Venonia

Venonia micarioides, copyright Angus Veitch.

Belongs within: Lycosidae.

Venonia is a genus of small, slender wolf spiders found in southeast Asia and Australasia that build small sheet-webs close to the ground (Yoo & Framenau 2006).

Characters (from Yoo & Framenau 2006): Coloration uniform, light brown to black, with posterodorsal white spot on abdomen; male pedipalp with strongly asymmetrical cymbium appearing apico-retrolaterally truncated, tegular apophysis membranous, embolic division reduced, pedipalp tibia rotated in relation to femur so that pedipalp bulbs are facing each other; epigyne with posterior central incision, lacking sclerotised plate.

<==Venonia Thorell 1894 YF06
    |--V. kokoda Lehtinen & Hippa 1979 YF06
    `--+--V. micarioides (Koch 1877) [=Aulonia micarioides, Artoria micarioides] YF06
       `--+--V. infundibulum Yoo & Framenau 2006 YF06
          |--V. kimjoopili Yoo & Framenau 2006 YF06
          |--V. nata Yoo & Framenau 2006 YF06
          |--V. sungahae Yoo & Framenau 2006 YF06
          |--V. vilkkii Lehtinen & Hippa 1979 YF06
          `--+--+--V. cinctipes (Simon 1898) [=Anomalomma cinctipes] YF06
             |  `--V. muju (Chrysanthus 1967) [=Anomalomma muju] YF06
             `--+--*V. coruscans Thorell 1894 YF06
                `--+--+--V. chaiwooi Yoo & Framenau 2006 YF06
                   |  `--V. joejim Yoo & Framenau 2006 YF06
                   `--+--V. micans (Simon 1898) [=Anomalomma micans; incl. V. gabrielae Barrion & Litsinger 1995] YF06
                      `--+--V. choiae Yoo & Framenau 2006 YF06
                         `--V. milla Lehtinen & Hippa 1979 YF06

*Type species of generic name indicated

REFERENCES

[YF06] Yoo, J.-S., & V. W. Framenau. 2006. Systematics and biogeography of the sheet-web building wolf spider genus Venonia (Araneae: Lycosidae). Invertebrate Systematics 20: 675–712.

Evippa

Evippa sp., copyright A. Nadolny.

Belongs within: Lycosidae.

Evippa is a genus of wolf spiders found from southern China to northern Africa. Male pedipalps of this genus have an embolus forming a slender functional conductor that projects beyond the tegular apophysis.

<==Evippa Simon 1882 [Evippinae] MGK03
    |--*E. arenaria (Audouin 1827) [=Lycosa arenaria] MGK03
    |--E. apsheronica Marusik, Guseinov & Koponen 2003 MGK03
    |--E. badchysica Šternbergs 1979 MGK03
    |--E. benevola (Pickard-Cambridge 1885) MGK03
    |--E. caucasica Marusik, Guseinov & Koponen 2003 MGK03
    |--E. concolor (Kroneberg 1875) [=Lycosa concolor] MGK03
    |--E. eltonica Dunin 1994 MGK03
    |--E. fujianensis Peng, Yin & Kim 1996 MGK03
    |--E. helanshangensis Peng, Yin & Kim 1996 MGK03
    |--E. jocquei Alderweireldt 1991 MGK03
    |--E. kronebergi Roewer 1955 (see below for synonymy) MGK03
    |--E. onager Simon 1895 MGK03
    |--E. praelongipes (Pickard-Cambridge 1870) MGK03
    |--E. schenkeli Šternbergs 1979 MGK03
    |--E. sjostedti Schenkel 1936 [incl. Xerolycosa brunneopicta Loksa 1965, E. potanini Schenkel 1963] MGK03
    |--E. sibirica Marusik 1995 MGK03
    `--E. turkmenica Šternbergs 1979 MGK03

Nomen nudum: Evippa beshkentica Andreeva 1976

Evippa kronebergi Roewer 1955 [=Lycosa aculeata Kroneberg 1875 non (Clerck 1757), Acantholycosa aculeata, Evippa aculeata, Pardosa aculeata]

*Type species of generic name indicated

REFERENCES

[MGK03] Marusik, Yu. M., E. F. Guseinov & S. Koponen. 2003. Spiders (Arachnida: Aranei) of Azerbaijan. 2. Critical survey of wolf spiders (Lycosidae) found in the country with description of three new species and brief review of Palaearctic Evippa Simon, 1885. Arthropoda Selecta 12 (1): 47–65.

Pardosa

Pardosa lugubris, copyright MdE.

Belongs within: Lycosinae.

Pardosa is a diverse genus of wolf spiders commonly found in damp habitats near ponds and streams. Species of this genus include P. lugubris, widespread in the Palaearctic region.

