Recifella

Dorsum and palp of female Recifella excavata, from Rosso de Ferradás & Böttger (1997).

Belongs within: Unionicolidae.

Recifella is a genus of unionicolid water mites found in Australia and South America, with complete dorsal and ventral shields and three or four pairs of glandularia on the dorsal shield.

<==Recifella Viets 1935 RFF05
    |--R. (Recifella) RFF05
    |    |--R. (R.) angulipes (Lundblad 1938) RFF05
    |    |--R. (R.) clavata (Lundblad 1937) RFF05
    |    |--R. (R.) gibberipalpis (Lundblad 1936) RFF05
    |    |--R. (R.) laminipes (Viets 1935) RFF05
    |    |--R. (R.) latipes (Lundblad 1936) RFF05
    |    |--R. (R.) lundbladi (Besseling 1949) RFF05
    |    `--R. (R.) sinuatipes (Lundblad 1941) RFF05
    |--R. (Eorecifella Cook 1980) RFF05
    |    |--R. (E.) balteata (Lundblad 1943) RFF05
    |    |--R. (E.) cruciata (Lundblad 1936) RFF05
    |    |    |--R. c. cruciata RFF05
    |    |    `--R. c. brevis (Lundblad 1943) RFF05
    |    |--R. (E.) diplothoracica (Lundblad 1937) RFF05
    |    |--R. (E.) elliptica (Walter 1919) RFF05
    |    |    |--R. e. elliptica RFF05
    |    |    |--R. e. amabilis (Lundblad 1943) RFF05
    |    |    |--R. e. schadei (Lundblad 1943) RFF05
    |    |    `--R. e. validipalpis (Lundblad 1937) RFF05
    |    |--R. (E.) excavata (Lundblad 1936) RFF05
    |    |--R. (E.) excavatoides (Viets 1975) RFF05
    |    |--R. (E.) opimipalpis (Viets 1977) RFF05
    |    |--R. (E.) pusilla (Lundblad 1937) RFF05
    |    `--R. (E.) undulata (Lunblad 1936) RFF05
    `--R. (Monokoenikea Lundblad 1941) RFF05
         `--R. (M.) melini (Lundblad 1930) RFF05

Recifella incertae sedis:
  R. agnosta Cook 1986 H98
  R. australica Cook 1986 H98
  R. baltoona Cook 1986 H98
  R. bifida Cook 1986 H98
  R. columbiana (Lundblad 1953) RFF05
  R. doomba Cook 1986 H98
  R. flagellata Cook 1986 H98
  R. inka (Lundblad 1944) RFF05
  R. paucipora (Lundblad 1943) RFF05
  R. pectinatus Smit 1992 H98
  R. plana Cook 1986 H98
  R. pulchra (Lundblad 1937) RFF05
  R. szalayi Cook 1986 H98
  R. tinka (Viets 1975) [=Koenikea tinka] H98
  R. triradiata Cook 1986 H98
  R. truncata (Besseling 1949) RFF05
  R. umala Cook 1986 H98
  R. vonjama Cook 1986 H98

*Type species of generic name indicated

REFERENCES

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

Koenikea

Koenikea sp., copyright David E. Reed.

Belongs within: Unionicolidae.

Koenikea is a genus of unionicolid water mites found in Australia and the Americas, with complete dorsal and ventral shields and five or six pairs of glandularia on the dorsal shield.

