Systasis

Female Systasis, from Bouček (1988).

Belongs within: Ormocerinae.

Systasis is a cosmopolitan genus of wasps of uncertain biology: they may parasitise insect larvae within galls, or they may be gall-formers themselves (Bouček 1988).

Characters (from Bouček 1988): Body with more or less distinct metallic tinge; antenna with seven segments between pedicel and clava; thoracic dorsum usually lacking coarse punctures, usually with dense reticulation (and then dull) or very dense but fine puncturation; propodeum deeply reticulate; forewing with rather sparse pilosity, with a large speculum anteriorly reaching virtually the distal end of the marginal vein, thus making conspicuous a row of longish hairs on the underside, below the marginal vein; marginal vein more than twice as long as stigmal, postmarginal vein usually only slightly longer than the stigmal; legs usually more or less infuscate.

<==Systasis Walker 1834 [incl. Guieralia Risbec 1951, Paruriella Girault 1913, Systasina Bouček 1955] B88
    |--*S. encyrtoides B88
    |--S. australiensis (Girault 1913) [=Paruriella australiensis] B88
    |--‘Miscogaster’ bato Walker 1839 B88
    |--S. cecidomyiae (Ashmead 1900) [=Asemantus cecidomyiae] B88
    |--S. cecili Girault 1929 B88
    |--S. cernus (Walker 1839) [=Miscogaster cernus, Seladerma cernum] B88
    |--S. dasyneurae A71
    |--S. dice (Walker 1839) [=Miscogaster dice, Semiotellus dice, Semiotus dice] B88
    |--S. doddi (Girault 1915) [=Schizonotus doddi] B88
    |--S. euctemon (Walker 1839) [=Pteromalus euctemon] B88
    |--S. flindersiae Girault 1934 G34
    |--S. graminis (Cameron 1912) (see below for synonymy) B88
    |--S. grotiusi Girault 1915 B88
    |--S. henrici Girault 1913 B88
    |--S. horridula (Girault 1915) [=Paruriella horridula] B88
    |--S. insularis Dodd & Girault in Girault 1915 B88
    |--S. keatsi Girault 1927 B88
    |--S. lelex (Walker 1839) [=Pteromalus lelex] B88
    |--‘Miscogaster’ letus Walker 1839 B88
    |--S. merula (Walker 1839) [=Miscogaster merula, Semiotellus merula, Semiotus merula] B88
    |--S. persimilis (Dodd & Girault in Girault 1915) [=Paruriella persimilis] B88
    |--S. punctativertex Girault 1915 B88
    |--S. seposita Girault 1915 [=S. sepositus] B88
    `--S. varipes Girault 1915 B88

Systasis graminis (Cameron 1912) [=Callimome graminis, Paruriella graminis; incl. C. reticulatus Cameron 1912, Systasis reticulata] B88

*Type species of generic name indicated

REFERENCES

Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

Diparinae

Lelaps striatus, from here.

Belongs within: Chalcidoidea.

The Diparinae are a group of chalcidoid wasps found primarily in leaf litter. Many species are strongly sexually dimorphic: males are usually macropterous, but females may be brachypterous or apterous (Desjardins 2007).

Characters (from Bouček 1988, Desjardins 2007): Antenna 13-segmented. Scutellum varied, but never smooth and convex. Metacoxa inserted unusually high, usually with transverse striations on posterior margin. Gastral tergite 1 usually expanded to cover at least half of metasoma. Cercal brush present. Females often apterous or brachypterous.

