Limnoriidea

Gribbles Limnoria lignorum, from NOAA.

Belongs within: Scuticoxifera.

The Limnoriidea are a group of marine isopods united by the absence of a mandibular molar (Brandt & Poore 2003). The gribble Limnoria lignorum is a significant pest of marine timbers, gnawing into exposed wood (Hatch 1947).

Characters (from Brandt & Poore 2003): Pleonites 1–5 or 1–4 free, pleonite 1 similar to others or narrower. Pleotelson underside flat, without ventrolateral ridges (pleopods not enclosed laterally). Antenna 1 with or without scale. Mandibular lacinia mobilis present or absent. Mandibular molar absent. Maxillipedal enditre reaching at least distal margin of palp article 4, non-tapering, slender (except in Keuphylia). Dorsal coxal plate 7 similar to others. Pleopods 3–5 endopods more or less oval or rectangular, rounded distally. Uropodal rami ventral to pleotelson, articulating from side to side along vertical axis or distal.

<==Limnoriidea [Limnorioidea] BP03
    |--Keuphylia [Keuphyliidae] BP03
    |    `--K. nodosa Bruce 1980 BP03
    `--+--Hadromastax Bruce 1988 [Hadromastacidae] BP03
       |    `--H. polynesica Bruce & Müller 1991 BP03
       `--Limnoriidae BP03
            |--Paralimnoria andrewsi (Calman 1910) BP03
            |--Lynseia [Lynseiidae] BP03
            `--Limnoria BP03
                 |--L. lignorum H47
                 |--L. pfefferi Stebbing 1904 MK83
                 |--L. platycauda Menzies 1957 MK83
                 |--L. quadripunctata Holthuis 1949 BP03
                 |--L. saseboensis Menzies 1957 MK83
                 |--L. simulata Menzies 1957 MK83
                 |--L. stephenseni BP03
                 `--L. tripunctata Menzies 1951 MK83

*Type species of generic name indicated

REFERENCES

[BP03] Brandt, A. & G. C. B. Poore. 2003. Higher classification of the flabelliferan and related Isopoda based on a reappraisal of relationships. Invertebrate Systematics 17: 893–923.

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[MK83] Menzies, R. J., & W. L. Kruczynski. 1983. Isopod Crustacea (exclusive of Epicaridea). Memoirs of the Hourglass Cruises 6 (1): 1–126.

Cirolaninae

Cirolana harfordi, copyright Peter J. Bryant.

Belongs within: Cymothoida.

The Cirolaninae are a group of mostly marine isopods with a secondary unguis present on the pereopodal dactyli (Kensley & Schotte 1989).

Characters (from Kensley & Schotte 1989): Frontal lamina short, flat. Clypeus flattened, not projecting. Antennal peduncular articles 4 and 5 subequal, longer than articles 1–3. Pereopods with secondary unguis on dactyli. Penes reduced or absent. Pleonite 5 always overlapped by pleonite 4. Pleopod 2 in male with copulatory stylet articulating basally.

<==Cirolaninae
    |--Parabathynomus BBN86
    |--Hansenolana sphaeromiformis (Hansen 1890) BBN86, MK83
    |--Neocirolana BBN86
    |--Cartetolana BBN86
    |--Gnatholana BBN86
    |--Oncilorpheus stebbingi Paul & Menzies 1971 BBN86, MK83
    |--Creaseriella Rioja 1953 BBN86
    |    `--C. anops (Creaser 1936) BBN86
    |--Saharolana Monod 1930 BBN86
    |    `--S. seurati Monod 1930 BBN86
    |--Antrolana Bowman 1964 BBN86
    |    `--A. lira Bowman 1964 BBN86
    |--Cirolanides Benedict 1896 BBN86
    |    `--C. texensis Benedict 1896
    |--Mexilana Bowman 1975 BBN86
    |    `--M. saluposi Bowman 1975 BBN86
    |--Faucheria Dollfus & Viré 1905 BBN86
    |    `--F. faucheri (Dollfus & Viré 1900) BBN86
    |--Bathynomus Milne Edwards 1879 BBN86, BP03
    |    |--B. giganteus Milne Edwards 1879 MK83
    |    `--B. immanis Bruce 1986 BP03
    |--Haptolana Bowman 1966 BBN86
    |    |--H. pholeta Bruce & Humphreys 1993 PH13
    |    |--H. somala Messana & Chelazzi 1984 BBN86
    |    `--H. trichostoma Bowman 1966 BBN86
    |--Bahalana Carpenter 1981 BBN86
    |    |--B. cardiopus Notenboom 1981 BBN86
    |    `--B. geracei Carpenter 1981 BBN86
    |--Turcolana Argano & Pesce 1980 BBN86
    |    |--T. cariae Argano & Pesce 1980 BBN86
    |    `--T. rhodica Botosaneanu, Boutin & Henry 1985 BBN86
    |--Skotobaena Ferrara & Monod 1972 BBN86
    |    |--S. monodi Ferrara & Lanza 1978 BBN86
    |    `--S. mortoni Monod 1972 BBN86
    |--Sphaerolana Cole & Minckley 1972 BBN86
    |    |--S. affinis Cole & Minckley 1970 BBN86
    |    `--S. interstitialis Cole & Minckley 1970 BBN86
    |--Anopsilana Paulian & Delamare Deboutteville 1956 (see below for synonymy) BBN86
    |    |--A. acanthura (Notenboom 1981) BBN86
    |    |--A. crenata Bowman & Franz 1982 BBN86
    |    |--A. cubensis (Hay 1903) BBN86
    |    |--A. poissoni Paulian & Delamare Deboutteville 1956 BBN86
    |    `--A. radicicola (Notenboom 1981) BBN86
    |--Speocirolana Bolivar y Pieltain 1950 BBN86
    |    |--S. bolivari (Rioja 1953) BBN86
    |    |--S. endeca Bowman 1981 BBN86
    |    |--S. guerrai Contreras-Balderas & Purata-Velarde 1982 BBN86
    |    |--S. pelaezi (Bolivar y Pieltain 1950) BBN86
    |    |--S. pubens Bowman 1981 BBN86
    |    `--S. thermydronis Cole & Minckley 1966 BBN86
    |--Sphaeromides Dollfus 1897 BBN86
    |    |--S. bureschi Strouhal 1963 BBN86
    |    |    |--S. b. bureschi BBN86
    |    |    `--S. b. serbica Pljakič 1968 BBN86
    |    |--S. polateni Angelov 1968 BBN86
    |    |--S. raymondi Dollfus 1897 BBN86
    |    `--S. virei (Brian 1923) [=Troglaega virei] BBN86
    |         |--S. v. virei BBN86
    |         |--S. v. mediodalmatina Sket 1964 BBN86
    |         `--S. v. montenigrina Sket 1957 BBN86
    |--Typhlocirolana Racovitza 1905 BBN86
    |    |--T. buxtoni Racovitza 1912 BBN86
    |    |--T. fontis (Gurney 1908) BBN86
    |    |--T. gurneyi Racovitza 1912 BBN86
    |    |--T. leptura Botosaneanu, Boutin & Henry 1985 BBN86
    |    |--T. lulli Pujiula 1911 BBN86
    |    |--T. moraguesi Racovitza 1905 BBN86
    |    |--T. reichi Por 1962 BBN86
    |    |--T. rifana Margalef 1958 BBN86
    |    `--T. steinitzi Strouhal 1960 BBN86
    `--Cirolana Leach 1818 MK83
         |--*C. cranchii Leach 1818 MK83
         |--C. borealis Lilljeborg 1851 [incl. C. spinipes Bate & Westwood 1868] MK83
         |--C. chiltoni H47
         |--C. gracilis Hansen 1890 MK83
         |--C. harfordi (Lockington 1877) HJ08
         |--C. kincaidi Hatch 1947 H47
         |--C. koepckei Bott 1954 B54
         |--C. lata BBN86
         |--C. leptanga Bruce 1994 BP03
         |--C. linguifrons B54
         |--C. minuta Hansen 1890 MK83
         |--C. obtruncata Richardson 1901 MK83
         |--C. parva Hansen 1890 [incl. C. albida Richardson 1901, C. diminuta Menzies 1962] MK83
         |--C. polita (Stimpson 1854) [=Aega polita, Conilera polita] MK83
         |--C. salvadorensis Schuster 1954 S54
         |--C. troglexuma FBI01
         `--C. vancouverensis [=C. chiltoni vancouverensis] H47