<==Koenikea Wolcott 1900 RFF05
    |--K. (Koenikea) V77
    |    |--K. (K.) acuta Lundblad 1943 RFF05
    |    |    |--K. a. acuta RFF05
    |    |    `--K. a. dentata Viets 1975 RFF05
    |    |--K. (K.) affinis Lundblad 1938 RFF05
    |    |--K. (K.) angustipalpis Viets 1975 RFF05
    |    |--K. (K.) bicornis Lundblad 1941 RFF05
    |    |--K. (K.) bipapillata Lundblad 1941 RFF05
    |    |--K. (K.) biscutata Daday 1905 RFF05
    |    |--K. (K.) brunnea Lundblad 1937 RFF05
    |    |--K. (K.) coerulescens Lundblad 1936 RFF05
    |    |    |--K. c. coerulescens RFF05
    |    |    |--K. c. rosea Lundblad 1936 RFF05
    |    |    `--K. c. vietsi Lundblad 1937 RFF05
    |    |--K. (K.) convexa Daday 1905 (n. d.) RFF05
    |    |--K. (K.) corniculata Viets 1975 RFF05
    |    |--K. (K.) falcaria Viets 1977 V77
    |    |--K. (K.) fluctuata Viets 1975 RFF05
    |    |--K. (K.) furcatimaculata Lundblad 1936 RFF05
    |    |--K. (K.) hastata Viets 1977 V77
    |    |--K. (K.) lata Viets 1975 RFF05
    |    |--K. (K.) longipes Lundblad 1936 RFF05
    |    |--K. (K.) medioposita Lundblad 1936 RFF05
    |    |    |--K. m. medioposita V77
    |    |    |--K. m. asterifera Lundblad 1943 V77
    |    |    `--K. m. transversa Lundblad 1943 V77
    |    |--K. (K.) octopapillata Lundblad 1930 RFF05
    |    |--K. (K.) opimipalpis Viets 1977 V77
    |    |--K. (K.) pallida Lundblad 1937 RFF05
    |    |--K. (K.) papillata Viets 1975 RFF05
    |    |--K. (K.) paraguayensis Rosso de Ferradás & Böttger 1998 RFF05
    |    |--K. (K.) plicata Lundblad 1936 RFF05
    |    |--K. (K.) procursa Viets 1977 V77
    |    |--K. (K.) quadricornuta Walter 1919 RFF05
    |    |--K. (K.) rectangulata Lundblad 1930 RFF05
    |    |--K. (K.) rectirostris Lundblad 1936 RFF05
    |    |    |--K. r. rectirostris RFF05
    |    |    |--K. r. crassoalata Lundblad 1943 RFF05
    |    |    |--K. r. ramosa Lundblad 1943 RFF05
    |    |    `--K. r. retractidens Lundblad 1938 RFF05
    |    |--K. (K.) retrocornuta Rosso de Ferradás & Böttger 1997 RFF05
    |    |--K. (K.) siolii Viets 1975 RFF05
    |    |--K. (K.) triangularis Lundblad 1938 RFF05
    |    |--K. (K.) unguiculata Lundblad 1943 RFF05
    |    `--K. (K.) victoriola Rosso de Ferradás 1998 RFF05
    |--K. (Diplokoenikea Lundblad 1936) RFF05
    |    |--K. (D.) clavigera Lundblad 1943 RFF05
    |    |    |--K. c. clavigera RFF05
    |    |    `--K. c. assimilis Lundblad 1943 RFF05
    |    |--K. (D.) curvirostris Walter 1919 RFF05
    |    |--K. (D.) flagellaris Lundblad 1938 RFF05
    |    |--K. (D.) grossa Lundblad 1938 RFF05
    |    |--K. (D.) pectinata Lundblad 1936 RFF05
    |    |--K. (D.) pectinifera Lundblad 1938 RFF05
    |    |--K. (D.) quinquemaculata Lundblad 1943 RFF05
    |    |--K. (D.) scutica Viets 1977 V77
    |    `--K. (D.) simulans Lundblad 1943 RFF05
    |--K. (Jurucuia Viets 1954) RFF05
    |    `--K. (J.) oblongata (Viets 1954) RFF05
    |--K. (Koenikella Lundblad 1936) RFF05
    |    |--K. (K.) crassipalpis Lundblad 1936 RFF05
    |    `--K. (K.) cyanozona Lundblad 1937 RFF05
    |--K. (Neokoenikea Lundblad 1936) RFF05
    |    `--K. (N.) armipes Lundblad 1936 RFF05
    |--K. (Notomideopsis Wolcott 1905) RFF05
    |    |--K. (N.) acanthifera Lundblad 1953 RFF05
    |    |--K. (N.) acutidens Lundblad 1943 RFF05
    |    |--K. (N.) brasiliensis Lundblad 1930 RFF05
    |    |--K. (N.) chilensis Cook 1988 RFF05
    |    |--K. (N.) circumcincta Lundblad 1937 RFF05
    |    |--K. (N.) consimilis Viets 1975 RFF05
    |    |--K. (N.) crassa Lundblad 1937 RFF05
    |    |--K. (N.) curicola Viets 1975 RFF05
    |    |--K. (N.) curvipes Lundblad 1938 RFF05
    |    |--K. (N.) diabolica Viets 1975 RFF05
    |    |--K. (N.) elegans Lundblad 1938 RFF05
    |    |--K. (N.) hamulata Lundblad 1936 RFF05
    |    |--K. (N.) imitata Lundblad 1930 RFF05
    |    |--K. (N.) lunipes Lundblad 1938 RFF05
    |    |--K. (N.) nordenskioldi Cook 1980 RFF05
    |    |--K. (N.) obscura Lundblad 1937 RFF05
    |    |--K. (N.) rectipes Lundblad 1943 RFF05
    |    |--K. (N.) soona Cook 1988 RFF05
    |    |--K. (N.) spinosa Daday 1905 RFF05
    |    `--K. (N.) tigrina Lundblad 1936 RFF05
    |         |--K. t. tigrina RFF05
    |         `--K. t. crassiseta Lundblad 1943 RFF05
    |--K. (Parakoenikea Lundblad 1936) RFF05
    |    |--K. (P.) curvipalpis Lundblad 1936 RFF05
    |    `--K. (P.) scutellum Besch 1969 RFF05
    |--K. (Pseudokoenikea Lundblad 1941) RFF05
    |    |--K. (P.) horrida Lundblad 1930 RFF05
    |    `--K. (P.) rutae Lundblad 1930 RFF05
    |         |--K. r. rutae RFF05
    |         `--K. r. furcaticornis Viets 1956 RFF05
    |--K. (Sespekoenikea Cook 1974) S86
    |    `--K. (S.) expansipalpis Cook 1974 S86
    `--K. (Tamboella Lundblad 1953) RFF05
         `--K. (T.) multiseta Lundblad 1953 RFF05