<==Diparinae [Diparides, Diparini, Lelapinae]
    |--+--Chimaerolelaps Desjardins 2007 D07
    |  |    `--*C. villosa Desjardins 2007 D07
    |  `--Dipara D07
    `--+--Netomocera Bouček 1954 D07, B88 [Netomocerini]
       |    |--*N. setifera Bouček 1954 D07
       |    |--N. africana Hedqvist 1971 D07
       |    |--N. alboscapus Hedqvist 1971 D07
       |    |--N. nearctica Yoshimoto 1977 D07
       |    |--N. nigra Sureshan & Narendran 1990 D07
       |    |--N. rufa Hedqvist 1971 D07
       |    `--N. sedlaceki Bouček 1988 B88
       `--+--Lelaps Walker 1843 (see below for synonymy) D07
          |    |--*L. pulchricornis [=*Dilaelaps pulchricornis, *Stenopistha pulchricornis] D07
          |    |--L. abdominalis Ashmead 1904 D07
          |    |--L. aeneiceps Ashmead 1904 D07
          |    |--L. affinis Ashmead 1904 D07
          |    |--L. albipes Cameron 1884 D07
          |    |--L. albofasciatus Hedqvist 1964 D07
          |    |--L. annulicornua (Strand 1911) [=Stenopistha annulicornua] D07
          |    |--L. apicalis Ashmead 1904 D07
          |    |--L. argenticoxa Girault 1916) [=*Spalangiolaelaps argenticoxa] D07
          |    |--L. avicula Haliday 1844 D07
          |    |--L. beckeri Yoshimoto 1977 D07
          |    |--L. bimaculata Ashmead 1904 D07
          |    |--L. callisto Marshall in Buysson & Marshall 1892 [=Laelaps (l. c.) callisto] D07
          |    |--L. caudatula (Strand 1911) [=Stenopistha caudatula] D07
          |    |--L. decorata Walker 1862 [=Laelaps (l. c.) decorata] D07
          |    |--L. ferrierei Hedqvist 1964 D07
          |    |--L. ferruginea Cameron 1884 D07
          |    |--L. flagellata (Strand 1911) [=Stenopistha flagellata] D07
          |    |--L. flavescens Ashmead in Riley et al. 1894 D07
          |    |--L. floridensis Yoshimoto 1977 D07
          |    |--L. halidayi Ashmead 1904 [incl. Stenopistha halidayi sobrina Strand 1911] D07
          |    |--L. insularis Mercet 1927 D07
          |    |--L. magnifica (Strand 1911) [=Stenopistha magnifica] D07
          |    |--L. melinus Yoshimoto 1977 D07
          |    |--L. noortii Desjardins 2007 D07
          |    |--L. ornata (Strand 1911) [=Stenopistha ornata] D07
          |    |--L. paraguayensis Girault 1912 D07
          |    |--L. picta Walker 1862 [=Laelaps (l. c.) picta] D07
          |    |--L. pulchella (Strand 1911) [=Stenopistha pulchella] D07
          |    |--L. pulchra Girault 1912 D07
          |    |--L. pulchricornis Walker 1843 D07
          |    |--L. pygata (Strand 1911) [=Stenopistha pygata; incl. S. pygata conjuncta Strand 1911] D07
          |    |--L. rectivitta (Strand 1911) [=Stenopistha rectivitta] D07
          |    |--L. rhomboidea (Strand 1911) [=Stenopistha rhomboidea] D07
          |    |--L. sadales (Walker 1839) [=Merostenus sadales] D07
          |    |--L. setifrons (Strand 1911) [=Stenopistha setifrons] D07
          |    |--L. simplex (Fabricius 1804) [=Chalcis simplex] D07
          |    |--L. striaticeps (Strand 1911) [=Stenopistha striaticeps] D07
          |    |--L. striatus Yoshimoto 1977 D07
          |    |--L. stylata Ashmead 1904 D07
          |    |--L. tapajoana (Schulz 1906) [=Dilaelaps tapajoana, L. ferruginea Ashmead 1904 non Cameron 1884] D07
          |    |--L. terebrans (Strand 1911) [=Stenopistha terebrans] D07
          |    |--L. viridiceps (Strand 1911) [=Stenopistha viridiceps] D07
          |    `--L. vittipennis (Strand 1911) [=Stenopistha vittipennis] D07
          `--+--Dozodipara Desjardins 2007 D07
             |    `--*D. insularis Desjardins 2007 D07
             `--+--Boeria Hedqvist 1969 D07
                |    `--*B. saetosa Hedqvist 1969 D07
                `--+--Cerodipara Desjardins 2007 D07
                   |    `--*C. sabensis Desjardins 2007 D07
                   `--+--+--Conodipara Hedqvist 1972 [=Turneria Hedqvist 1969 non Forel 1895] D07
                      |  |    `--*C. scutellata (Hedqvist 1969) [=*Turneria scutellata] D07
                      |  `--+--Conophorisca Hedqvist 1969 D07
                      |     |    |--*C. annulata Hedqvist 1969 D07
                      |     |    |--C. grisselli Desjardins 2007 D07
                      |     |    `--C. littoriticus Desjardins 2007 D07
                      |     `--Myrmicolelaps Hedqvist 1969 [incl. Dolichodipara Hedqvist 1969] D07
                      |          |--+--*M. paradoxus Hedqvist 1969 D07
                      |          |  `--M. aurantius Desjardins 2007 D07
                      |          `--+--M. iridius Desjardins 2007 D07
                      |             `--M. scutellata (Hedqvist 1969) [=*Dolichodipara scutellata] D07
                      `--+--Pyramidophoriella Hedqvist 1969 D07
                         |    |--*P. albiclava Hedqvist 1969 D07
                         |    `--P. brunnea Hedqvist 1969 D07
                         `--+--Nosodipara Bouček 1988 D07
                            |    |--*N. monteithorum Bouček 1988 D07
                            |    `--N. ferrana Desjardins 2007 D07
                            `--Pseudoceraphron Dodd 1924 [incl. Dipareta Bouček 1988, Malinka Bouček 1988] D07
                                 |--P. regieri Desjardins 2007 D07
                                 `--+--P. albifrons (Bouček 1988) [=*Dipareta albifrons] D07
                                    `--+--+--*P. pulex Dodd 1924 D07
                                       |  `--P. burwelli Desjardins 2007 D07
                                       `--+--P. fijiensis Desjardins 2007 D07
                                          `--P. nana (Bouček 1988) [=*Malinka nana] D07

