. centrocampta Ulenberg in Ulenberg & Pelt 1985 B88
    |--A. channingi (Girault 1913) [=*Idarnoides channingi] B88
    |--A. macroseta Ulenberg in Ulenberg & Pelt 1985 B88
    |--A. mega Ulenberg in Ulenberg & Pelt 1985 B88
    |--A. megischia Ulenberg in Ulenberg & Pelt 1985 B88
    `--A. meromassa Ulenberg in Ulenberg & Pelt 1985 B88
  Critogaster B88
  Dobunabaa Bouček 1988 B88
    `--*D. flava Bouček 1988 B88
  Tenka Bouček 1988 B88
    `--*T. percaudata Bouček 1988 B88
  Watshamiella Wiebes in Bouček, Watsham & Wiebes 1981 [=Metidarnes Riek 1970 (n. n.)] B88
    |--*W. alata B88
    `--W. aurea (Girault 1919) [=Philotrypesis aurea] B88
  Calolelaps Timberlake 1925 D07
    |--*C. basalis Timberlake 1925 D07
    `--C. coeruleus Timberlake 1925 D07
  Mesolelaps Ashmead 1901 D07 [=Mesolaelaps Schulz 1906 B88; Mesolelapini]
    `--*M. cyaneiventris Ashmead 1901 D07
  Neolelaps Ashmead 1901 D07
    |--*N. hawaiiensis Ashmead 1901 D07
    `--N. flavipes Ashmead 1901 D07
  Stictolelaps Timberlake 1925 D07
    |--*S. flaviventris Timberlake 1925 D07
    `--S. stigmatus Timberlake 1925 D07
  Eutelisca D07
  Dinarmolaelaps Masi 1917 D07
    |--*D. protus Masi 1917 D07
    `--D. vatomandryi Risbec 1952 D07

Agiommatus viridis (Girault 1913) [=*Pterosemella viridis; incl. *Polycystella fasciativentris Girault 1913] B88

Anisopteromalus calandrae (Howard 1881) [=Pteromalus calandrae; incl. Neocatolaccus australiensis Girault 1913, Aplastomorpha australiensis, *Ap. pratti, Meraporus vandinei, Ap. vandinei] B88

Apocrypta Coquerel 1855 [incl. Agaonella Baker 1913, Goniogaster Mayr 1885, Goniogastrella Girault & Dodd in Girault 1915, Idarnoides Girault 1913; Apocryptini] B88

Cryptoprymna Förster 1856 [=Prosodes Walker 1833 non Eschscholz 1829; incl. Polycystelomorpha Girault 1915] B88

Cryptoprymna australiensis (Girault 1913) [=Syntomopus australiensis; incl. *Polycystelomorpha flavifemur Girault 1915] B88

Dinarmus Thomson 1878 [incl. Bruchobius Ashmead 1904, Metastenoides Girault 1915, Oedaule Waterston 1922, Sphaerakis Masi 1924] B88

Homoporus Thomson 1878 MH11, B88 [incl. Merisoporus Masi 1924 B88, Parapteromalus Ashmead 1904 B88, Phaenacra Förster 1878 B88, Pseudomerisus Erdös & Novitzky 1953 B88]

Homoporus chalcidiphagus (Walsh & Riley 1869) [=Semiotellus chalcidiphagus, Merisoporus chalcidiphagus] B88

Lariophagus distinguendus (Förster 1841) [=Pteromalus distinguendus; incl. Nasonia miltoni Girault 1929, *Uriellomyia resoluta Girault 1915] B88

Merismomorpha Girault 1913 [incl. Epipolycystus Girault 1915, Giorgionia Girault 1933, Neopolycystella Girault 1915] B88

‘Merisus’ subapterus Riley 1885 G17d, GM79 [=*Nemicromelus subapterus G17d; incl. M. fulvipes (Forbes 1885) G17d, GM79]

Nasonia vitripennis (Walker 1836) B88 [=Pteromalus vitripennis B88, Mormoniella vitripennis A71; incl. *Nasonia brevicornis Ashmead 1904 B88]

*Neocylus nigraeneus (Girault 1913) [=Polycysteloides nigraeneus; incl. Aplastomorpha reticulata Dodd in Girault 1915] B88