Anopsilana Paulian & Delamare Deboutteville 1956 [incl. Haitilana Notenboom 1981, Troglocirolana Rioja 1956] BBN86

*Type species of generic name indicated

REFERENCES

[BBN86] Botosaneanu, L., N. Bruce & J. Notenboom. 1986. Isopoda: Cirolanidae. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 412–422. E. J. Brill/Dr. W. Backhuys: Leiden.

[B54] Bott, R. 1954. Ein neuer litoraler Isopod aus Peru. Senckenbergiana Biologica 35: 107–108.

[BP03] Brandt, A. & G. C. B. Poore. 2003. Higher classification of the flabelliferan and related Isopoda based on a reappraisal of relationships. Invertebrate Systematics 17: 893–923.

[FBI01] Fosshagen, A., G. A. Boxshall & T. M. Iliffe. 2001. The Epacteriscidae, a cave-living family of calanoid copepods. Sarsia 86: 245–318.

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[HJ08] Huisman, J. M., D. S. Jones, F. E. Wells & T. Burton. 2008. Introduced marine biota in Western Australian waters. Records of the Western Australian Museum 25 (1): 1–44.

Kensley, B., & M. Schotte. 1989. Guide to the Marine Isopod Crustaceans of the Caribbean. Smithsonian Institution Press: Washington.

[MK83] Menzies, R. J., & W. L. Kruczynski. 1983. Isopod Crustacea (exclusive of Epicaridea). Memoirs of the Hourglass Cruises 6 (1): 1–126.

[PH13] Poore, G. C. B. & W. F. Humphreys. 2013. Bunderanthura bundera gen. et sp. nov. from Western Australia, first anchialine Leptanthuridae (Isopoda) from the Southern Hemisphere. Records of the Western Australian Museum 28 (1): 21–29.

[S54] Schuster, O. 1954. Zwei neue Crustaceen von der pazifischen Küste Mittel-Amerikas (Amphipoda und Isopoda). Senckenbergiana Biologica 35: 103–105.

Bopyridae

Dorsal (left) and ventral (centre) views of female and dorsal view of male (right) of Parabopyrella richardsonae, from Marilyn Schotte.

Belongs within: Epicaridea.

The Bopyridae are a group of isopods parasitic as adults on decapod crustaceans, while larvae (where known) are parasites of calanoid copepods (Markham 1985).

Characters (from Markham 1985): Female distinctly segmented, slightly to greatly asymmetrical, depressed. Head occasionally fused with first pereomere, sometimes enclosed by frontal lamina; antenna rudimentary; maxilliped of two segments, often with anteromedial palp; posteroventral border of head usually with one or two lateral projections on each side. All seven pereomeres usually distinct; pereopods reduced, usually of seven pairs, though up to six occasionally absent from one side; pereopods prehensile, dorsal to lateral, isomorphic; oostegites of five (rarely seven) pairs often produced into lateral plates resembling pleopodal rami; pleopods generally present on all but final pleomere; pleopods rudimentary or modified for respiration, uniramous or biramous, isomorphic; uropods, when present, terminal, uniramous or biramous, often of same form as pleopods or lateral plates. Male much smaller than female, at least twice as long as wide, symmetrical, distinctly segmented. Head rounded anteriorly, occasionally fused with first pereomere; antennae often prominent. Pereon of seven distinct pereomeres; pair of ventral prehensile pereopods on each pereomere, usually isomorphic except first and sometimes second rarely conspicuously larger than others. Pleon of one to six pleomeres; if unsegmented, usually lacking appendages; if multisegmented, often with sessile to flaplike uniramous or biramous pleopods on each pleomere but last; uropods, if present, uniramous or biramous and terminal.