Koenikea incertae sedis:
  K. australica Lundblad 1941 H98
  K. balteata Lundblad 1943 V77
  K. bispina WL09
  K. branacha Cook 1986 H98
  K. colorata Viets 1980 [=Recifella colorata] H98
  K. crinita Cook 1986 H98
  K. curtiseta Cook 1986 H98
  K. decapora Cook 1986 H98
  K. distans Viets 1980 H98
  K. elliptica Walter 1919 V77
    |--K. e. elliptica V77
    |--K. e. amabilis Lundblad 1943 V77
    |--K. e. schadei Lundblad 1943 V77
    `--K. e. validipalpis Lundblad 1937 V77
  K. haldemani PW01
  K. jacunda Cook 1986 H98
  K. lemba Cook 1986 H98
  K. longiuscula Lundblad 1936 RFF05
  K. pauciseta Cook 1980 RFF05
  K. piota Cook 1986 H98
  K. plaumanni Lundblad 1937 RFF05
  K. pseudodistans Cook 1986 H98
  K. pulchra Lundblad 1937 V77
  K. purpurea Smit 1992 H98
  K. rodosa Cook 1986 H98
  K. rutala Cook 1986 H98
  K. schubarti Viets 1977 V77
  K. sorpresa Cook 1986 H98
  K. spinipes PW01
  K. timmsi Viets 1977 H98
  K. verrucosa Lundblad 1947 H98
  K. voloma Cook 1986 H98

*Type species of generic name indicated

REFERENCES

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[PW01] Proctor, H., & K. Wilkinson. 2001. Coercion and deceit: water mites (Acari: Hydracarina) and the study of intersexual conflict. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10^th International Congress pp. 155–169. CSIRO Publishing: Melbourne.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

[S86] Schwoerbel, J. 1986. Acari: “Hydrachnellae”. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 652–696. E. J. Brill/Dr W. Backhuys: Leiden.