Norbanus Walker 1843 [incl. Amicromelus Girault 1913, Apirene Girault 1914, Arthrolysis Förster 1856, Epicaudonia Girault 1914, Neocaudonia Dodd in Girault 1915, Picroscytus Thomson 1878, Stylophorella Ashmead 1904] B88

Notoglyptus scutellaris (Dodd & Girault in Girault 1915) [=Merismus scutellaris; incl. *N. niger Masi 1917, M. squamosus Girault 1915, N. virescens Masi 1917] B88

Pachycrepoideus vindemmiae (Rondani 1875) [=Pteromalus vindemmiae, Pa. vindemiae (l. c.); incl. *Anisopteromalia crassinervis, Toxeumella dissimilis Dodd & Girault in Girault 1915, Pterosemoidea drosophilae Dodd 1917, *Pachycrepoideus dubius Ashmead 1904, *Toxeumopsis nigra Girault 1915] B88

Pachyneuron Walker 1833 MH11, B88 [incl. Atrichoptilus Delucchi 1955 B88, Eupachyneuron Blanchard 1948 B88, Nepachyneuron Girault 1917 B88, Propachyneuronia Girault 1917 B88, Serimus Brèthes 1913 B88]

Propicroscytus mirificus (Girault 1915) [=Arthrolysis mirificus; incl. A. flaviventris Girault & Dodd in Girault 1915, Propicroscytus flaviventris, P. indicus Subba Rao 1981, Arthrolysis trilongifasciatus Girault 1915, *P. trilongifasciatus] B88

Pseudanogmus silanus (Walker 1843) [=Micromelus silanus, M. silvanus (l. c.), *Pseudomicromelus silanus, *Pseudomicromelus silvanus (l. c.); incl. Pseudanogmus fasciipennis Dodd in Girault 1915, Pseudanogmus fuscipes Girault 1915, Pteromalus stironotus Cameron 1912] B88

Psilocera Walker 1833 [incl. Dichalysis Förster 1856, Eupsilocera Westwood 1839, Metopon Walker 1834, Parapolycystus Girault & Dodd in Girault 1915, Polycystoides Girault 1913] B88

Psilocera tennysoni (Girault 1913) [=*Polycystoides tennysoni; incl. Parapolycystus clavicornis Dodd in Girault 1915] B88

Pteromalinae [Bruchobiini, Dinarmini, Merisinae, Mischogastrides, Miscogasteridae, Miscogasterinae, Miscogasteroidae, Pteromalina, Pteromalini, Sphegigasterinae, Sycoryctinae, Sycoryctini, Trigonoderini] B88

Rhaphitelus Walker 1834 [incl. Eucerchysius Brèthes 1913, Storthygocerus Ratzeburg 1848, Styloceras Ratzeburg 1844; Rhaphitelini] B88

Stinoplus etearchus (Walker 1848) [=Pteromalus etearchus; incl. Etroxys (Stinoplus) aureolus (Thomson 1878)] B88

Trichomalopsis Crawford 1913 [incl. Eupteromalus Kurdjumov 1913, Metadicylus Girault 1926, Nemicromelus Girault 1917] B88

*Type species of generic name indicated

REFERENCES

[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[D07] Desjardins, C. A. 2007. Phylogenetics and classification of the world genera of Diparinae (Hymenoptera: Pteromalidae). Zootaxa 1647: 1-88.

[GJR97] Giordano, R., J. J. Jackson & H. M. Robertson. 1997. The role of Wolbachia bacteria in reproductive incompatibilities and hybrid zones of Diabrotica beetles and Gryllus crickets. Proceedings of the National Academy of Sciences of the USA 94: 11439-11444.

[G17a] Girault, A. A. 1917a. New Javanese Hymenoptera. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 59-71).

[G17b] Girault, A. A. 1917b. Descriptiones stellarum novarum. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 80-101).

[G17c] Girault, A. A. 1917c. Descriptiones hymenopterorum chalcidoidicarum variorum cum observationibus. III. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 102-111).

[G17d] Girault, A. A. 1917d. Descriptiones hymenopterorum chalcidoidicarum variorum cum observationibus. V. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 116-131).

[G17e] Girault, A. A. 1917e. Chalcidoidea nova Marilandensis. III. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 132-137).

[G17f] Girault, A. A. 1917f. Descriptiones hymenopterorum chalcidoidicarum variorum cum observationibus. V. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 116-131).

[G26] Girault, A. A. 1926. New pests from Australia. IV. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 205).

[GM79] Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 1-400.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.

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