<==Bopyridae [Bopyrina]
    |  i. s.: Leidya distortus (Leidy 1855) [=Cepon distortus] M85
    |         Munidion parvum H47
    |         Phyllodurus abdominalis H47
    |--Athelginae M85
    |--Ione H47 [Ioninae M85]
    |    `--I. cornuta [incl. I. brevicauda Bonnier 1900] H47
    |--Argeia Dana 1852 [Argeiinae] M85
    |    |--*A. pugettensis Dana 1852 M85
    |    `--A. atlantica Markham 1977 M85
    |--Pseudioninae M85
    |    |--Pontobopyrus Markham 1979 M85
    |    |    `--*P. abyssorum Markham 1979 M85
    |    `--Pseudione Kossmann 1881 M85
    |         |--*P. callianassae Kossmann 1881 M85
    |         |--P. affinis (Sars 1882) [=Pleurocrypta affinis; incl. Palaegyge plesionikae Barnard 1920] M85
    |         |--P. cognata Markham 1985 M85
    |         |--P. elongata M85
    |         |--P. galacanthae H47
    |         |--P. giardi H47
    |         |--P. indica M85
    |         |--P. magna Shiino 1951 M85
    |         `--P. tattersalli M85
    |--Hemiarthrinae [Phrixinae, Phryxinae] M85
    |    |--Allodiplophryxus Markham 1985 M85
    |    |    `--*A. floridanus Markham 1985 M85
    |    |--Loki Markham 1972 M85
    |    |    `--*L. circumsaltanus Markham 1972 M85
    |    |--Azygopleon Markham 1985 M85
    |    |    `--*A. schmitti (Pearse 1932) [=Hemiarthrus schmitti] M85
    |    |--Eophrixus Caroli 1930 [incl. Hypophryxus Shiino 1934] M85
    |    |    |--*E. lysmatae (Caroli 1930) [=Phrixus (Eophrixus) lysmatae] M85
    |    |    `--E. subcaudalis (Hay 1917) (see below for synonymy) M85
    |    |--Dicropleon Markham 1972 M85
    |    |    |--*D. periclimenis Markham 1972 M85
    |    |    `--D. morator Markham 1982 M85
    |    |--Hyperphrixus Nierstrasz & Brender à Brandis 1931 M85
    |    |    |--*H. tattersalli Nierstrasz & Brender à Brandis 1931 M85
    |    |    `--H. castrensis Markham 1985 M85
    |    |--Metaphrixus Nierstrasz & Brender à Brandis 1931 M85
    |    |    |--*M. carolii Nierstrasz & Brender à Brandis [=Metaphryxus carolii] M85
    |    |    `--M. intutus Bruce 1965 M85
    |    |--Hemiarthrus Giard & Bonnier 1887 [=Phryxus (preoc.), Phrixus Caroli 1930] M85
    |    |    |--*H. abdominalis (Krøyer 1840-1841) (see below for synonymy) M85
    |    |    |--H. nematocarcini Stebbing 1914 M85
    |    |    `--H. synalphei (Pearse 1950) [=Diplophryxus synalphei] M85
    |    `--Diplophryxus Richardson 1904 M85
    |         |--*D. jordani Richardson 1904 M85
    |         |--D. alphei Shiino 1934 M85
    |         |--D. kempi Chopra 1930 M85
    |         `--D. richardsoni Chopra 1930 M85
    `--Bopyrinae M85
         |--Schizobopyrina M85
         |--Bopyrus Latreille 1802 M85
         |    `--*B. crangorum (Fabricius 1798) [=Monoculus crangorum, B. squillarum Latreille 1802] M85
         |--Bathygyge Hansen 1897 M85
         |    `--*B. grandis Hansen 1897 M85
         |--Probopyrinella Nierstrasz & Brender à Brandis 1929 M85
         |    `--*P. latreuticola (Gissler 1882) (see below for synonymy) M85
         |--Urobopyrus Richardson 1904 M85
         |    `--*U. processae Richardson 1904 [=U. processi (l. c.); incl. U. provisorius Rouch & Taberly 1961] M85
         |--Parabopyriscus Markham 1985 M85
         |    `--*P. stellatus Markham 1985 M85
         |--Probopyria Markham 1985 M85
         |    `--*P. alphei (Richardson 1900) (see below for synonymy) M85
         |--Ovobopyrus Markham 1985 M85
         |    `--*O. alphezemiotes Markham 1985 M85
         |--Bopyriscus Richardson 1905 M85
         |    `--*B. calmani Richardson 1905 M85
         |--Bopyroides Stimpson 1864 M85
         |    |--B. hippolytes (Krøyer 1838) (see below for synonymy) M85
         |    `--B. cluthae M85
         |--Bopyrinella Nierstrasz & Brender à Brandis 1925 M85
         |    |--B. thorii (Richardson 1904) (see below for synonymy) M85
         |    |--B. albida Shiino 1958 M85
         |    `--B. nipponica Shiino 1936 [=B. antillensis var. nipponica] M85
         |--Probopyrus Giard & Bonnier 1888 [incl. Palaegyge Giard & Bonnier 1888, Palegyge] M85
         |    |--*P. ascendens (Semper 1881) [=Bopyrus ascendens] M85
         |    |--‘*Palaegyge’ borrei Giard & Bonnier 1888 M85
         |    |--P. palaemoni Lemos de Castro & Brasil Lima 1974 M85
         |    `--P. pandalicola (Packard 1879) (see below for synonymy) M85
         |--Bopyrione Bourdon & Markham 1980 M85
         |    |--*B. synalphei Bourdon & Markham 1980 M85
         |    |--B. longicapitata Markham 1982 M85
         |    |--B. toloensis Markham 1982 M85
         |    `--B. woodmasoni (Chopra 1923) M85
         |--Synsynella Hay 1917 [incl. Bopyro Pearse 1932, Prosynsynella Nierstrasz & Brender à Brandis 1929] M85
         |    |--*S. deformans Hay 1917 [=Bopyrella deformans] M85
         |    |--‘*Prosynsynella’ hayi Nierstrasz & Brender à Brandis 1929 (see below for synonymy) M85
         |    |--‘Bopyrella’ inoi Shiino 1949 M85
         |    `--S. integra Bourdon 1981 M85
         |--Bopyrella Bonnier 1900 M85
         |    |--*B. thomsoni Bonnier 1900 M85
         |    |--B. harmopleon Bowman 1956 M85
         |    |--B. macginitiei M85
         |    `--B. malensis M85
         |--Bopyrina Kossmann 1881 M85
         |    |--*B. ocellata (Czerniavsky 1869) [=Bopyrus ocellatus; incl. Bopyrina giardi] M85
         |    |--B. abbreviata Richardson 1904 M85
         |    |--B. choprae M85
         |    |--B. crangona Pearse 1953 S69
         |    |--B. gigas M85
         |    |--B. pleurocephala M85
         |    `--B. sewelli M85
         `--Parabopyrella Markham 1985 M85
              |--*P. mortenseni (Nierstrasz & Brender à Brandis 1929) [=Bopyrella mortenseni] M85
              |--P. delagoae (Bourdon 1982) [=Bopyrella delagoae] M85
              |--P. incisa (Bourdon 1982) [=Bopyrella incisa] M85
              |--P. lata (Nierstrasz & Brender à Brandis 1929) [=Bopyrella lata] M85
              |--‘Bopyrella’ nierstraszi M85
              |--P. richardsonae (Nierstrasz & Brender à Brandis 1929) [=Bopyrella richardsonae] M85
              |--P. saronae (Bourdon & Bruce 1979) [=Bopyrella saronae] M85
              `--P. thomasi (Nierstrasz & Brender à Brandis 1929) [=Bopyrella thomasi] M85