Lelaps Walker 1843 [=Laelaps (l. c.), Dilaelaps Schulz 1906; incl. Spalangiolaelaps Girault 1916, Stenopistha Strand 1910] D07

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[D07] Desjardins, C. A. 2007. Phylogenetics and classification of the world genera of Diparinae (Hymenoptera: Pteromalidae). Zootaxa 1647: 1-88.

Terobiella

Antenna of male Terobiella nigriceps, from Bouček (1988).

Belongs within: Ormocerinae.

Terobiella is an Australian genus of wasps causing galls on leaves of Eucalyptus (Bouček 1988).

Characters (from Bouček 1988): Body compact, non-metallic. Antennae 12-segmented, with seven segments between pedicel and clava, bearing long branches in the males, females with most funicular segments dorsally only slightly produced appearing serrate. Scutellum anteriorly pointed, axillae converging medially.

<==Terobiella Ashmead 1900 (see below for synonymy) B88
    |--*T. flavifrons Ashmead 1900 [incl. *Melanosomella flavipes Girault 1913] B88
    |--T. adolphi (Girault 1931) [=Perilampoides adolphi] B88
    |--T. bicolor (Girault 1913) [=*Perilampoides bicolor] B88
    |--T. cinctitibiae (Girault 1931) [=Perilampoides cinctitibiae] B88
    |--T. dilutiventris (Girault 1913) [=Epiperilampus dilutiventris] B88
    |--T. eucalypti (Gallard 1930) [=Tepperella eucalypti] B88
    |--T. nigriceps (Ashmead 1904) (see below for synonymy) B88
    |--T. particolor (Girault 1915) [=Perilampoides particolor] B88
    |--T. regalis (Girault 1931) [=Perilampoides regalis] B88
    |--T. scutata (Girault 1931) [=Perilampoides scutatus] B88
    |--T. similis (Girault 1917) [=Perilampoides similis] B88
    `--T. tennysoni (Girault 1920) [=Perilampoides tennysoni] B88

Terobiella Ashmead 1900 [incl. Melanosomella Girault 1913, Paraterobia Ashmead 1904, Perilampoides Girault 1913, Tepperella Cameron 1912] B88

Terobiella nigriceps (Ashmead 1904) [=*Paraterobia nigriceps; incl. Perilampoides aurantiscutum anna Girault 1931, Pe. aurantiscutum Girault 1916, Pe. flavithorax Girault 1915, *Tepperella maculiscutis Cameron 1912, Pe. aurantiscutum regis Girault 1931, Pe. aurantiscutum rex Girault 1931] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

Trichilogaster

Female Trichilogaster signiventris, photographed by John Hoffman.

Belongs within: Ormocerinae.

Trichilogaster is a genus of wasps forming galls on Acacia species (Bouček 1988).

Characters (from Bouček 1988): Body robust, often partly rufous or yellowish brown, with thorax short and convex, sometimes broader than head, always sculptured, sometimes rather coarsely. Head with broadly concave occiput and convex frons; scrobes not large but sometimes reaching ocellus. Clypeus produced, bilobate. Antennae at centre of face or slightly above, formula 11263; anelli large, flagellum in female subfiliform. clava symmetric. Pronotum nearly as broad as scutum, medially short; notauli narrow but deep; axillae slightly advanced, medially almost meeting. Scutellum slightly oblong, with admarginai groove, apex closely applied to raised middle of dorsellum (in side view). Propodeum often with longitudinal rugae or at least with weak median carina and curved plicae posteriorly. Hind tibia with 2 spurs. Forewing with veins rather short, the marginal and base of postmarginal slightly thickened, stigma often knobbed; often infumation present below parastigma. Gaster short and high but collapsing rather irregularly; hind margins of tergites 1 and 2 emarginate, exposing large third tergite, this often with coarse sculpture.

