Heydenia pretiosa, from Entomology WSL.

Belongs within: Chalcidoidea.
Contains: Cleonymus.

The Cleonyminae was originally established for pteromalid wasps with a large conical pronotum. Most species are parasites of beetle larvae in wood (Bouček 1988).

Characters (from Bouček 1988): Occipital carina indistinct; pronotum large and subconical, dorsally rounded or at least without transverse edge, medially often with smooth strip or raised longitudinal line; hind margin only moderately emarginate; thorax usually punctured; body without long sparse hairs; scutellum without sublateral grooves, without distinct frenal line or frenal bristles; propodeum without deep petiolar emargination.

<==Cleonyminae [Cleonymidae, Cleonymides] B88
    |  i. s.: Agrilocida B88
    |         Bouckekius D07
    |--Heydeniini B88
    |    |--Heydeniopsis B88
    |    `--Heydenia Förster 1856 (see below for synonymy) B88
    |         |--*H. pretiosa B88
    |         |--H. burgeoni [=Lycisca burgeoni, *Risbecisca burgeoni] B88
    |         |--H. cristatipennis (Girault 1924) [=Paraheydenia cristatipennis] B88
    |         |--H. longicollis (Cameron 1912) [=*Paraheydenia longicollis] B88
    |         |--*Heydenisca’ mateui B88
    |         |--*Pterooderella’ ornata B88
    |         `--H. trinodis Bouček 1988 B88
    `--Cleonymini B88
         |  i. s.: Amarisca Delucchi 1962 B88
         |         Pannoniella Erdös 1960 B88
         |         Paracleonymus B88
         |         Notanisus Walker 1837 B88 [incl. Bekiliella Risbec 1952 D07]
         |           |--*N. versicolor B88
         |           |--N. cyanea (Risbec 1952) [=*Bekiliella cyanea] D07
         |           |--N. richteri (Girault 1922) [=Systolomorphella richteri; incl. S. dinotipennis Girault 1922] B88
         |           `--N. zebra Bouček 1988 B88
         |         Zolotarewskya Risbec 1955 [incl. Pseudocleonymus Hedqvist 1967] B88
         |           |--*Z. seyrigi B88
         |           |--Z. shakespearei (Girault 1926) [=Systolomorphella shakespearei] B88
         |           `--*Pseudocleonymus’ spinifer B88
         |--+--Cleonymus MH11
         |  `--Dasycleonymus MH11
         `--Callocleonymus Masi 1940 MH11, B88
              |--*C. pulcher B88
              |--C. ferrierei Kerrich 1957 B88
              `--C. swezeyi (Yoshimoto & Ishii 1965) [=Ptinobius swezeyi] B88

Heydenia Förster 1856 [incl. Heydenisca Hedqvist 1967, Paraheydenia Cameron 1912, Pterooderella Risbec 1952, Risbecisca Hedqvist 1960] B88

*Type species of generic name indicated


[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[D07] Desjardins, C. A. 2007. Phylogenetics and classification of the world genera of Diparinae (Hymenoptera: Pteromalidae). Zootaxa 1647: 1-88.

[GM79] Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 1-400.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.

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