Bopyrinella thorii (Richardson 1904) [=Bopyrina thorii; incl. *Bopyrinella antillensis Nierstrasz & Brender à Brandis 1925]

Bopyroides hippolytes (Krøyer 1838) [=Bopyrus hippolytes, Bobyroides (l. c.) hippolytes, Gyge hippolytes, Pleurocrypta hippolytes; incl. *Bopyroides acutimarginatus Stimpson 1864] M85

Eophrixus subcaudalis (Hay 1917) [=Phryxus subcaudalis, Hemiarthrus subcaudalis, Phrixus (Paraphrixus) subcaudalis] M85

*Hemiarthrus abdominalis (Krøyer 1840-1841) [=Bopyrus abdominalis, *Phrixus abdominalis, *Phryxus abdominalis; incl. Phryxus hippolytes Rathke 1843] M85

*Probopyria alphei (Richardson 1900) [=Bopyrus alphei, Bopyrella alphei, Probopyrus alphei; incl. Capitetragonia asperotibialis Pearse 1953] M85

*Probopyrinella latreuticola (Gissler 1882) [=Bopyroides latreuticola, Bopyrina latreuticola, Probopyrus latreuticola; incl. Bopyrella asymmetrica Nierstrasz & Brender à Brandis 1929, Bopyrus latreutis Bate 1888] M85

Probopyrus pandalicola (Packard 1879) [=Bopyrus pandalicola, Pr. pandicola (l. c.); incl. Pr. bithynis Richardson 1904, Pr. bythinis (l. c.), Pr. creaseri Pearse 1936, Pr. floridensis Richardson 1904, Pr. bithynis var. gigas Nierstrasz & Brender à Brandis 1929, Pr. floridensis var. gigas, Bopyrus manhattensis Gissler 1881, Palaegyge meeki Richardson 1912, Probopyrus meeki, Pr. oviformis Nierstrasz & Brender à Brandis 1929, Pa. oviformis, Bopyrus palaemoneticola Gissler 1882, Probopyrus palaemoneticola, Pr. panamensis Richardson 1912, Pr. papaloapanensis Rioja 1949, Pr. papaloanensis (l. c.)] M85

‘*Prosynsynella’ hayi Nierstrasz & Brender à Brandis 1929 [incl. *Bopyro choprae Pearse 1932, Bopyrina choprae, Bopyrus chopre (l. c.)] M85

*Type species of generic name indicated

REFERENCES

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[M85] Markham, J. C. 1985. A review of the bopyrid isopods infesting caridean shrimps in the northwestern Atlantic Ocean, with special reference to those collected during the Hourglass Cruises in the Gulf of Mexico. Memoirs of the Hourglass Cruises 7 (3): 1–156.

S69] Steyskal, G. C. 1969. The mistreatment of the Latin genitive case in forming names of parasites. Systematic Zoology 18 (3): 339–342.

Asellus (Asellus)

Water hoglouse Asellus aquaticus, copyright Peter Pfeiffer.

Belongs within: Asellidae.

The subgenus Asellus of the genus freshwater isopod genus Asellus is characterised by a lack of spines on the propodite of the male first legs (Hatch 1947).

Characters (from Hatch 1947): Body usually elongate, depressed. Head usually narrower and shorter than first thoracic segment; eyes usually present; mandible with three-segmented palp. Abdomen usually not longer than broad. Male with inner margin of posterior surface of propodite of first legs not produced in teeth or immovable spines.

<==Asellus Geoffroy 1762 (Asellus) HLM86
    |--A. (A.) amamiensis Matsumoto 1961 HLM86
    |--A. (A.) aquaticus (Linnaeus 1758) HLM86
    |    |--A. a. aquaticus HLM86
    |    |--A. a. carsicus Karaman 1952 HLM86
    |    |--A. a. cavernicolus Racovitza 1925 HLM86
    |    |--A. a. cyclobranchialis Sket 1965 HLM86
    |    |--A. a. fribergensis Schneider 1887 HLM86
    |    `--A. a. longicornis Sket 1965 HLM86
    |--A. (A.) birsteini Levanidov 1976 HLM86
    |--A. (A.) ezoensis Matsumoto 1962 HLM86
    |--A. (A.) hilgendorfi Bovallius 1886 HLM86
    |    |--A. h. hilgendorfi Bovallius 1886 HLM86
    |    `--A. h. ryukyuensis Ueno 1938 HLM86
    |--A. (A.) hyugaensis Matusmoto 1960 HLM86
    |--A. (A.) kumaensis Matsumoto 1960 HLM86
    |--A. (A.) monticola [incl. A. monticola var. fontinalis Birstein 1936] HLM86
    |--A. (A.) musashiensis Matsumoto 1961 HLM86
    |--A. (A.) shikokuensis Matsumoto 1960 HLM86
    |--A. (A.) tamaensis Matsumoto 1961 HLM86
    `--A. (A.) tomalensis H47

*Type species of generic name indicated

REFERENCES

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[HLM86] Henry, J.-P., J. J. Lewis & G. Magniez. 1986. Isopoda: Asellota: Aselloidea, Gnathostenetroidoidea, Stenetrioidea. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 434–464. E. J. Brill/Dr. W. Backhuys: Leiden.

Asellidae

One-spotted waterlouse Proasellus meridianus, copyright Keith Lugg.

Belongs within: Asellota.
Contains: Asellus (Asellus).

The Asellidae are a group of freshwater crustacean of the Holarctic region (Henry et al. 1986).