<==Trichilogaster Mayr 1906 [incl. Epiperilampus Girault 1913] B88
    |--*T. maideni (Froggatt 1892) [=Cynips maideni] B88
    |--T. acaciaelongifoliae (Froggatt 1892) [=Cynips acaciaelongifoliae, T. longifoliae Mayr 1905] B88
    |--T. arabica B88
    |--T. atra (Girault 1915) [=Epiperilampus ater] B88
    |--T. atricorpus (Girault 1931) [=Epiperilampus atricorpus; incl. E. atricorpus socratis Girault 1931] B88
    |--T. channingi (Girault 1913) [=Epiperilampus channingi] B88
    |--T. esculenta Ferrière in Gahan & Ferrière 1947 [=T. esculentus] B88
    |--T. flavivena (Girault 1931) [=Coelocybelloides flavivena] B88
    |--T. pendulae Mayr 1905 B88
    |--T. significata (Girault 1913) [=Epiperilampus significatus] B88
    |--T. signiventris (Girault 1931) (see below for synonymy) B88
    `--T. xanthocephala (Girault 1913) [=*Epiperilampus xanthocephalus] B88

Trichilogaster signiventris (Girault 1931) [=Perilampella signiventris; incl. Epiperilampus adolphi Girault 1931, Trichilogaster adolphi, Coelocybelloides atraticorpus Girault 1931] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

Encyrtocephalus

Female Encyrtocephalus albiclava, from Bouček (1988).

Belongs within: Ormocerinae.

Encyrtocephalus is a genus of wasps found in Africa and Australia that cause galls in trees such as Acacia, Eucalyptus and Brachychiton (Bouček 1988).

Characters (from Bouček 1988): Antenna with eight segments between pedicel and clava, first two segments of flagellum subequal in size, shorter than following ones; upper margin of occiput sometimes carinate or edged, rarely rounded; body dull, without metallic gloss; thorax with conspicuous pilosity and coarse puncturation (latter sometimes less conspicuous in small specimens); prepectus large and broadly triangular; forewing often without infumation, sometimes infumate just at stigma, sometimes with double or almost treble infumation; fringe well developed; stigmal vein strongly arched, at its perpendicular base more or less triangularly thickened, its base and apex usually darkened, distinctly shorter than marginal vein; gaster usually sessile, very rarely on conspicuous petiole.

<==Encyrtocephalus Ashmead 1900 (see below for synonymy) B88
    |--*E. simplicipes Ashmead 1900 B88
    |--E. albiclava (Girault 1915) [=Neorileyella (*Pseudorileya) albiclava] B88
    |--E. albipilum Girault 1931 B88
    |--E. atrativentris Girault 1931 B88
    |--E. fasciatus (Girault 1915) [=*Neorileyella fasciata] B88
    |--E. gallicola (Ashmead 1904) [=*Decatomothorax gallicola] B88
    |--E. hyalinus (Girault 1915) [=Neorileyella hyalina] B88
    |--E. io Girault 1931 B88
    |--E. katrina (Girault 1931) [=Neorileyella katrina] B88
    |--E. mozarti Girault 1931 B88
    |--E. notabilis [=*Eucyrtonotus notabilis] B88
    `--E. ophelia (Girault 1931) [=Neorileyella ophelia] B88

Encyrtocephalus Ashmead 1900 [incl. Decatomothorax Ashmead 1904, Encyrtonotus (l. c.), Eucyrtonotus Silvestri 1915, Neorileyella Girault 1915, Pseudorileya Girault 1915] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

Colotrechninae

Ovipositing Cameronella, from here.

Belongs within: Chalcidoidea.

The Colotrechninae are a group of chalcidoid wasps found in Australia and South America. Host data are known for only a small number of species, but colotrechnines include species associated with plant galls and seed pods (Bouček 1988).

Characters (from Bouček 1988): Occipital carina absent; large axillulae present, separated from scutellum by (mostly) outcurving grooves or lines visible dorsally.