<==Pardosa Koch 1847 [Pardosinae] MGK03
    |--+--+--P. brevivulva PY07
    |  |  `--P. hedini PY07
    |  `--+--P. laura PY07
    |     `--P. takahashii (Saito 1936) PY07, VMP02
    `--+--+--P. paludicola (Clerck 1757) PY07, MGK03
       |  `--+--+--P. alacris PY07
       |     |  `--P. lugubris (Walckenaer 1802) PY07, MGK03 [=Lycosa lugubris V09]
       |     `--+--P. amentata (Clerck 1758) PY07, F08 (see below for synonymy)
       |        `--+--P. riparia PY07
       |           `--+--P. prativaga (Koch 1870) PY07, K55
       |              `--P. pullata (Clerck 1757) PY07, MGK03 [=Lycosa pullata V09]
       `--+--P. nigra PY07
          `--+--+--P. astrigera Koch 1878 PY07, KS02
             |  `--+--P. isago PY07
             |     `--P. lyrifera PY07
             `--+--P. hortensis (Thorell 1872) PY07, MGK03
                `--+--+--P. oreophila PY07
                   |  `--P. saturatior PY07
                   `--+--P. blanda PY07
                      `--+--P. agrestis (Westring 1861) PY07, MGK03
                         `--P. palustris (Linnaeus 1758) PY07, F08 [=Aranea palustris F08]

Pardosa incertae sedis:
  P. adumbrata (Urquhart 1887) [=Lycosa adumbrata] NS00
  P. adustella Roewer 1951 MGK03
  P. aenigmatica Tongiorgi 1966 MGK03
  P. agricola (Thorell 1856) MGK03
  P. aquila Buchar & Thaler 1998 MGK03
  P. atrata (Thorell 1873) K02
  P. azerifalcata Marusik, Guseinov & Koponen 2003 MGK03
  P. bifasciata (Koch 1834) MGK03
  P. birmanica (Simon 1884) HU08
  P. buchari Ovtsharenko 1979 MGK03
  P. caucasica Ovtsharenko 1979 MGK03
  P. colchica Mcheidze 1947 MGK03
  P. feldmanni (Strand 1907) (n. d.) [=Lycosa feldmanni] J98
  P. fuscula CM07
  P. goyeni FF99
  P. groenlandica CM07
  P. gusarensis Marusik, Guseinov & Koponen 2003 MGK03
  P. hanrasanensis Jo & Paik 1984 MGK03
  P. hyperborea (Thorell 1872) K02
  P. ibex Buchar & Thaler 1998 MGK03
  P. incerta Nosek 1905 MGK03
  P. injucunda P.-Cambridge 1876 [incl. Lycosa feldmanni xanthippe Strand 1916, P. feldmanni xanthippe] J98
  P. italica Tongiorgi 1966 MGK03
  P. jergeniensis Ponomarev 1979 MGK03
  P. josemitensis (Strand 1908) [=Lycosa josemitensis] J98
  P. lapidicina (Emerton 1885) KP14
  P. lignaria K01
  P. luctinosa Simon 1876 MGK03
  P. mackenziana (Keyserling 1877) PS08
  P. modica CM07
  P. mongolica Kulczyński 1901 K01
  P. monticola (Clerck 1757) K55
  P. morosa (Koch 1870) MGK03
  P. naevioides (Strand 1916) [=Lycosa naevioides] J98
  P. nebulosa (Thorell 1872) MGK03
  P. nigriceps (Thorell 1856) PG08 [=Lycosa nigriceps V09]
  P. paracolchica Zyuzin & Logunov 2000 MGK03
  P. pirculiensis Zyuzin & Logunov 2000 MGK03
  P. pontica (Thorell 1875) MGK03
  P. proxima (Koch 1847) MGK03 [=Lycosa proxima V09]
  P. proximella Strand 1906 (n. d.) [=Lycosa proximella] J98
    |--P. p. proximella J98
    `--P. p. messingerae (Strand 1916) [=Lycosa proximella messingerae] J98
  P. purbeckensis (Westring 1861) PG08
  P. saltans Töpfer-Hofmann 2000 PG08
  P. saltuaria (Koch 1870) MGK03
  P. schenkeli Lessert 1904 MGK03
  P. seydi (Strand 1908) [=Lycosa seydi] J98
  P. sphagnicola (Dahl 1908) MGK03
  P. strigillata (Simon 1876) K55
  P. tasevi Buchar 1968 MGK03
  P. tatarica (Thorell 1875) MGK03
  P. vicaria S99
  P. vittata (Keyserling 1863) MGK03

Pardosa amentata (Clerck 1758) PY07, F08 [=Lycosa amentata V09; incl. Aranea fumigata Linnaeus 1758 F08, P. amentata f. ileachensis J02, A. saccata Linnaeus 1758 F08]

*Type species of generic name indicated

REFERENCES

[CM07] Cokendolpher, J. C., & P. G. Mitov. 2007. Natural enemies. In: Pinto-da-Rocha, R., G. Machado & G. Giribet (eds) Harvestmen: The Biology of Opiliones pp. 339–373. Harvard University Press: Cambridge (Massachusetts).