Characters (from Henry et al. 1986): Antenna bearing a squama; maxillipedal oostegite reduced to a "Wasserstrudelapparat" (water swirling apparatus); pleonites 1 and 2 reduced both in length and width, forming a stalk between pereion and pleotelson; male copulatory organ (endopodite of pleopods 2) secondarily uniarticulate, but complex.

<==Asellidae
    |--Baicalasellus HLM86
    |--Gallasellus Henry & Magniez 1977 HLM86
    |    `--G. heilyi (Legrand 1956) HLM86
    |--Stygasellus Chappuis 1943 HLM86
    |    `--S. phreaticus (Chappuis 1943) [=Stygonectes phreaticus] HLM86
    |--Uenasellus Matsumoto 1962 HLM86
    |    `--U. iyoensis (Matsumoto 1960) HLM86
    |--Salmasellus Bowman 1975 HLM86
    |    `--S. steganothrix Bowman 1975 HLM86
    |--Psammasellus Braga 1968 HLM86
    |    `--P. capitatus Braga 1968 HLM86
    |--Calasellus Bowman 1981 HLM86
    |    |--C. californicus (Miller 1933) HLM86
    |    `--C. longus Bowman 1981 HLM86
    |--Lirceolus Bowman & Longley 1976 HLM86
    |    |--L. pilus (Steeves 1968) HLM86
    |    `--L. smithii (Ulrich 1902) HLM86
    |--Lirceus Rafinesque 1820 HLM86
    |    |--L. culveri Holsinger & Bowman 1973 HLM86
    |    |--L. fontinalis HLM86
    |    `--L. usdagalum Estes & Holsinger 1976 HLM86
    |--Nipponasellus Matsumoto 1962 HLM86
    |    |--N. aioii (Chappuis 1955) HLM86
    |    |--N. hubrichti (Matsumoto 1956) HLM86
    |    |--N. kagaensis (Msatsumoto 1958) HLM86
    |    |--N. takefuensis (Matsumoto 1961) HLM86
    |    `--N. tonensis (Matsumoto 1961) HLM86
    |--Bragasellus Henry & Magniez 1968 HLM86
    |    |--B. aireyi Henry & Magniez 1980 HLM86
    |    |--B. boui Henry & Magniez 1974 HLM86
    |    |--B. comasi Henry & Magniez 1976 HLM86
    |    |--B. comasioides Magniez & Brehier 2004 F05
    |    |--B. conimbricensis (Braga 1946) HLM86
    |    |--B. escolai Henry & Magniez 1978 HLM86
    |    |--B. frontellum (Braga 1964) HLM86
    |    |--B. incurvatus Afonso 1984 HLM86
    |    |--B. lagari Henry & Magniez 1973 HLM86
    |    |--B. pauloae (Braga 1958) HLM86
    |    `--B. seabrai (Braga 1943) HLM86
    |--Asellus Geoffroy 1762 HLM86
    |    |  i. s.: A. cavaticus T30
    |    |         A. meridianus N29
    |    |         A. nipponensis Nicholls 1929 N29
    |    |--A. (Asellus) HLM86
    |    |--A. (Conasellus) communis H47
    |    |--A. (Mesoasellus Birstein 1939) HLM86
    |    |    |--A. (M.) dentifer Birstein & Levanidov 1952 HLM86
    |    |    `--A. (M.) strinatii Chappuis 1955 HLM86
    |    `--A. (Phreatosellus Matsumoto 1962) HLM86
    |         |--A. (P.) akiyoshiensis (Ueno 1927) HLM86
    |         |--A. (P.) higoensis Matsumoto 1960 HLM86
    |         |--A. (P.) iriei Matsumoto 1978 HLM86
    |         |--A. (P.) kawamurai (Tattersall 1921) HLM86
    |         |--A. (P.) minatoi Matsumoto 1978 HLM86
    |         |--A. (P.) miurai Chappuis 1955 HLM86
    |         |--A. (P.) uenoi Matsumoto 1978 HLM86
    |         `--A. (P.) yoshinoensis Matsumoto 1960 HLM86
    |--Synasellus Braga 1944 HLM86
    |    |--S. albicastrensis Braga 1960 HLM86
    |    |--S. barcelensis Noodt & Galhano 1969 HLM86
    |    |--S. brigantinus Braga 1959 HLM86
    |    |--S. exiguus Braga 1944 HLM86
    |    |--S. fragilis Braga 1946 HLM86
    |    |--S. insignis Afonso 1984 HLM86
    |    |--S. lafonensis Braga 1959 HLM86
    |    |--S. longicauda Braga 1959 HLM86
    |    |--S. mariae (Braga 1942) HLM86
    |    |    |--S. m. mariae HLM86
    |    |    `--S. m. favaiensis Eiras 1974 HLM86
    |    |--S. mateusi Braga 1954 HLM86
    |    |--S. meirelesi Braga 1959 HLM86
    |    |--S. minutus Braga 1967 HLM86
    |    |--S. nobrei Braga 1967 HLM86
    |    |--S. pireslimai Braga 1959 HLM86
    |    |--S. serranus Braga 1967 HLM86
    |    `--S. transmontanus Braga 1959 HLM86
    |--Caecidotea Packard 1871 HLM86
    |    |--C. acuticarpa Mackin & Hubricht 1940 HLM86
    |    |--C. adenta Mackin & Hubricht 1940 HLM86
    |    |--C. alabamensis Stafford 1911 HLM86
    |    |--C. ancyla (Fleming 1972) HLM86
    |    |--C. antricola (Creaser 1931) HLM86
    |    |--C. barri (Steeves 1965) HLM86
    |    |--C. beattyi Lewis & Bowman 1981 HLM86
    |    |--C. bicrenata (Steeves 1963) HLM86
    |    |    |--C. b. bicrenata HLM86
    |    |    `--C. b. whitei Lewis & Bowman 1981 HLM86