[V77] Viets, K. O. 1977. Neue Koenikea-Arten aus Brasilien (Acari: Hydrachnellae: Unionicolidae). Senckenbergiana Biologica 57 (4-6): 341–358.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

Piona

Male Piona nodata, from here.

Belongs within: Pionidae.

Piona is a genus of pionid water mites in which males bear a dorsal concavity on the tibia of leg IV containing several stout, peg-like setae.

<==Piona Koch 1842 RFF05
    |--P. aculeatipalpis Viets 1954 RFF05
    |--P. acutidens Lundblad 1941 RFF05
    |--P. alpicola WP99
    |--P. amazonica Viets 1954 RFF05
    |--P. anisitsi Daday 1905 RFF05
    |--P. atrorubra Lundblad 1937 RFF05
    |--P. australica Viets 1980 H98
    |--P. carnea (Koch 1836) L91
    |--P. chilensis Viets 1968 RFF05
    |--P. coccinea (Koch 1836) PH91
    |--P. confinis Lundblad 1943 RFF05
    |--P. conglobata BK91
    |--P. constricta PH91
    |--P. cumberlandensis (Rainbow 1906) (see below for synonymy) H98
    |--P. davidcooki Rosso de Ferradás & Böttger 1997 RFF05
    |--P. deformis Lundblad 1941 RFF05
    |--P. dejecta Viets 1937 RFF05
    |--P. dissipata Lundblad 1938 RFF05
    |--P. erratica Marshall 1940 RFF05
    |--P. fissa Viets 1940 RFF05
    |--P. indica WL09
    |--P. interrupta WL09
    |--P. juncta Lundblad 1936 RFF05
    |    |--P. j. juncta RFF05
    |    `--P. j. inconspicua Lundblad 1943 RFF05
    |--P. lapointei WL09
    |--P. limnetica WL09
    |--P. longipalpis WP99
    |--P. marchanti Harvey 1996 H98
    |--P. mitchelli WL09
    |--P. murleyi Harvey 1996 H98
    |--P. neumani (Koenike 1883) PH91
    |--P. nodata (Müller 1776) L91
    |--P. oxyura Viets 1954 RFF05
    |--P. panniculata Viets 1940 RFF05
    |--P. paucipora (Thor 1897) PH91
    |--P. plaumanni Lundblad 1943 RFF05
    |--P. plenipalpis Viets 1954 RFF05
    |--P. praegracilis Viets 1937 RFF05
    |--P. (Tetrapiona) pseudouncata (Piersig 1906) SL71
    |--P. punctatissima Viets 1954 RFF05
    |--P. puripalpis Viets 1984 H98
    |--P. pusilla (Neumann 1875) PH91
    |--P. rectimarginata Viets 1954 RFF05
    |--P. reflexidens Viets 1940 RFF05
    |--P. robustipalpis Viets 1954 RFF05
    |--P. rotunda (Kramer 1879) RFF05
    |    |--P. r. rotunda RFF05
    |    |--P. r. acutipes Viets 1954 RFF05
    |    `--P. r. rotundiformis Lundblad 1938 RFF05
    |--P. rotundoides (Thor 1897) PH91
    |--P. serratiseta Viets 1956 RFF05
    |--P. setipes Cook 1980 RFF05
    |--P. sicaria Lundblad 1937 RFF05
    |--P. stjoerdalensis (Thor 1897) PH91
    |--P. sudamericana Viets 1910 RFF05
    |--P. tenuipalpis Viets 1954 RFF05
    |--P. thoracica Lundblad 1941 RFF05
    |--P. (Piona) uncataexigua Viets 1949 SL71
    |--P. ungularis Lundblad 1943 RFF05
    |--P. unidens Viets 1954 RFF05
    |--P. variabilis BK91
    `--P. venezuelensis Viets 1956 RFF05

Piona cumberlandensis (Rainbow 1906) [=Atax cumberlandensis, Unionicola cumberlandensis, Unionocola (l. c.) cumberlandensis; incl. Piona uncatiformis Lundblad 1941] H98

*Type species of generic name indicated

REFERENCES

[BK91] Biesiadka, E., & W. Kowalik. 1991. Water mites (Hydracarina) as indicators of trophy and pollution in lakes. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 475–481. SPB Academic Publishing: The Hague.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[L91] Leimann, J. 1991. Structure and formation of the sperm package of Piona carnea (Koch, 1836) (Prostigmata, Hydrachnidia), a copulating water mite. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 449–454. SPB Academic Publishing: The Hague.