[FF99] Forster, R., & L. Forster. 1999. Spiders of New Zealand and their World-wide Kin. University of Otago Press: Dunedin (New Zealand).

[F08] Framenau, V. W. 2008. A new wolf spider species of the genus Artoria from Western Australia (Araneae: Lycosidae). Records of the Western Australian Museum 24 (4): 363–368.

[HU08] Hore, U., & V. P. Uniyal. 2008. Diversity and composition of spider assemblages in five vegetation types of the Terai Conservation Area, India. Journal of Arachnology 36 (2): 251–258.

[J98] Jäger, P. 1998. Das Typenmaterial der Spinnentiere (Arachnida: Acari, Amblypygi, Araneae, Opiliones, Pseudoscorpiones, Scorpiones, Uropygi) aus dem Museum Wiesbaden. Jahrbuecher des Nassauischen Vereins fuer Naturkunde 119: 81–91.

[J02] Jocqué, R. 2002. Genitalic polymorphism—a challenge for taxonomy. Journal of Arachnology 30 (2): 298–306.

[KP14] Keiser, C. N., & J. N. Pruitt. 2014. Submersion tolerance in a lakeshore population of Pardosa lapidicina (Araneae: Lycosidae). Journal of Arachnology 42 (2): 192–194.

[KS02] Kiss, B., & F. Samu. 2002. Comparison of autumn and winter development of two wolf spider species (Pardosa, Lycosidae, Araneae) having different life history patterns. Journal of Arachnology 30 (2): 409–415.

[K02] Koponen, S. 2002. Ground-living spiders in bogs in northern Europe. Journal of Arachnology 30 (2): 262–267.

[K55] Kraus, O. 1955. Spinnen von Korsika, Sardinien und Elba (Arach., Araneae). Senckenbergiana Biologica 36: 371–394.

[K01] Kulczyński, V. 1901. Arachnoideák [Arachnoidea]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 311–369. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[MGK03] Marusik, Yu. M., E. F. Guseinov & S. Koponen. 2003. Spiders (Arachnida: Aranei) of Azerbaijan. 2. Critical survey of wolf spiders (Lycosidae) found in the country with description of three new species and brief review of Palaearctic Evippa Simon, 1885. Arthropoda Selecta 12 (1): 47–65.

[NS00] Nicholls, D. C., P. J. Sirvid, S. D. Pollard & M. Walker. 2000. A list of arachnid primary types held in Canterbury Museum. Records of the Canterbury Museum 14: 37–48.

[PY07] Park, Y. C., J.-S. Yoo, M. P. Schwarz, N. Murphy & J.-P. Kim. 2007. Molecular phylogeny of east Asian wolf spiders (Araneae: Lycosidae) inferred from mitochondrial 12S ribosomal DNA. Annals of the Entomological Society of America 100 (1): 1–8.

[PG08] Pétillon, J., & A. Garbutt. 2008. Success of managed realignment for the restoration of salt-marsh biodiversity: preliminary results on ground-active spiders. Journal of Arachnology 36 (2): 388–393.

[PS08] Pinzón, J., & J. Spence. 2008. Performance of two arboreal pitfall trap designs in sampling cursorial spiders from tree trunks. Journal of Arachnology 36 (2): 280–286.

[S99] Simon, E. 1899. Ergebnisse einer Reise nach dem Pacific (Schauinsland 1896–1897). Zoologische Jahrbücher, Abteilung für Systematik, Geographie un Biologie der Thiere 12 (4): 411–437.

[V09] Verdcourt, B. (ed.) 2009. Additions to the Wild Fauna and Flora of the Royal Botanic Gardens, Kew. XXVI. Miscellaneous records. Kew Bulletin 64 (1): 183–194.

[VMP02] Vink, C. J., A. D. Mitchell & A. M. Paterson. 2002. A preliminary molecular analysis of phylogenetic relationships of Australasian wolf spider genera (Araneae, Lycosidae). Journal of Arachnology 30 (2): 227–237.

Last updated: 23 June 2017.

Piratinae

Pirata hygrophilus, copyright Kjetil Fjellheim.

Belongs within: Lycosidae.

The Piratinae is a group of wolf spiders characterised by male pedipalps in which the functional conductor is combined with the short thin embolus into a sickle-shaped complex that rests in a deep, narrow ascending tegular groove (Yoo & Framenau 2006). Members of the genus Pirata, the pirate wolf spiders, are found in marshy habitats and may be seen running across water surfaces.