    |    |--C. bisetus (Steeves 1968) HLM86
    |    |--C. bowmani Lewis 1980 HLM86
    |    |--C. brevicauda HLM86
    |    |--C. cannula (Steeves 1963) HLM86
    |    |--C. carolinensis Lewis & Bowman 1977 HLM86
    |    |--C. catachaetus (Fleming & Steeves 1972) HLM86
    |    |--C. chiapas Bowman 1975 HLM86
    |    |--C. circulus (Steeves 1968) HLM86
    |    |--C. cyrtorhynchus (Fleming & Steeves 1972) HLM86
    |    |--C. dentadactyla HLM86
    |    |--C. dimorpha Mackin & Hubricht 1940 HLM86
    |    |--C. extensolingualus (Fleming 1972) HLM86
    |    |--C. filicispeluncae Bowman & Hobbs 1983 HLM86
    |    |--C. franzi Lewis 1981 HLM86
    |    |--C. fustis Lewis 1981 HLM86
    |    |--C. henroti (Bresson 1955) HLM86
    |    |--C. hobbsi Maloney 1939 HLM86
    |    |--C. holsingeri (Steeves 1963) HLM86
    |    |--C. incurva (Steeves 1968) HLM86
    |    |--C. jordani (Eberly 1965) HLM86
    |    |--C. kendeighi (Steeves & Seidenbergh 1971) HLM86
    |    |--C. lesliei Lewis & Bowman 1981 HLM86
    |    |--C. macropropoda Chase & Blair 1937 [incl. C. ozarkana Chase & Blair 1937] HLM86
    |    |--C. metcalfi (Fleming 1972) HLM86
    |    |--C. mitchelli Argano 1972 HLM86
    |    |--C. nickajackensis Packard 1881 HLM86
    |    |--C. nortoni (Steeves 1966) HLM86
    |    |--C. oculata Mackin & Hubricht 1940 HLM86
    |    |--C. packardi Mackin & Hubricht 1940 HLM86
    |    |--C. parva (Steeves 1964) HLM86
    |    |--C. pasquinii Argano 1972 HLM86
    |    |--C. paurotrigonus (Fleming 1972) HLM86
    |    |--C. pricei Levi 1949 HLM86
    |    |--C. recurvata (Steeves 1963) HLM86
    |    |--C. reddelli (Steeves 1968) HLM86
    |    |--C. richardsonae Hay 1901 HLM86
    |    |--C. rotunda Bowman & Lewis 1984 HLM86
    |    |--C. salemensis Lewis 1981 HLM86
    |    |--C. scrupulosa HLM86
    |    |--C. scyphus (Steeves 1968) HLM86
    |    |--C. sequoiae Bowman 1975 HLM86
    |    |--C. serrata (Fleming 1972) HLM86
    |    |--C. simonini (Bresson 1955) HLM86
    |    |--C. sinuncus (Steeves 1965) HLM86
    |    |--C. spatulata Mackin & Hubricht 1940 HLM86
    |    |--C. steevesi (Fleming 1972) HLM86
    |    |--C. stiladactyla Mackin & Hubricht 1940 HLM86
    |    |--C. stygia Packard 1871 HLM86
    |    |--C. teresae Lewis 1982 HLM86
    |    |--C. tridentata Hungerford 1922 HLM86
    |    |--C. vandeli (Bresson 1955) HLM86
    |    |--C. vomeroi Argano 1972 HLM86
    |    `--C. zullinii Argano 1972 HLM86
    `--Proasellus Dudich 1925 HLM86
         |--P. acutianus Argano & Henry 1972 HLM86
         |--P. adriaticus Argano & Pesce 1979 HLM86
         |--P. albigensis (Magniez 1965) HLM86
         |--P. ambracicus Pesce & Argano 1980 HLM86
         |--P. amiterninus Argano & Pesce 1979 HLM86
         |--P. anophtalmus (Karaman 1934) HLM86
         |    |--P. a. anophtalmus HLM86
         |    |--P. a. bosnicus (Sket 1965) HLM86
         |    |--P. a. dalmatinus (Karaman 1955) HLM86
         |    |--P. a. longisetus (Remy 1941) HLM86
         |    |--P. a. radikanus (Karaman 1955) HLM86
         |    |--P. a. rhausinus (Remy 1941) HLM86
         |    `--P. a. serbicus (Karaman 1955) HLM86
         |--P. aquaecalidae (Racovitza 1922) HLM86
         |--P. arnautovici HLM86
         |    |--P. a. arnautovici HLM86
         |    `--P. a. elongatus (Karaman 1953) HLM86
         |--P. arthrodilus (Braga 1945) HLM86
         |--P. bagradicus Henry & Magniez 1972 HLM86
         |--P. bardaunii Alouf, Henry & Magniez 1982 HLM86
         |--P. basnosanui (Codreanu & Codreanu 1962) HLM86
         |--P. bellesi Henry & Magniez 1982 HLM86
         |--P. beroni Henry & Magniez 1968 HLM86
         |--P. boui Henry & Magniez 1969 HLM86
         |--P. burgundus Henry & Magniez 1969 HLM86
         |--P. cantabricus Henry & Magniez 1968 HLM86
         |--P. cavaticus (Leydig 1871) HLM86
         |--P. chappuisi Henry & Magniez 1968 HLM86
         |--P. chauvini Henry & Magniez 1978 HLM86
         |--P. coiffaiti Henry & Magniez 1972 HLM86
         |--P. comasi Henry & Magniez 1982 HLM86
         |--P. coxalis (Dollfus 1892) HLM86
         |    |--P. c. coxalis HLM86
         |    |--P. c. africanus (Monod 1924) HLM86
         |    |--P. c. banyulensis (Racovitza 1919) HLM86
         |    |--P. c. cephallenus (Strouhal 1942) HLM86
