Dorsal view of Carabodes dubius, from Murvanidze (2008).

Belongs within: Circumdehiscentiae.
Contains: Odontocepheus (Odontocepheus), Diplobodes, Yoshiobodes, Gibbicepheus (Gibbicepheus), Austrocarabodes (Austrocarabodes), Carabodes (Carabodes), Carabodes (Klappariches).

The Carabodidae are a family of oribatid mites with more or less parallel lamellae on the prodorsum (Balogh & Balogh 1992). Though diverse in adult biology, all species are probably endophagous as nymphs, burrowing in substrates such as decaying wood. Adults of many species are arboreal, feeding on fungi and lichens (Norton & Behan-Pelletier 2009). In a number of genera the notogaster bears longitudinal rows of large tubercles, such as in Gibbicepheus or Diplobodes. The genera Spathulocepheus and Trichocarabodes are distinctive in the high number of setae on their genital plates: eight pairs in Trichocarabodes, ten in Spathulocepheus. Other genera have at most seven pairs of genital setae (Balogh & Balogh 1992).

Characters (from Balogh & Balogh 1992): Prodorsum with true lamellae; lamellae marginal, almost parallel or weakly converging; interlamellar setae well developed. Notogaster usually pentagonal, with more or less parallel sides and large humeral tubercle.

    |--Gibbicepheus Aoki 1958 S04 [Gibbicepheinae CH98]
    |    |--G. (Gibbicepheus) S04
    |    `--G. (Gibbibodes Mahunka 1986) S04
    |         `--G. (*G.) similis (Mahunka 1986) [=*Gibbibodes similis] S04
    `--Carabodinae CH98
         |--Austrocarabodes Hammer 1966 CH98
         |    |--A. (Austrocarabodes) S04
         |    |--A. (Baloghodes Mahunka 1986) S04
         |    |    |--A. (*B.) heterotrichosus (Mahunka 1986) [=*Baloghodes heterotrichosus] S04
         |    |    `--A. (B.) secundus (Mahunka 1993) [=Baloghodes secundus] S04
         |    `--A. (Uluguroides Mahunka 1983) [incl. Ngorongobodes Balogh & Balogh 1992] S04
         |         |--A. (*U.) trichosus (Mahunka 1983) [=*Uluguroides trichosus] S04
         |         |--A. (U.) grandis (Mahunka 1984) [=Ulugurozetes grandis] S04
         |         |--A. (U.) incrustatus (Wallwork 1977) [=Carabodes incrustatus] S04
         |         |--A. (U.) minitricha (Mahunka 1993) [=Uluguroides minitricha] S04
         |         `--A. (U.) pentatrichus (Balogh 1962) S04 (see below for synonymy)
         |              |--A. p. pentatrichus S04
         |              `--A. p. ‘clavatus’ (Wallwork 1977) (see below for synonymy) S04
         `--Carabodes Koch 1835 S04
              |  i. s.: C. elongatus S22
              |--C. (Carabodes) S04
              |--C. (Flexa Kulijev 1977) S04
              |    |--C. (*F.) dubius Kulijev 1968 S04
              |    |--C. (F.) bidens (Djaparidze 1990) [=Flexa bidens] S04
              |    |--C. (F.) breviclava Aoki 1970 S04
              |    |--C. (F.) horeo (Djaparidze 1990) [=Flexa horeo] S04
              |    |--C. (F.) intermedius Willmann 1951 S04
              |    `--C. (F.) scopulae Kulijev 1968 S04
              |--C. (Klapperiches) S04
              `--C. (Phyllocarabodes Balogh & Mahunka 1969) (see below for synonymy) S04
                   |--C. (*P.) octogonalis (Balogh & Mahunka 1969) [=*Phyllocarabodes octogonalis] S04
                   |--C. (P.) calcaratus (Mahunka 1985) [=Antillobodes calcaratus] S04
                   |--C. (P.) inopinatus (Mahunka 1985) [=Antillobodes inopinatus] S04
                   |--C. (P.) insolitus (Balogh 1984) S04 [=*Pentabodes insolitus BB92]
                   `--C. (P.) ‘ornatus’ (Balogh 1986) non C. ornatus Storkan 1925 [=Phyllocarabodes ornatus] S04