<==Piratinae
    |--Hygrolycosa Dahl 1908 YF06
    |    `--H. rubrofasciata Ohlert 1865 FY06
    |--Aulonia Koch 1847 YF06
    |    |--*A. albimana (Walckenaer 1805) MGK03
    |    `--A. kratochvili Dunin, Buchar & Absolon 1986 MGK03
    `--Pirata Sundevall 1833 YF06
         |  i. s.: P. insularis Emerton 1885 MGK03
         |         P. latitans (Blackwall 1841) MGK03
         |         P. piraticus (Clerck 1757) MGK03 [=Araneus piraticus S99, Lycosa (Pirata) piratica K01]
         |         P. piratimorpha (Strand 1908) [=Tarentula piratimorpha, Piratula piratimorpha] J98
         |         P. piscatorius (Clerck 1757) K02 [=Lycosa (Pirata) piscatoria K01]
         |         P. uliginosus (Thorell 1856) RKD02
         |--P. subpiraticus (Bösenberg & Strand 1906) PY07, YF06
         `--+--P. piratoides PY07
            `--+--P. hygrophilus Thorell 1872 PY07, VMP02
               `--P. knorri (Scopoli 1763) PY07, VMP02

*Type species of generic name indicated

REFERENCES

[FY06] Framenau, V. W., & J.-S. Yoo. 2006. Systematics of the new Australian wolf spider genus Tuberculosa (Araneae: Lycosidae). Invertebrate Systematics 20: 185–202.

[J98] Jäger, P. 1998. Das Typenmaterial der Spinnentiere (Arachnida: Acari, Amblypygi, Araneae, Opiliones, Pseudoscorpiones, Scorpiones, Uropygi) aus dem Museum Wiesbaden. Jahrbuecher des Nassauischen Vereins fuer Naturkunde 119: 81–91.

[K02] Koponen, S. 2002. Ground-living spiders in bogs in northern Europe. Journal of Arachnology 30 (2): 262–267.

[K01] Kulczyński, V. 1901. Arachnoideák [Arachnoidea]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 311–369. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[MGK03] Marusik, Yu. M., E. F. Guseinov & S. Koponen. 2003. Spiders (Arachnida: Aranei) of Azerbaijan. 2. Critical survey of wolf spiders (Lycosidae) found in the country with description of three new species and brief review of Palaearctic Evippa Simon, 1885. Arthropoda Selecta 12 (1): 47–65.

[PY07] Park, Y. C., J.-S. Yoo, M. P. Schwarz, N. Murphy & J.-P. Kim. 2007. Molecular phylogeny of east Asian wolf spiders (Araneae: Lycosidae) inferred from mitochondrial 12S ribosomal DNA. Annals of the Entomological Society of America 100 (1): 1–8.

[RKD02] Relys, V., S. Koponen & D. Dapkus. 2002. Annual differences and species turnover in peat bog spider communities. Journal of Arachnology 30 (2): 416–424.

[S99] Simon, E. 1899. Ergebnisse einer Reise nach dem Pacific (Schauinsland 1896–1897). Zoologische Jahrbücher, Abteilung für Systematik, Geographie un Biologie der Thiere 12 (4): 411–437.

[VMP02] Vink, C. J., A. D. Mitchell & A. M. Paterson. 2002. A preliminary molecular analysis of phylogenetic relationships of Australasian wolf spider genera (Araneae, Lycosidae). Journal of Arachnology 30 (2): 227–237.

[YF06] Yoo, J.-S., & V. W. Framenau. 2006. Systematics and biogeography of the sheet-web building wolf spider genus Venonia (Araneae: Lycosidae). Invertebrate Systematics 20: 675–712.

Last updated: 23 June 2017.

Lycosinae

Female Geolycosa sp. on egg-sac, copyright Fir0002.

Belongs within: Lycosidae.
Contains: Venatrix, Pardosa, Trochosa, Lycosa, Alopecosa.

The Lycosinae is a group of wolf spiders characterised by male pedipalps bearing a transverse tegular apophysis carrying a ventrally directed spur and a sinuous channel on its dorsal surface (Framenau 2006). Members of this group include the burrowing wolf spiders of the genus Geolycosa, which ambush prey directly from the burrow rather than emerging to hunt in the open in the manner of other wolf spiders. The cosmopolitan genus Hogna includes the largest of wolf spiders, with some species reaching over an inch in length. Hoggicosa is a genus of large wolf spiders found in arid regions of Australia, some species of which exhibit a strong dimorphism in colour pattern between males and females (Langlands & Framenau 2010). The northern Australian genus Tuberculosa is distinguished by the presence in males of a field of modified, tubercle-like setae on the underside of the third coxa (Framenau & Yoo 2006).