         |    |--P. c. cyrenaicus (Arcangeli 1942) HLM86
         |    |--P. c. gabriellae (Margalef 1950) HLM86
         |    |--P. c. leucadius (Strouhal 1942) HLM86
         |    |--P. c. lucifugus (Deeleman-Reinhold 1965) HLM86
         |    |--P. c. perarmatus (Remy 1936) HLM86
         |    `--P. c. peyerimhoffi (Racovitza 1919) HLM86
         |--P. cretensis Pesce & Argano 1980 HLM86
         |--P. danubialis (Codreanu & Codreanu 1962) HLM86
         |--P. delhezi Henry & Magniez 1973 HLM86
         |--P. deminutus (Sket 1959) HLM86
         |    |--P. d. deminutus HLM86
         |    |--P. d. intermedius (Sket 1965) HLM86
         |    `--P. d. vulgaris (Sket 1965) HLM86
         |--P. elegans (Codreanu & Codreanu 1962) HLM86
         |--P. escolai Henry & Magniez 1982 HLM86
         |--P. espanoli Henry & Magniez 1982 HLM86
         |--P. exiguus Afonso 1983 HLM86
         |--P. franciscoloi (Chappuis 1955) HLM86
         |--P. gardinii (Arcangeli 1942) HLM86
         |--P. gauthieri (Monod 1924) HLM86
         |    |--P. g. gauthieri HLM86
         |    `--P. g. theodori (Birstein 1951) [incl. P. gauthieri numidicus Monod in Gauthier 1928 (n. n.)] HLM86
         |--P. gjorgjevici (Karaman 1933) HLM86
         |--P. gourbaultae Henry & Magniez 1981 HLM86
         |--P. hercegovinensis (Karaman 1933) HLM86
         |--P. hermallensis (Arcangeli 1938) HLM86
         |--P. hussoni (Remy 1941) HLM86
         |    |--P. h. hussoni HLM86
         |    `--P. h. curvifrons (Remy 1941) HLM86
         |--P. hypogeus (Racovitza 1922) HLM86
         |--P. infirmus (Birstein 1936) HLM86
         |--P. istrianus (Stammer 1932) HLM86
         |--P. jaloniacus Henry & Magniez 1977 HLM86
         |--P. karamani (Remy 1934) HLM86
         |--P. lagari Henry & Magniez 1982 HLM86
         |--P. lescherae Henry & Magniez 1977 HLM86
         |--P. linearis (Birstein 1967) HLM86
         |--P. ljovuschkini (Birstein 1967) HLM86
         |--P. lusitanicus (Frade 1938) HLM86
         |--P. lykaonicus Argano & Pesce 1978 HLM86
         |--P. maleri Henry 1977 HLM86
         |--P. margalefi Henry & Magniez 1982 HLM86
         |--P. mateusorum Afonso 1982 HLM86
         |--P. meridianus (Racovitza 1919) HLM86
         |    |--P. m. meridianus HLM86
         |    `--P. m. belgicus (Arcangeli 1935) HLM86
         |--P. minoicus Pesce & Argano 1980 HLM86
         |--P. monodi (Strouhal 1942) HLM86
         |--P. monsferratus (Braga 1948) HLM86
         |--P. montenigrinus (Karaman 1934) HLM86
         |    |--P. m. montenigrinus HLM86
         |    `--P. m. macedonicus (Karaman 1955) HLM86
         |--P. nolli (Karaman 1952) HLM86
         |--P. notenboomi Henry & Magniez 1982 HLM86
         |--P. parvulus (Sket 1960) HLM86
         |--P. patrizzi (Arcangeli 1952) HLM86
         |--P. pavani (Arcangeli 1942) HLM86
         |    |--P. p. pavani HLM86
         |    `--P. p. orientalis (Sket 1965) HLM86
         |--P. racovitzai Henry & Magniez 1972 HLM86
         |--P. rectangulatus Afonso 1982 HLM86
         |--P. rectus Afonso 1982 HLM86
         |--P. remyi [incl. P. remyi f. nudus (Karaman 1953)] HLM86
         |--P. rouchi Henry 1980 HLM86
         |--P. similis (Birstein 1967) HLM86
         |--P. sketi Henry 1975 HLM86
         |--P. slavus (Remy 1948) HLM86
         |    |--P. s. slavus HLM86
         |    |--P. s. histriae (Sket 1963) HLM86
         |    |--P. s. salisburgensis (Strouhal 1957) HLM86
         |    |--P. s. serbiae (Sket 1963) HLM86
         |    |--P. s. styriacus (Sket 1963) HLM86
         |    |--P. s. vindobonensis (Karaman 1955) HLM86
         |    `--P. s. zeii (Sket 1963) HLM86
         |--P. slovenicus (Sket 1957) HLM86
         |--P. solanasi Henry & Magniez 1972 HLM86
         |--P. spelaeus (Racovitza 1922) HLM86
         |--P. spinipes Afonso 1979 HLM86
         |--P. strouhali (Karaman 1955) HLM86
         |    |--P. s. strouhali HLM86
         |    `--P. s. puteanus (Henry 1966) HLM86
         |--P. synaselloides Henry 1963 HLM86
         |--P. thermonyctophilus (Monod 1924) HLM86
         |--P. valdensis (Chappuis 1948) HLM86
         |--P. vandeli Magniez & Henry 1969 HLM86
         |--P. variegatus Afonso 1982 HLM86
         |--P. vignai Argano & Pesce 1977 HLM86
         `--P. walteri (Chappuis 1948) HLM86

