Melyridae

Malachius bipustulatus, copyright Frederik81.


Belongs within: Cleroidea.
Contains: Dasytes, Laius, Hypattalus, Carphurus, Helcogaster.

The Melyridae are a group of relatively soft-bodied beetles that are usually predacious or scavengers as larvae but commonly flower-feeders as adults (Lawrence & Britton 1991).

Characters (from Lawrence & Britton 1991): Oblong to elongate, somewhat flattened, soft-bodied, usually clothed with decumbent hairs and scattered erect setae, either uniformly black or grey or bicoloured with red or yellow and blue or black . Labrum well sclerotised; tarsal claws toothed or provided with a pair of fleshy appendages; elytra sometimes very short, exposing several abdominal tergites; eversible vesicles often on prothorax, mesothorax and abdomen. Larvae elongate, slightly flattened, lightly sclerotised, except for head, T9 and thoracic plates, red or pink in colour; head with long epicranial stem, no median endocarina, and lightly sclerotised gular and para-gular areas, so that head capsule appears to be open behind ventral mouthparts; mesal appendage at base of mandible.

Melyridae [Malachiidae]
    |  i. s.: Simoderus MC13
    |         Listrus MW15
    |--Dasytinae [Dasytidae] B14
    |    |--Enicopus (Parahenicopus) lopezcoloni Bahillo de la Puebla 2005 F05
    |    |--Trichochrous BM76
    |    |--Amecocerus cervicalis BM76
    |    `--Dasytes L09
    |--Melyrinae B14
    |    |--Astylus [Astylini] B14
    |    |    `--A. atromaculatus (Blanchard 1843) B14
    |    |--Melyrodes B14
    |    |    |--M. basalis (LeConte 1852) [=Dasytes basalis] B14
    |    |    |--M. cribatus B14
    |    |    `--M. floridiana B14
    |    `--Melyris G89
    |         |--M. bicolor G89
    |         |--M. oblonga B14
    |         |--M. rubripes Lucas 1847 E12
    |         `--M. viridis L02
    `--Malachiinae [Malachiides] B14
         |--Laius L09
         |--Hypattalus L09
         |--Carphurus L09
         |--Helcogaster L09
         |--Flabellolaius LB91
         |--Dicranolaius LB91
         |    |--D. bellulus LB91
         |    `--D. villosus LB91
         |--Collops B14
         |    |--C. balteatus LeConte 1852 B14
         |    |--C. punctulatus B14
         |    |--C. quadrimaculatus B14
         |    `--C. versatilis B14
         |--Neocarphurus Lea 1898 L09
         |    |--N. basizonis Lea 1901 L09
         |    |--N. chlorops Lea 1898 L09
         |    |--N. coatesi Lea 1909 L09
         |    |--N. impunctatus (Lea 1895) [=Carphurus impunctatus] L09
         |    |--N. sobrinus Lea 1909 L09
         |    `--N. tuberculatus (Lea 1895) [=Helcogaster tuberculatus] L09
         |--Malachius E12
         |    |--M. aeneus (Linnaeus 1758) B14
         |    |--M. affinis Lucas 1847 E12
         |    |--M. angusticollis Lucas 1847 E12
         |    |--M. bipustulatus C01
         |    |--M. maculicollis Lucas 1847 E12
         |    |--M. marginicollis Lucas 1847 E12
         |    |--M. mauritanicus Lucas 1847 E12
         |    |--M. tristis Lucas 1847 E12
         |    `--M. viridis V09
         `--Balanophorus non Briganti 1825 L09
              |--B. ater Lea 1909 L09
              |--B. biplagiatus Fairm. 1879 [=Carphurus biplagiatus] L09
              |--B. brevipennis (Germ. 1848) [=Carphurus brevipennis] L09
              |--B. janthinipennis Fairm. 1879 L09
              |--B. macleayi Lea 1895 L09
              |--B. mastersi Macleay 1872 M86
              |--B. megalops Lea 1901 L09
              |--B. pictus Lea 1909 L09
              |--B. scapulatus (Fairm. 1879) [=Carphurus scapulatus] L09
              `--B. victoriensis Lea 1909 L09

*Type species of generic name indicated

REFERENCES

[BM76] Bohart, R. M. & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] Band II. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[L09] Lea, A. M. 1909. Revision of the Australian and Tasmanian Malacodermidae. Trans. Ent. Soc. Lond. 1909 (1): 45–251, pls 2–6.

[MC13] Majer, J. D., S. K. Callan, K. Edwards, N. R. Gunawardene & C. K. Taylor. 2013. Baseline survey of the terrestrial invertebrate fauna of Barrow Island. Records of the Western Australian Museum Supplement 83: 13–112.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part IV. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 259–380.

[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

[V09] Verdcourt, B. (ed.) 2009. Additions to the Wild Fauna and Flora of the Royal Botanic Gardens, Kew. XXVI. Miscellaneous records. Kew Bulletin 64 (1): 183–194.

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