<==Lycosinae FL13
    |--+--Venatrix PY07
    |  `--+--Pardosa PY07
    |     `--‘Alopecosa’ cinnameopilosa PY07
    `--+--+--Trochosa PY07
       |  `--+--+--Lycosa PY07
       |     |  `--Varacosa Chamberlin & Ivie 1942 PY07, VMP02
       |     |       `--V. avara (Keyserling 1877) VMP02
       |     `--+--Dingosa Roewer 1955 LF10, FL13
       |        |    |--D. serrata (Koch 1877) LF10
       |        |    `--D. simsoni (Simon 1898) LF10
       |        `--+--Geolycosa Montgomery 1904 PY07, FL13
       |           |    |--G. dunini Zyuzin & Logunov 2000 MGK03
       |           |    |--G. goderffroyi K90
       |           |    |--G. hubbelli Wallace 1942 LF10
       |           |    |--G. kijabica (Strand 1916) [=Tarentula kijabica, Scaptocosa kijabica] J98
       |           |    |--G. missouriensis (Banks 1895) LF10
       |           |    |--G. rafaelana Cr90
       |           |    |--G. rogersi Wallace 1942 VMP02
       |           |    `--G. wrightii K90
       |           `--Mainosa Framenau 2006 FL13
       |                `--M. longipes (Koch 1878) LF10
       `--Alopecosa PY07

Lycosinae incertae sedis:
  Hogna Simon 1885 FL13
    |--*H. radiata (Latreille 1817) LF10 [=Lycosa radiata MGK03]
    |--H. alticeps (Kroneberg 1875) [=Tarentula alticeps, Lycosa alticeps] MGK03
    |--H. bergsoei (Thorell 1875) [=Alopecosa bergsoei] MGK03
    |--H. birabenae (Mello-Leitão 1941) [=Birabenia birabenae] Ca90
    |--H. crispipes (Koch 1877) FL13 [incl. Geolycosa tongatabuensis (Strand 1911) PVD10]
    |--H. exigua (Roewer 1960) [=Hoggicosa exigua, Lycosa exigua] LF10
    |--H. gumia (Petrunkevitch 1911) FM11
    |--H. ingens (Blackwall 1857) JD-S07
    |--H. kuyani Framenau, Gotch & Austin 2007 LF10
    |--H. lenta (Hentz 1844) [=Lycosa lenta] SB14
    |--H. ocellata (Koch 1878) [=Lycosa ocellata, Arctosa ocellata] MGK03
    |--H. sternalis (Bertkau 1880) FM11
    `--H. yauliensis (Strand 1908) [=Tarentula yauliensis, Lynxosa yauliensis] J98
  Tuberculosa Framenau & Yoo 2006 FL13
    |--T. monteithi Framenau & Yoo 2006 FY06
    `--+--T. hoggi (Framenau & Vink 2001) [=Venatrix hoggi] FY06
       `--+--*T. harveyi Framenau & Yoo 2006 FY06
          `--T. austini Framenau & Yoo 2006 FY06
  Hoggicosa Roewer 1960 FL13
    |  i. s.: H. natashae Langlands & Framenau 2010 LF10
    |--H. wolodymyri Langlands & Framenau 2010 LF10
    `--+--H. forresti (McKay 1973) [=Lycosa forresti] LF10
       `--+--+--H. castanea (Hogg 1905) (see below for synonymy) LF10
          |  |--H. alfi Langlands & Framenau 2010 LF10
          |  `--H. duracki (McKay 1975) [=Lycosa duracki] LR10
          `--+--H. brennani Langlands & Framenau 2010 LF10
             `--+--H. storri (McKay 1973) [=Lycosa storri] LF10
                `--+--H. bicolor (McKay 1973) [=Lycosa bicolor, Allocosa bicolor] LF10
                   `--H. snelli (McKay 1975) [=Lycosa snelli] LF10
  Tasmanicosa Roewer 1959 FY06 [incl. Orthocosa Roewer 1960 (n. n.) FL13]
    `--*T. tasmanica (Hogg 1905) LF10
  Venator Hogg 1900 FL13
    |--*V. spenceri Hogg 1900 LF10
    `--‘Hogna’ immansueta (Simon 1909) LF10
  Hippasa Simon 1885 F07
    `--H. greenalliae (Blackw. 1867) (see below for synonymy) K92
  Tapetosa Framenau, Main et al. 2009 FL13
    `--*T. darwini Framenau, Main et al. 2009 FM09
  Pavocosa Roewer 1960 FY06
    `--P. gallopavo (Mello-Leitão 1941) LF10
  Costacosa Framenau & Leung 2013 FL13
    |--*C. torbjorni Framenau & Leung 2013 FL13
    `--C. dondalei Framenau & Leung 2013 FL13
  Knoelle Framenau 2006 FL13
    `--*K. clara (Koch 1877) [=Lycosa clara, Allocosa clara] F06
  Cynosa Caporiacco 1933 FL13
  Orinocosa Chamberlin 1916 FL13
  Rabidosa punctulata (Hentz 1844) LF10, SB14
  Allocosa Banks 1900 PY07, VMP02 (see below for synonymy)
    |--‘Glieschiella’ alticeps Mello-Leitão 1944 Ca90
    |--‘Moenkhausiana’ argentinensis Mello-Leitão 1938 (n. d.) Ca90
    |--A. brasiliensis (Petrunkevitch 1910) (see below for synonymy) Ca90
    |--A. funerea Ca90
    |--A. georgicola (Walckenaer 1837) VMP02
    |--‘*Glieschiella’ halophila Mello-Leitão 1932 (n. d.) Ca90
    |--A. hasseltii F06
    |--A. mutillata Ca90
    |--A. palabunda (Koch 1877) [=Lycosa palabunda] E07
    |--A. paraguayensis Ca90
    |--‘Glieschiella’ senex Mello-Leitão 1945 Ca90
    `--A. tuberculipalpa JD-S07