[PH91] Punčochár, P., & J. Hrbáček. 1991. Water mites in the plankton of Hubenov Reservoir and their relations to fish stock composition. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 449–457. SPB Academic Publishing: The Hague.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

[SL71] Spain, A. V., & M. Luxton. 1971. Catalog and bibliography of the Acari of the New Zealand subregion. Pacific Insects Monograph 25: 179–226.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

[WP99] Walter, D. E., & H. C. Proctor. 1999. Mites: Ecology, Evolution and Behaviour. CABI Publishing: Wallingford (UK).

Gretacarus

Selected features of Gretacarus australicus, from Viets (1978).

Belongs within: Neohydrachnidia.

Gretacarus is an Australian genus of water mites characterised by the presence of a large, robust seta set on an anteriorly directed protuberance on the palp tibia, and numerous pairs of acetabula set in several rows.

<==Gretacarus Viets 1978 [Gretacaridae, Gretacarinae] H98
    |--G. abnormipalpis Cook 1986 H98
    |--G. australicus Viets 1978 H98
    |--G. clodus Cook 1986 H98
    |--G. concavus Cook 1986 H98
    |--G. crankus Cook 1986 H98
    |--G. evidus Cook 1986 H98
    |--G. expansisetus Cook 1986 H98
    |--G. kurtvietsi Cook 1986 H98
    |--G. kyphopalpis Cook 1986 H98
    |--G. mutilus Viets 1978 H98
    `--G. oldus Cook 1986 H98

*Type species of generic name indicated

REFERENCES

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

Homocrinidae

Ectenocrinus simplex assemblage, copyright James St. John.

Belongs within: Inadunata.

The Homocrinidae are a group of crinoids with small, conical cups known from the Middle Ordovician to the Middle Silurian (Moore et al. 1978).

Characters (from Moore et al. 1978): Cup small, conical; B, C, and E radials compound: anal X incorporated in cup between C and D radials.

Homocrinidae [Homocrinacea, Homocrinicae, Homocrinida]
    |--Ibexocrinus Lane 1970 ML78
    |    `--*I. lepton Lane 1970 A98, ML78
    |--‘Ascocrinus’ Jaekel 1918 non Barrande 1887 (n. d.) J18
    |    `--*A. bohemicus Jaekel 1918 J18
    |--Daedalocrinus Ulrich 1925 ML78
    |    `--*D. kirki Ulrich 1925 ML78
    |--Drymocrinus Ulrich 1925 ML78
    |    `--*D. geniculatus (Ulrich 1879) [=Heterocrinus geniculatus] ML78
    |--Sygcaulocrinus Ulrich 1925 ML78
    |    `--*S. typus Ulrich 1925 ML78
    |--Homocrinus Hall 1852 ML78
    |    |--*H. parvus Hall 1852 ML78
    |    `--H. rhenanus Jaekel 1918 J18
    `--Ectenocrinus Miller 1889 ML78
         |--*E. simplex (Hall 1847) [=Heterocrinus simplex] ML78
         `--E. grandis ML78

*Type species of generic name indicated

REFERENCES

[A98] Ausich, W. I. 1998. Early phylogeny and subclass division of the Crinoidea (phylum Echinodermata). Journal of Paleontology 72 (3): 499–510.

[J18] Jaekel, O. 1918. Phylogenie und System der Pelmatozoen. Paläontologische Zeitschrift 3: 1–128.

[ML78] Moore, R. C., N. G. Lane, H. L. Strimple, J. Sprinkle & R. O. Fay. 1978. Inadunata. In: Moore, R. C., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt T. Echinodermata 2. Crinoidea vol. 2 pp. T520–T759. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

Parasitengonina

Calyptostoma larvae on crane fly, copyright Steve Begin.