Carabodidae incertae sedis:
  Apotomocepheus Aoki 1965 S04
    `--*A. gressitti Aoki 1965 S04
  Meriocepheus Aoki 1973 S04
    `--*M. peregrinus Aoki 1973 S04
  Sagittabodes Balogh & Balogh 1992 S04
    `--*S. sagitta (Balogh & Mahunka 1966) [=Machadocepheus sagitta] S04
  Tansocepheus Mahunka 1983 S04
    `--*T. serratus Mahunka 1983 S04
  Tectocarabodes Mahunka 1988 S04
    `--*T. monstruosus Mahunka 1988 S04
  Tuberocepheus Balogh & Mahunka 1969 S04
    `--*T. longus (Balogh 1964) [=Machadocepheus longus] S04
  Singabodes Mahunka 1998 S04
    `--*S. rarus Mahunka 1998 S04
  Archegocepheus Aoki 1965 S04
    |--*A. imadatei Aoki 1965 S04
    `--A. macrofoliatus Wen 1997 S04
  Bathocepheus Aoki 1978 [incl. Philippobodes Balogh & Balogh 1992] S04
    |--*B. concavus Aoki 1978 S04
    `--B. manguiati (Corpuz-Raros 1979) S04 [=Machadocepheus manguiati S04, *Philippobodes manguiati BB92]
  Cavernocarabodes Mahunka 1974 S04
    |--*C. perreti Mahunka 1974 S04
    `--C. trigonosternum (Pérez-Íñigo 1976) [=Carabodes trigonosternum] S04
  Spathulocepheus Balogh & Mahunka 1969 S04
    |--*S. amazonicus Balogh & Mahunka 1969 S04
    `--S. plumiger Balogh & Mahunka 1977 S04
  Malgasodes Mahunka 2000 S04
    |--*M. curvisetus Mahunka 2000 S04
    `--M. hungarorum Mahunka 2000 S04
  Gymnobodes Balogh 1965 S04
    |--*G. fraterculus (Balogh 1963) [=Carabodes fraterculus] S04
    |--G. semengok Mahunka 1996 S04
    `--G. subnudus (Balogh 1963) [=Carabodes subnudus] S04
  Pseudocarabodes Mahunka 1991 [incl. Bakobodes Mahunka 1996, Yemenobodes Mahunka 2000] S04
    |--*P. xenus Mahunka 1991 S04
    |--P. improvisus (Mahunka 2000) [=Yemenobodes improvisus] S04
    `--P. orangutan (Mahunka 1996) [=Bakobodes orangutan] S04
  Trichocarabodes Balogh 1961 [incl. Ceylobodes Balogh & Balogh 1992] S04
    |--*T. celisi (Balogh 1958) [=Carabodes celisi] S04
    |--T. capillatus Balogh 1970 S04 [=*Ceylobodes capillatus BB92]
    |--T. costulatus Mahunka 1984 S04
    `--T. hettigei Balogh 1970 S04
  Hardybodes Balogh 1970 [incl. Carabodella Mahunka 1986] S04
    |--*H. mirabilis Balogh 1970 S04
    |--H. calcaratus (Mahunka 1986) S04 [=*Carabodella calcarata BB92]
    |--H. flabellatus Mahunka 1995 S04
    `--H. penicillatus Mahunka 1995 S04
  Cubabodes Balogh & Mahunka 1974 S04
    |--*C. hexagonalis Balogh & Mahunka 1974 S04
    |--C. confertus Balogh & Mahunka 1980 S04
    |--C. radiatus Balogh & Mahunka 1974 S04
    |--C. spathulatus Balogh & Mahunka 1980 S04
    `--C. verrucatus Balogh & Mahunka 1980 S04
  Machadocepheus Balogh 1958 S04
    |--*M. excavatus Balogh 1958 S04
    |--M. filiferus Mahunka 1986 S04
    |--M. pauliani Balogh 1962 S04
    |--M. phyllophorus Balogh 1970 S04
    |--M. taprobanicus Balogh 1970 S04
    `--M. tuberculosus Mahunka 1987 S04
  Odontocepheus Berlese 1913 S04
    |--O. (Odontocepheus) S04
    |--O. (Aokiella Balogh & Mahunka 1967) S04
    |    |--O. (*A.) florens (Balogh & Mahunka 1967) [=*Aokiella florens] S04
    |    `--O. (A.) rotundus (Hammer 1979) [=Aokiella rotunda] S04
    `--O. (Indotocepheus Mondal & Kundu 1999) S04
         `--O. (*I.) himalayensis (Mondal & Kundu 1999) [=*Indotocepheus himalayensis] S04
  Pasocepheus Aoki 1976 S04
    |--P. (Pasocepheus) [incl. Opisthocepheus Aoki 1976] S04
    |    |--*P. (P.) triarcuatus Aoki 1976 S04
    |    |--P. (P.) bako Mahunka 1996 S04
    |    |--P. (P.) eremaeozetoides Mahunka 1995 S04
    |    `--P. (P.) kirai (Aoki 1976) S04 [=*Opisthocepheus kirai BB92]
    `--P. (Guineobodes Mahunka 1987) S04
         `--P. (*G.) papuanus (Balogh 1970) [=Machadocepheus papuanus] S04
  Congocepheus Balogh 1958 S04
    |--*C. heterotrichus Balogh 1958 S04
    |--C. hauseri Mahunka 1989 S04
    |--C. involutus Mahunka 1997 S04
    |--C. latilamellatus Mahunka 1984 S04
    |--C. orientalis Mahunka 1989 S04
    |--C. ornatus Mahunka 1983 S04
    |--C. taurus Balogh 1961 S04
    `--C. velatus Mahunka 1986 S04
  Diplobodes S04
  Yoshiobodes S04

Austrocarabodes (Uluguroides) pentatrichus (Balogh 1962) S04 [=Carabodes pentatrichus S04, *Ngorongobodes pentatrichus BB92]

Austrocarabodes (Uluguroides) pentatrichus ‘clavatus’ (Wallwork 1977) [=Carabodes pentatrichus clavatus nec C. labyrinthicus clavatus Warburton 1912 nec C. gibbiceps clavatus Jacot 1938] S04

Carabodes (Phyllocarabodes Balogh & Mahunka 1969) [incl. Antillobodes Mahunka 1985, Pentabodes Balogh 1984] S04

*Type species of generic name indicated


[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

Murvanidze, M. 2008. Checklist and key to species of Carabodes (Acari, Oribatida, Carabodidae) of the Caucasian region, with description of a new species. Acarina 16 (2): 177–186.

[S22] Schweizer, J. 1922. Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandl. Naturf. Ges. Basel 33: 23–112, 4 pls.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

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