Allocosa Banks 1900 PY07, VMP02 [incl. Glieschiella Mello-Leitão 1932 Ca90, Moenkhausiana Ca90; Allocosinae]

Allocosa brasiliensis (Petrunkevitch 1910) [=*Moenkhausiana brasiliensis; incl. Araucaniocosa difficilis Mello-Leitão 1951] Ca90

Hippasa greenalliae (Blackw. 1867) [=Lycosa greenalliae; incl. Pirata agelenoides Simon 1884, Diapontia agalenoides] K92

Hoggicosa castanea (Hogg 1905) [=Lycosa castanea, Allocosa castanea; incl. L. errans Hogg 1905, *H. errans; incl. L. perinflata Pulleine 1922, L. skeeti Pulleine 1922] LF10

*Type species of generic name indicated

REFERENCES

[Ca90] Capocasale, R. M. 1990. Las especies de la subfamilia Hippasinae de America del Sur (Araneae, Lycosidae). Journal of Arachnology 18: 131–142.

[Cr90] Crawford, C. S. 1990. Scorpiones, Solifugae, and associated desert taxa. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 421–475. John Wiley & Sones: New York.

[E07] Elliott, M. G. 2007. Annotated catalogue of the Pompilidae (Hymenoptera) of Australia. Zootaxa 1428: 1–83.

[F06] Framenau, V. W. 2006. Knoelle, a new monotypic wolf spider genus from Australia (Araneae: Lycosidae). Zootaxa 1281: 55–67.

[F07] Framenau, V. W. 2007. Revision of the new Australian genus Artoriopsis in a new subfamily of wolf spiders, Artoriinae (Araneae: Lycosidae). Zootaxa 1391: 1–34.

[FL13] Framenau, V. W., & A. E. Leung. 2013. Costacosa, a new genus of wolf spider (Araneae, Lycosidae) from coastal north-west Western Australia. Records of the Western Australian Museum Supplement 83: 173–184.

[FM09] Framenau, V. W., B. Y. Main, M. S. Harvey & J. M. Waldock. 2009. Tapetosa, a new monotypic wolf spider genus from Western Australia (Araneae: Lycosidae: Lycosinae). Records of the Western Australian Museum 25 (3): 309–314.

[FY06] Framenau, V. W., & J.-S. Yoo. 2006. Systematics of the new Australian wolf spider genus Tuberculosa (Araneae: Lycosidae). Invertebrate Systematics 20: 185–202.

[FM11] Freire-Jr, G. de B., & P. C. Motta. 2011. Effects of experimental fire regimes on the abundance and diversity of cursorial arachnids of Brazilian savannah (cerrado biome). Journal of Arachnology 39 (2): 263–272.

[J98] Jäger, P. 1998. Das Typenmaterial der Spinnentiere (Arachnida: Acari, Amblypygi, Araneae, Opiliones, Pseudoscorpiones, Scorpiones, Uropygi) aus dem Museum Wiesbaden. Jahrbuecher des Nassauischen Vereins fuer Naturkunde 119: 81–91.

[JD-S07] Jocqué, R., & A. S. Dippenaar-Schoeman. 2007. Spider Families of the World. Royal Museum for Central Africa: Tervuren (Belgium).

[K92] Karsch, F. 1892. Arachniden von Ceylon und von Minikoy gesammelt von den Herren Doctoren P. und F. Sarasin. Berliner Entomologische Zeitschrift 36 (2): 267–310.

[K90] Kotzman, M. 1990. Annual activity patterns of the Australian tarantula Selenocosmia stirlingi (Araneae, Theraphosidae) in an arid area. Journal of Arachnology 18: 123–130.

[LF10] Langlands, P. R., & V. W. Framenau. 2010. Systematic revision of Hoggicosa Roewer, 1960, the Australian 'bicolor' group of wolf spiders (Araneae: Lycosidae). Zoological Journal of the Linnean Society 158: 83–123.

[MGK03] Marusik, Yu. M., E. F. Guseinov & S. Koponen. 2003. Spiders (Arachnida: Aranei) of Azerbaijan. 2. Critical survey of wolf spiders (Lycosidae) found in the country with description of three new species and brief review of Palaearctic Evippa Simon, 1885. Arthropoda Selecta 12 (1): 47–65.

[PVD10] Paquin, P., C. J. Vink & N. Dupérré. 2010. Spiders of New Zealand: annotated family key and species list. Manaaki Whenua Press: Lincoln (New Zealand).

[PY07] Park, Y. C., J.-S. Yoo, M. P. Schwarz, N. Murphy & J.-P. Kim. 2007. Molecular phylogeny of east Asian wolf spiders (Araneae: Lycosidae) inferred from mitochondrial 12S ribosomal DNA. Annals of the Entomological Society of America 100 (1): 1–8.