Belongs within: Prostigmata.
Contains: Trombellidae, Leeuwenhoekiidae, Johnstonianidae, Neothrombiidae, Chyzeriidae, Trombidiidae, Microtrombidiidae, Trombiculidae, Erythraeoidea, Hydrachnidia.

The Parasitengonina are a diverse group of mites with parasitic larvae and predatory deutonymphs and adults, with the active instars separated by inactive calyptostatic stages. Subgroups include the Trombidiae, terrestrial Parasitengonina in which the predatory adults have chelicerae with a movable digit that is usually bladelike and not retractable, and the parasitic larvae have a prodorsal plate (scutum) and urstigmata present between the contiguous coxal plates I–II. A pregenital tubercle is present in adults of the family Tanaupodidae but absent in other families (Walter et al. 2009).

<==Parasitengonina (see below for synonymy) DPD16
    |  i. s.: Allotanaupodidae [Allotanaupodoidea] ZF11
    |--Trombidiae [Trombidiina] DPD16
    |    |  i. s.: Amphotrombium WL09 [Amphotrombiidae, Amphotrombioidea ZF11]
    |    |           `--A. jenseni WL09
    |    |         Yurebilla Southcott 1996 H98 [Yurebillidae, Yurebilloidea ZF11]
    |    |           `--Y. gracilis Southcott 1996 H98
    |    |--Tanaupodidae [Tanaupodoidea] W91
    |    |    |--Lassenia Newell 1957 [Lasseniinae] W91
    |    |    |--Polydiscia Methlagl 1928 [Polydisciinae] W91
    |    |    `--Tanaupodinae W91
    |    |         |--Rhinothrombium Berlese 1910 T35
    |    |         |    `--R. nemoricola S51
    |    |         |--Tanaupodus Haller 1882 W91
    |    |         |    |--*T. steudeli Haller 1882 SB63
    |    |         |    `--T. passimpilosus Berlese 1910 SB63
    |    |         `--Eothrombium Berlese 1910 T35
    |    |              |--E. lasseni WL09
    |    |              `--E. nemoricola WL09
    |    `--Trombidioidea [Microtrombidioidea, Trombiculoidea] LKW09
    |         |  i. s.: Georgiothrombium pulcherrimum W01
    |         |         Parawenhoekia Paoli 1937 S61
    |         |           `--*P. dectici Paoli 1937 S61
    |         |         Endotrombicula Ewing 1931 S61
    |         |         Achaemenothrombiidae ZF11
    |         |         Trombellidae W91
    |         |         Audyana Womersley 1954 H98 [Audyanidae W91]
    |         |           `--A. thompsoni Womersley 1954 H98
    |         |         Neotrombidiidae W91
    |         |           |  i. s.: Monunguis streblida WL09
    |         |           |--Anomalothrombiinae WL09
    |         |           `--Neotrombidium Leonardi 1901 [Neotrombidiinae] S61
    |         |                |--N. anuroporum WL09
    |         |                |--N. barringunense Hirst 1928 H98
    |         |                |--N. beeri WL09
    |         |                |--N. gracilare Womersley 1963 H98
    |         |                |--N. gracilipes Womersley 1963 H98
    |         |                |--N. neptunium Southcott 1961 S61 (see below for synonymy)
    |         |                |--N. papuanum (Canestrini 1884) [=Trombidium papuanum, Trombicula papuana] H98
    |         |                |--N. samsinaki WL09
    |         |                `--N. tridentifer (Ewing 1909) [=Rhyncholophus tridentifer] S61
    |         |         Leeuwenhoekiidae W91
    |         |--Johnstonianidae DPD16
    |         |--Neothrombiidae W91
    |         `--+--+--Chyzeriidae DPD16
    |            |  `--Trombidiidae DPD16
    |            `--+--Microtrombidiidae DPD16
    |               `--Trombiculidae DPD16
    `--+--Erythraeoidea DPD16
       `--+--Hydrachnidia DPD16
          `--Calyptostoma Cambridge 1875 S61 (see below for synonymy)
               |--C. expalpe (Hermann 1804) S61 (see below for synonymy)
               |--C. caelatum (Berlese 1904) [=Smaris caelata, C. caelata] S76
               |--C. extranea (Koch 1867) [=Smaridia extranea] H98
               |--C. latiseta Shiba 1976 S76
               |--C. lyncaea (Berlese 1887) [=Smaris lyncaea; incl. S. impressa Canestrini 1882 non Koch 1837] S61
               |--C. simplexa Shiba 1976 S76
               `--C. velutinum (Müller 1776) [=Acarus velutinus] S61