<==Colotrechninae [Colotrechnides] B88
    |--Uzka Bouček 1988 [Uzkini] B88
    |    `--*U. bethyloides Bouček 1988 B88
    |--Divna Bouček 1988 [Divnini] B88
    |    `--*D. hirtura Bouček 1988 B88
    |--Amerostenini B88
    |    |--Errolia Bouček 1988 B88
    |    |    `--*E. cyanea Bouček 1988 B88
    |    |--Glorimontana Bouček 1988 B88
    |    |    `--*G. flaveola Bouček 1988 B88
    |    |--Omphalodipara Girault 1923 [incl. Froudeana Girault 1928] B88
    |    |    |--*O. splendida Girault 1923 B88
    |    |    `--O. clavicornis (Dodd 1924) [=Hetreulophus clavicornis, *Froudeana clavicornis] B88
    |    `--Amerostenus Girault 1913 B88
    |         |--*A. australiensis Girault 1913 B88
    |         |--A. aereipes Girault 1915 B88
    |         |--‘Miscogaster’ athanis Walker 1839 B88
    |         |--A. theope (Walker 1839) [=Miscogaster theope, Semiotellus theope, Semiotus theope] B88
    |         |--A. tridentatus Girault 1928 B88
    |         `--A. varidentatus Girault 1929 B88
    `--Colotrechnini [Cremneulophini, Dvaliniinae] B88
         |--Bofuria Hedqvist 1978 B88
         |--Bomburia Hedqvist 1978 B88
         |--Dvalinia Hedqvist 1977 B88
         |--Elachertodomyia Girault 1917 B88
         |--Pachyneuronella Girault 1913 B88
         |    `--*P. viridis Girault 1913 B88
         |--Uriellopteromalus Girault 1915 B88
         |    |--*U. subplanithorax Girault 1915 B88
         |    `--U. aeneus Bouček 1988 B88
         |--Colotrechnus Thomson 1878 [incl. Zanonia Masi 1921] B88
         |    |--*C. subcoeruleus B88
         |    |--C. agromyzae B88
         |    |--C. ignotus B88
         |    |--C. notalauris Bouček 1988 B88
         |    `--C. viridis B88
         `--Cameronella Dalla Torre 1898 (see below for synonymy) B88
              |--*C. blackburni (Cameron 1888) [=*Panthalis blackburni] B88
              |--C. apiomorphae (Girault 1939) [=Dinoura apiomorphae] B88
              |--C. auriventris (Ashmead 1900) [=*Dinoura auriventris] B88
              |--C. cyanea (Ashmead 1900) [=Dinoura cyanea] B88
              |--C. eucalypti (Girault 1917) [=Dinoura eucalypti] B88
              |--C. gigantea (Girault 1916) [=*Cremneulophus giganteus] B88
              `--C. pulchra (Girault 1927) [=Dinoura pulchra] B88

Cameronella Dalla Torre 1898 [=Panthalis Cameron 1888 non Kinberg 1854;incl. Cremneulophus Girault 1916, Dinoura Ashmead 1900] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

Cleonymus

Unidentified Cleonymus, photographed by Kim Fleming.

Belongs within: Cleonyminae.

Cleonymus is a near-cosmopolitan genus of wasps (unrecorded from South America), parasitic on wood-boring beetles (Bouček 1988).

Characters (from Bouček 1988): Antenna with eight distinct segments between pedicel and clava; male antenna unbranched, with almost symmetrical segments covered by regular dense pilosity; preclava in female with distinct lateral process closely applied to side of clava; notaular grooves becoming indistinct in very dense reticulation-puncturation on posterior half of scutum; stigmal vein in both sexes fairly long, at very low angle, fully twice as long as distance of stigmal knob from postmarginal vein; hind femur often serrate or with a subapical tooth on ventral margin; gaster in female sessile.