[SB14] Suter, R. B., & K. Benson. 2014. Nocturnal, diurnal, crepuscular: activity assessments of Pisauridae and Lycosidae. Journal of Arachnology 42 (2): 178–191.

[VMP02] Vink, C. J., A. D. Mitchell & A. M. Paterson. 2002. A preliminary molecular analysis of phylogenetic relationships of Australasian wolf spider genera (Araneae, Lycosidae). Journal of Arachnology 30 (2): 227–237.

Last updated: 22 January 2022.

Venatrix

Venatrix lapidosa, copyright David Paul.

Belongs within: Lycosidae.

Venatrix is a genus of wolf spiders found in Australasia, usually characterised by the presence of two longitudinal light bands (or row of light spots) on the dark venter of the abdomen. Males have a tubercle on the outer edge of the fangs, and pedipalps with claw-like macrosetae on the tip of the cymbium (Framenau 2006).

<==Venatrix Roewer 1960 F06
    |--V. funesta (Koch 1847) [incl. Venator fuscus Hogg 1900, *Venatrix fusca] F06
    |--V. allopictiventris Framenau & Vink 2001 F06
    |--V. amnicola Framenau 2006 F06
    |--V. archookoora Framenau & Vink 2001 F06
    |--V. arenaris (Hogg 1905) F06
    |--V. australiensis Framenau & Vink 2001 F06
    |--V. brisbanae (Koch 1878) [=Lycosa brisbanae, L. brisbanensis (l. c.), Pirata brisbanae] F06
    |--V. esposica Framenau & Vink 2001 F06
    |--V. fontis Framenau & Vink 2001 F06
    |--V. furcillata (Koch 1867) [=Lycosa furcillata, Allocosa furcillata] F06
    |--V. hickmani Framenau & Vink 2001 F06
    |--V. konei (Berland 1924) (see below for synonymy) F06
    |--V. koori Framenau & Vink 2001 F06
    |--V. kosciuskoensis (McKay 1974) [=Lycosa kosciuskoensis] F06
    |--V. lapidosa (McKay 1974) F06
    |--V. magkasalubonga (Barrion & Litsinger 1995) (see below for synonymy) F06
    |--V. mckayi Framenau & Vink 2001 F06
    |--V. ornatula (Koch 1877) [=Lycosa ornatula, Allocosa ornatula; incl. V. forsteri Framenau & Vink 2001] F06
    |--V. palau Framenau 2006 F06
    |--V. penola Framenau & Vink 2001 F06
    |--V. pictiventris (Koch 1877) F06 [=Lycosa pictiventris E07]
    |--V. pseudospeciosa Framenau & Vink 2001 F06
    |--V. pullastra (Simon 1909) (see below for synonymy) F06
    |--V. roo Framenau & Vink 2001 F06
    |--V. speciosa (Koch 1877) F06
    |--V. summa (McKay 1974) [=Lycosa summa] F06
    `--V. tinfos Framenau 2006 F06

Venatrix konei (Berland 1924) [=Lycosa konei, Arctosa konei; incl. L. goyderi Hickman 1944, A. goyderi, Mustelicosa goyderi, Piratosa goyderi, Venatrix goyderi, Lycosa howensis McKay 1979] F06

Venatrix magkasalubonga (Barrion & Litsinger 1995) [=Pardosa magkasalubonga; incl. P. daniloi Barrion & Litsinger 1995, P. hawakana Barrion & Litsinger 1995, P. sacayi Barrion & Litsinger 1995] F06

Venatrix pullastra (Simon 1909) [=Lycosa pullastra, Hogna pullastra, L. pallustra (l. c.); incl. L. marcentior Simon 1909, Hogna marcentior, L. percauta Simon 1909, Allocosa percauta, L. propitia Simon 1909, A. propitia, L. segregis Simon 1909, Hogna segregis] F06

*Type species of generic name indicated

REFERENCES

[E07] Elliott, M. G. 2007. Annotated catalogue of the Pompilidae (Hymenoptera) of Australia. Zootaxa 1428: 1–83.

[F06] Framenau, V. W. 2006. The wolf spider genus Venatrix Roewer: new species, synonymies and generic transfers (Araneae, Lycosidae). Records of the Western Australian Museum 23 (2): 145–166.

Artoriinae

Male Artoria cingulipes, copyright Volker Framenau.

Belongs within: Lycosidae.

The Artoriinae are a group of wolf spiders found in Australasia and insular southeast Asia, characterised by male pedipalps bearing a distinct apophysis at the base of the embolus. Members of the subfamily vary from small to large in size, and the shape of the basoembolic apophysis may vary from thin and lamellar to strongly sclerotised (Framenau 2007).