Calyptostoma Cambridge 1875 S61 [Calyptostomatidae DPD16, Calyptostomatoidea, Calyptostomidae, Calyptostomidoidea]

Calyptostoma expalpe (Hermann 1804) S61 [=Trombidium expalpe S61, Smaris expalpis S61; incl. *C. hardii Cambridge 1875 S61, C. hardyi S61, Smaris impressa Koch 1837 S61, S. plana Koch 1879 T04]

Neotrombidium neptunium Southcott 1961 S61 [=N. tridentifer Southcott 1957 non Rhyncholophus tridentifer Ewing 1909 S61, N. bidentifer (l. c.) H98]

Parasitengonina [Apobolostigmata, Engonostigmata, Erythraeina, Erythraiae, Parasitengona, Parasitengonae, Parasitenini, Trombidia, Trombidina] DPD16

*Type species of generic name indicated

REFERENCES

[DPD16] Dabert, M., H. Proctor & J. Dabert. 2016. Higher-level molecular phylogeny of the water mites (Acariformes: Prostigmata: Parasitengonina: Hydrachnidiae). Molecular Phylogenetics and Evolution 101: 75–90.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[K90] Kethley, J. 1990. Acarina: Prostigmata (Actinedida). In: Dindal, D. L. (ed.) Soil Biology Guide pp. 667–756. John Wiley & Sones: New York.

[LKW09] Lindquist, E. E., G. W. Krantz & D. E. Walter. 2009. Classification. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 97–103. Texas Tech University Press.

[S51] Schweizer, J. 1951. Die Landmilben des schweizerischen Nationalparkes. 2. Teil: Trombidiformes. Ergebnisse der wissenschaftlichen Untersuchung des schweizerischen Nationalparks [Résultats des recherches scientifiques entreprises au Parc National Suisse] 3 (Neue Folge): 51–172.

[SB63] Schweizer, J., & C. Bader. 1963. Die Landmilben der Schweiz (Mittelland, Jura und Alpen): Trombidiformes Reuter, mit 217 Arten und Unterarten und 193 Originalzeichnungen. Denkschriften der Schweizerischen Naturforschenden Gesellschaft [Mémoires de la Société Helvétique des Sciences Naturelles] 84 (2): i–vi, 209–378.

[S76] Shiba, M. 1976. Taxonomic investigation on free-living Prostigmata from the Malay Peninsula. Nature and Life in Southeast Asia 7: 83–229.

[S61] Southcott, R. V. 1961. Studies on the systematics and biology of the Erythraeoidea (Acarina), with a critical revision of the genera and subfamilies. Australian Journal of Zoology 9: 367–610.

[T35] Thor, S. 1935. Übersicht und Einteilung der Familie Trombidiidae W. E. Leach 1814 in Unterfamilien. Zoologischer Anzeiger 109 (5–6): 107–112.

[T04] Trägårdh, I. 1904. Fauna Arctica: Eine Zusammenstellung der arktischen Tierformen, mit besonderer Berücksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das Nördliche Eismeer im Jahre 1898 vol. 4 pt 1. Monographie der arktischen Acariden. Gustav Fischer: Jena.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

[W91] Welbourn, W. C. 1991. Phylogenetic studies of the terrestrial Parasitengona. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 163–170. SPB Academic Publishing: The Hague.

