Madras harp Harpa davidus, copyright H. Zell.

Belongs within: Siphonogastropoda.

The Harpidae, harp shells, are a group of predatory gastropods with usually colourfully patterned, polished shells bearing strong axial ribs.

Characters (from Wilson & Gillett 1971): Shell usually shiny, strikingly coloured and patterned, sculptured with strong axial ribs more highly polished than remainder of shell. Anterior siphon canal a deep, wide notch; columella lacking folds; operculum absent. Respiratory siphon and eye stalks long; foos broad with front end greatly expanded laterally to form wide, flat, muscular organ.

    |--Cryptochorda Mörch 1858 [Cryptochordidae, Cryptochordinae] BR05
    |--Eocithara Fischer 1883 WG71
    |--Austroharpa Finlay 1931 WG71
    |    |--A. exquisita Iredale 1931 WG71
    |    |--A. punctata Verco 1906 WG71
    |    `--A. wilsoni Rehder 1973 MG-H11
    |--Morum Röding 1798 [Moruminae] BR05
    |    |  i. s.: M. vicdani Emerson 1995 BC01
    |    `--M. (Oniscidia Mörch 1852) [incl. Cancellomorum Emerson & Old 1963] BC01
    |         |--M. (O.) joelgreenei Emerson 1981 BC01
    |         |--M. (O.) macdonaldi Emerson 1981 BC01
    |         |--M. (O.) matthewsi Emerson 1967 [=M. (Cancellomorum) matthewsi] BC01
    |         |--M. (O.) ninomiyai Emerson 1986 BC01
    |         `--M. (O.) veleroae Emerson 1968 [=M. (Cancellomorum) veleroae] BC01
    `--Harpa Röding 1798 [Harpinae] BR05
         |--*H. harpa Linnaeus 1758 [incl. H. nobilis Röding 1798] WG71
         |--H. amouretta Röding 1798 [incl. H. minor Lamarck 1816] WG71
         |--H. articularis Lamarck 1822 WG71
         |--H. davidus Röding 1798 WG71
         |--H. goodwini Rehder 1993 BC01
         |--H. gracilis Broderip & Sowerby 1829 WG71
         |--H. kajiyamai Habe & Kosuge 1970 PK11
         |--H. major Röding 1798 WG71
         |--H. neozelanica [=Eocithara neozelanica] A27
         `--H. ventricosa H04

*Type species of generic name indicated


[A27] Allan, R. S. 1927. The geology and palaeontology of the Lower Waihao Basin, south Canterbury, New Zealand. Transactions and Proceedings of the New Zealand Institute 57: 265–309.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[H04] Haeckel, E. 1899–1904. Kunstformen der Natur. Bibliographisches Institut: Leipzig und Wien.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[PK11] Puillandre, N., Y. I. Kantor, A. Sysoev, A. Couloux, C. Meyer, T. Rawlings, J. A. Todd & P. Bouchet. 2011. The dragon tamed? A molecular phylogeny of the Conoidea (Gastropoda). Journal of Molluscan Studies 77: 259–272.

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.

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