<==Artoriinae F07
    |--+--Notocosa Vink 2002 VMP02, F07
    |  |    `--*N. bellicosa (Goyen 1887) [=Pardosa bellicosa] F07
    |  `--Artoria Thorell 1877 VMP02, F08 [incl. Lycosula Roewer 1960 F07]
    |       |--*A. parvula Thorell 1877 F08
    |       |--A. albopilata (Urquhart 1893) F07
    |       |--A. cingulipes Simon 1909 F08
    |       |--A. flavimana Simon 1909 F08
    |       |--A. hebridisiana (Berland 1938) [=Lycosula hebridisiana] F07
    |       |--A. hospita Vink 2002 PVD10
    |       |--A. impedita (Simon 1909) FM09 [=Lycosa impedita F07, Trochosa expolita impedita F07]
    |       |--A. linnaei Framenau 2008 F08
    |       |--A. minima (Berland 1938) [=Lycosula minima] F07
    |       |--A. segrega Vink 2002 PVD10
    |       |--A. separata Vink 2002 PVD10
    |       `--A. thorelli (Berland 1929) [=*Lycosula thorelli] F07
    `--Anoteropsis Koch 1878 VMP02, F07
         |  i. s.: *A. flavescens Koch 1878 F07
         |         A. aerescens (Goyen 1887) PVD10
         |         A. alpina Vink 2002 PVD10
         |         A. arenivaga (de Dalmas 1917) PVD10
         |         A. blesti Vink 2002 PVD10
         |         A. canescens (Goyen 1887) PVD10
         |         A. cantuaria Vink 2002 PVD10
         |         A. forsteri Vink 2002 PVD10
         |         A. hallae Vink 2002 PD10
         |         A. hilaris (Koch 1877) PVD10
         |         A. insularis Vink 2002 PVD10
         |         A. litoralis Vink 2002 PVD10
         |         A. montana Vink 2002 PVD10
         |         A. okatainae Vink 2002 PVD10
         |         A. ralphi (Simon 1905) PVD10
         |         A. urquharti (Simon 1898) PVD10
         |         A. westlandica Vink 2002 PVD10
         |--A. senica (Koch 1877) VMP02, PVD10
         `--+--A. adumbrata (Urquhart 1887) VMP02
            `--A. lacustris Vink 2002 VMP02

Artoriinae incertae sedis:
  Artoriopsis Framenau 2007 F07
    |--*A. expolita (Koch 1877) (see below for synonymy) F07
    |--A. anacardium Framenau 2007 F07
    |--A. eccentrica Framenau 2007 F07
    |--A. joergi Framenau 2007 F07
    |--A. klausi Framenau 2007 F07
    |--A. melissae Framenau 2007 F07
    `--A. whitehouseae Framenau 2007 F07
  Diahogna Roewer 1960 F07
    `--*D. martensii (Karsch 1878) [=Lycosa martensii] F07
  Lycosella Thorell 1890 F07
    `--*L. tenera Thorell 1890 F07
  Syroloma Simon 1900 F07
    `--*S. major Simon 1900 F07
  Tetralycosa Roewer 1960 F07
    |--T. oraria (Koch 1877) FL13 [incl. Lycosa meracula Simon 1909 FL13, *T. meracula F07]
    `--T. eyrei F07
  ‘Trochosa’ alboguttulata (Koch 1878) F07
  ‘Trochosa’ tristicula (Koch 1877) F07
  ‘Lycosa’ properipes Simon 1909 F07

*Artoriopsis expolita (Koch 1877) [=Lycosa expolita, Avicosa expolita, Schizocosa expolita, Trochosa expolita] F07

*Type species of generic name indicated

REFERENCES

[F07] Framenau, V. W. 2007. Revision of the new Australian genus Artoriopsis in a new subfamily of wolf spiders, Artoriinae (Araneae: Lycosidae). Zootaxa 1391: 1–34.

[F08] Framenau, V. W. 2008. A new wolf spider species of the genus Artoria from Western Australia (Araneae: Lycosidae). Records of the Western Australian Museum 24 (4): 363–368.

[FL13] Framenau, V. W., & A. E. Leung. 2013. Costacosa, a new genus of wolf spider (Araneae, Lycosidae) from coastal north-west Western Australia. Records of the Western Australian Museum Supplement 83: 173–184.

[FM09] Framenau, V. W., B. Y. Main, M. S. Harvey & J. M. Waldock. 2009. Tapetosa, a new monotypic wolf spider genus from Western Australia (Araneae: Lycosidae: Lycosinae). Records of the Western Australian Museum 25 (3): 309–314.

[PVD10] Paquin, P., C. J. Vink & N. Dupérré. 2010. Spiders of New Zealand: annotated family key and species list. Manaaki Whenua Press: Lincoln (New Zealand).

[VMP02] Vink, C. J., A. D. Mitchell & A. M. Paterson. 2002. A preliminary molecular analysis of phylogenetic relationships of Australasian wolf spider genera (Araneae, Lycosidae). Journal of Arachnology 30 (2): 227–237.

Last updated: 22 January 2022.