*Type species of generic name indicated

REFERENCES

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[HLM86] Henry, J.-P., J. J. Lewis & G. Magniez. 1986. Isopoda: Asellota: Aselloidea, Gnathostenetroidoidea, Stenetrioidea. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 434–464. E. J. Brill/Dr. W. Backhuys: Leiden.

[N29] Nicholls, G. E. 1929. A new species of Asellus (A. nipponensis) from Japan. Archives de Zoologie Expérimentale et Générale 68 (1): 33–38.

[T30] Tattersall, W. M. 1930. Asellus cavaticus, a blind isopod new to the British fauna, from a well in Hampshire. Proceedings of the Linnean Society of London 141: 7.

Crinocheta

Nagurus cristatus, copyright Arp Kruihof.

Belongs within: Oniscidea.
Contains: Oniscidae, Philosciidae, Armadillidae, Porcellio.

The Crinocheta are a group of terrestrial isopods characterised by an 'embolophore' type of male genitalia with the two vasa deferentia opening by separate pores on either side of the penis towards its apex (Hatch 1947, as Embolophora). They are divided between the Armadilloidea, in which the pleopods carry respiratory pseudotracheae, and the Oniscoidea in which such structures are absent.

<==Crinocheta [Embolophora] MD01
    |--Oniscoidea MD01
    |    |  i. s.: Tasmanoniscus Vandel 1973 V73
    |    |           `--*T. evansi Vandel 1973 V73
    |    |--Berytoniscidae MD01
    |    |--Olibrinus Budde-Lund 1913 JP13 [Olibrinidae MD01]
    |    |--Pudeoniscidae MD01
    |    |--Speleoniscidae MD01
    |    |--Sphaeroniscidae MD01
    |    |--Stenoniscidae MD01
    |    |--Tendosphaeridae MD01
    |    |--Oniscidae MD01
    |    |--Philosciidae MD01
    |    |--Halophiloscia PP64 [Halophilosciidae MD01]
    |    |    `--H. couchii PP64
    |    |--Australoniscus Vandel 1973 V73 [Bathytropidae MD01]
    |    |    `--*A. springetti Vandel 1973 V73
    |    |--Deto V73 [Detonidae MD01]
    |    |    `--D. marina (Chilton 1884) V73
    |    |--Rhyscotus M90 [Rhyscotidae MD01]
    |    |    `--R. texensis M90
    |    |--Platyarthrus M90 [Platyarthridae MD01]
    |    |    |--P. hoffmannseggi M90
    |    |    `--P. schoebli M90
    |    `--Scyphacidae MD01
    |         |--Scyphacella arenicola M90
    |         |--Detonella [Scyphacinae] H47
    |         |    |--D. papillicornis [=Trichoniscus papillicornis; incl. D. lohmanderi] H47
    |         |    `--D. sachalina H47
    |         `--Armadilloniscus [Armadilloniscinae] H47
    |              |--A. ellipticus M90
    |              |--A. holmesi M90
    |              |--A. lindahli M90
    |              `--A. tuberculatus [=Actoniscus tuberculatus] H47
    `--Armadilloidea MD01
         |--Atlantididae MD01
         |--Balloniscidae MD01
         |--Eubelidae MD01
         |--Periscyphicidae MD01
         |--Armadillidae MD01
         |--Cylisticus M90 [Cylisticidae MD01]
         |    |--C. convexus M90 [incl. C. convexus var. confluentus H47, Porcellio glaber H47]
         |    `--C. rotabilis D01
         |--Armadillidium M90 [Armadillidiidae MD01]
         |    |--A. (Armadillidium) vulgare [incl. Armadillo pilularis Say 1818] H47
         |    `--A. (Pseudosphaerium) nasatum H47
         |--Actaecia Dana 1853 V73 [incl. Cylloma Budde-Lund 1885 V73; Actaeciidae MD01]
         |    |--A. euchroa Chilton 1901 NB26
         |    |--A. opihensis Chilton 1901 NB26
         |    `--A. pallida Nicholls & Barnes 1926 V73
         |--Porcellionidae MD01
         |    |--Porcellio S77
         |    |--Agabiformius lentus (Budde-Lund 1885) V73
         |    `--Porcellionides Miers 1877 JP13
         |         `--P. pruinosus (Brandt 1833) JP13 [=Metoponorthus pruinosus M90; incl. P. immaculatus H47]
         `--Trachelopodidae [Trachelipidae] MD01
              |--Trachelipus [incl. Tracheoniscus] M90
              |    `--T. rathkei M90 (see below for synonymy)
              `--Nagurus Holthuis 1949 [incl. Nagara Budde-Lund 1908 (preoc.), Nagaroides Wahrberg 1922] V73
                   |--N. aegaeus Schmalfuss 1977 S77
                   |--N. carinatus [incl. N. hermonensis Vandel 1955] S77
                   |--N. cristatus (Dollfus 1889) V73
                   |--N. declivus (Wahrberg 1922) V73
                   |--N. gracillimus (Wahrberg 1922) V73
                   |--N. longiflagellatus (Wahrberg 1922) V73
                   |--N. rhodiensis S77
                   `--N. tumidus (Wahrberg 1922) V73

Nomen nudum: Australoniscus alticolus Vandel 1973 V73

Trachelipus rathkei M90 [incl. Porcellio dorsalis H47, P. lateralis H47, P. limatus H47, P. limbalis H47, P. marginatus H47, P. mixtus H47, P. multiguttatus H47, T. rathkei var. ochraceus H47, Porcellio variegatus H47, T. rathkei var. varius H47]

*Type species of generic name indicated

REFERENCES

[D01] Dollfus, A. 1901. Isopodák [Isopodes]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 371–374. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[JP13] Judd, S., & G. Perina. 2013. An illustrated key to the morphospecies of terrestrial isopods (Crustacea: Oniscidea) of Barrow Island, Western Australia. Records of the Western Australian Museum Supplement 83: 185–207.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1–124.

[M90] Muchmore, W. B. 1990. Terrestrial Isopoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 805–817. John Wiley & Sones: New York.

[NB26] Nicholls, G. E., & H. M. Barnes. 1926. A description of two new terrestrial isopods from Western Australia. Journal of the Royal Society of Western Australia 12 (17): 149–159.

[PP64] Peres, J. M., & J. Picard. 1964. Nouveau manuel de bionomie benthique de la mer Mediterranee. Recueil des Travaux de la Station Marine d'Endoume, Bulletin 31 (27): 5–137.

[S77] Schmalfuss, H. 1977. Eine neue Nagurus-Art aus der Süd-Agäis (Crustacea: Isopoda: Oniscoidea: Trachelipidae). Senckenbergiana Biologica 57 (4–6): 359–365.

[V73] Vandel, A. 1973. Les isopodes terrestres de l'Australie. Étude systématique et biogéographique. Mémoires du Muséum National d'Histoire Naturelle, Série A, Zoologie 82: 1–171.

Last updated: 29 October 2017.

Mesidoteinae

Austridotea annectens, copyright Paddy Ryan.

Belongs within: Valvifera.

The Mesidoteinae are a group of relatively unspecialised valviferans exhibiting a benthic lifestyle (Nicholls 1937).

Characters (from Nicholls 1937): Body flattened and ovate; eyes small; distinct coxal plates present on six of the seven free mesosomatic segments; three or four pleonites present of at least I and II are free.

<==Mesidoteinae
    |--Mesidotea N37
    |    |--M. entomon N37
    |    |--M. sabini N37
    |    `--M. sibirica N37
    `--Austridotea Nicholls 1937 N37
         |--A. (Austridotea) N37
         |    |--A. (A.) annectens Nicholls 1937 N37
         |    `--A. (A.) benhami Nicholls 1937 N37
         `--A. (Notidotea Nicholls 1937) N37
              |--A. (N.) lacustris (Thomson 1879) [=Idotea lacustris, Pentidotea lacustris] N37
              `--A. (N.) rotundicauda (Miers 1881) [=Idotea rotundicauda] N37

*Type species of generic name indicated

REFERENCES

[N37] Nicholls, G. E. 1937. On the freshwater Idoteidae of New Zealand (Crustacea, Isopoda). Annals and Magazine of Natural History, series 10, 19: 113–136.