[W01] Wohltmann, A. 2001. Closely related species of Parasitengonae (Acari: Prostigmata) inhabiting the same areas: features facilitating coexistence. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10^th International Congress pp. 121–135. CSIRO Publishing: Melbourne.

[ZF11] Zhang, Z.-Q., Q.-H. Fan, V. Pesic, H. Smit, A. V. Bochkov, A. A. Khaustov, A. Baker, A. Wohltmann, T. Wen, J. W. Amrine, P. Beron, J. Lin, G. Gabryś & R. Husband. 2011. Order Trombidiformes Reuter, 1909. In: Zhang, Z.-Q. (ed.) Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness. Zootaxa 3148: 129–138.

Chyzeriidae

Chyzeria sp., copyright Matias G.

Belongs within: Parasitengonina.

The Chyzeriidae are a group of parasitengonine mites in which adults usually bear numerous setiferous, finger-like projections on the idionotum (Walter et al. 2009).

Characters (from Walter et al. 2009): Adult gnathosoma not retractable into idiosoma; cheliceraewith movable digit bladelike and curved, palp bearing enlarged spinelike setae adjacent to tibial claw; prodorsal area bearing one pair of trichobothria on prodorsal plate and two pairs of lateral eyes; dorsal cuticle peripherally with several pairs of long papillae densely covered with plumose setae, strongly reticulate medially and bearing moderately dense pelage of simple setae; with three pairs of genital papillae, pregenital tubercle absent. Larva heteromorphic; palpgenu with one seta; idionotum with prodorsal plate (scutum) bearing four pairs of setae including one pair of trichobothria but lacking scutellum, setae c1 located on paired platelets; ventrally with sessile urstigmata and well-defined anus, anal plate bearing two pairs of setae; coxal plates I–III bearing 2, 1, and 1 setae, respectively, plates I–II contiguous; legs with genua each bearing four setae.

<==Chyzeriidae [Chyzeriniini, Chyzerioidea]
    |--Cretessenia leoni WL09
    |--Napassenia aginapaica WL09
    |--Perumaropta [Perumaroptinae] WL09
    |    `--P. mirsadi WL09
    |--Pteridopus Newell & Vercammen-Grandjean 1964 [Pteridopodinae] W91
    |    |--P. auditor WL09
    |    `--P. pseudhannemania WL09
    |--Ralphaudyana Vercammen-Grandjean et al. 1974 [Ralphaudyaninae] W91
    |    |--R. amamiensis WL09
    |    `--R. iranensis WL09
    `--Chyzeria Canestrini 1897 [incl. Grossia Womersley 1954; Chyzeriinae] H98
         |--C. armigera Hirst 1929 H98
         |--C. australiensis Hirst 1928 [=C. australiense] H98
         |--C. derricki Southcott 1982 H98
         |--C. flindersi Southcott 1982 H98
         |--C. hirsti Womersley 1934 [=C. australiense var. hirsti, C. australiensis var. hirsti] H98
         |--C. insulana Hirst 1929 H98
         |--C. montana Hirst 1928 H98
         |--C. musgravei Hirst 1928 [=C. australiense var. musgravei] H98
         |--C. novaezealandiae Hirst 1924 QH01 [=C. novaehollandiae (l. c.) W34]
         |--C. occidentalis Hirst 1929 [=C. australiense var. occidentalis, C. australiensis var. occidentalis] H98
         |--C. onychia (Womersley 1954) [=Grossia onychia] H98
         |--C. poivrei WL09
         `--C. queenslandica Womersley 1942 H98

*Type species of generic name indicated

REFERENCES

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[QH01] Qin, T.-K., & R. C. Henderson. 2001. Taxonomy of New Zealand Prostigmata: past, present and future. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10^th International Congress pp. 35–39. CSIRO Publishing: Melbourne.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

[W91] Welbourn, W. C. 1991. Phylogenetic studies of the terrestrial Parasitengona. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 163–170. SPB Academic Publishing: The Hague.

Womersley, H. 1934. A revision of the trombid and erythraeid mites of Australia with descriptions of new genera and species. Records of the South Australian Museum 5 (2): 179–254.