<==Cleonymus Latreille 1809 (see below for synonymy) B88
    |--*C. depressus [=Diplolepis depressa] B88
    |--C. californicus Girault 1917 GM79
    |--C. dentatifemur (Girault 1926) [=Aplatygerrhus dentatifemur] B88
    |--C. imperialis (Dodd 1924) [=Aplatygerrhus imperialis] B88
    |--C. laticinctus (Girault 1926) [=Systolomorphella laticincta] B88
    |--C. magnificus (Ashmead 1888) [=Charitopus magnificus, *Ptinobius magnificus] B88
    |--C. magnus Bouček 1988 [=*Aplatygerrhus magnificus Girault 1913 non Charitopus magnificus Ashmead 1888] B88
    |--C. nigriclavus Girault 1917 G17
    |--C. pentlandi (Girault 1922) [=Systolomorphella pentlandi] B88
    |--C. regalis (Dodd 1924) [=Aplatygerrhus regalis] B88
    |--C. rufiscapus (Girault 1925) [=Aplatygerrhus rufiscapus] B88
    |--C. silvifilia (Girault 1927) [=Systolomorphella silvifilia] B88
    |--C. trifasciatipennis (Girault 1915) [=*Systolomorphella trifasciatipennis] B88
    |--C. unfasciatipennis (Dodd & Girault in Girault 1915) [=Systolomorphella unfasciatipennis] B88
    |--C. unnotipennis (Girault 1915) [=Systolomorphella unnotipennis] B88
    `--‘*Beharella’ viridicyanea B88

Cleonymus Latreille 1809 [incl. Aplatygerrhus Girault 1913, Beharella Risbec 1952, Ptinobius Ashmead 1896, Systolomorphella Girault 1915] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[G17] Girault, A. A. 1917. Descriptiones hymenopterorum chalcidoidicarum variorum cum observationibus. III. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 102-111).

[GM79] Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 1-400.

Cleonyminae

Heydenia pretiosa, from Entomology WSL.

Belongs within: Chalcidoidea.
Contains: Cleonymus.

The Cleonyminae was originally established for pteromalid wasps with a large conical pronotum. Most species are parasites of beetle larvae in wood (Bouček 1988).

Characters (from Bouček 1988): Occipital carina indistinct; pronotum large and subconical, dorsally rounded or at least without transverse edge, medially often with smooth strip or raised longitudinal line; hind margin only moderately emarginate; thorax usually punctured; body without long sparse hairs; scutellum without sublateral grooves, without distinct frenal line or frenal bristles; propodeum without deep petiolar emargination.

<==Cleonyminae [Cleonymidae, Cleonymides] B88
    |  i. s.: Agrilocida B88
    |         Bouckekius D07
    |--Heydeniini B88
    |    |--Heydeniopsis B88
    |    `--Heydenia Förster 1856 (see below for synonymy) B88
    |         |--*H. pretiosa B88
    |         |--H. burgeoni [=Lycisca burgeoni, *Risbecisca burgeoni] B88
    |         |--H. cristatipennis (Girault 1924) [=Paraheydenia cristatipennis] B88
    |         |--H. longicollis (Cameron 1912) [=*Paraheydenia longicollis] B88
    |         |--‘*Heydenisca’ mateui B88
    |         |--‘*Pterooderella’ ornata B88
    |         `--H. trinodis Bouček 1988 B88
    `--Cleonymini B88
         |  i. s.: Amarisca Delucchi 1962 B88
         |         Pannoniella Erdös 1960 B88
         |         Paracleonymus B88
         |         Notanisus Walker 1837 B88 [incl. Bekiliella Risbec 1952 D07]
         |           |--*N. versicolor B88
         |           |--N. cyanea (Risbec 1952) [=*Bekiliella cyanea] D07
         |           |--N. richteri (Girault 1922) [=Systolomorphella richteri; incl. S. dinotipennis Girault 1922] B88
         |           `--N. zebra Bouček 1988 B88
         |         Zolotarewskya Risbec 1955 [incl. Pseudocleonymus Hedqvist 1967] B88
         |           |--*Z. seyrigi B88
         |           |--Z. shakespearei (Girault 1926) [=Systolomorphella shakespearei] B88
         |           `--‘*Pseudocleonymus’ spinifer B88
         |--+--Cleonymus MH11
         |  `--Dasycleonymus MH11
         `--Callocleonymus Masi 1940 MH11, B88
              |--*C. pulcher B88
              |--C. ferrierei Kerrich 1957 B88
              `--C. swezeyi (Yoshimoto & Ishii 1965) [=Ptinobius swezeyi] B88

Heydenia Förster 1856 [incl. Heydenisca Hedqvist 1967, Paraheydenia Cameron 1912, Pterooderella Risbec 1952, Risbecisca Hedqvist 1960] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[D07] Desjardins, C. A. 2007. Phylogenetics and classification of the world genera of Diparinae (Hymenoptera: Pteromalidae). Zootaxa 1647: 1-88.

[GM79] Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 1-400.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.

Parepistenia

Dorsal view of body of female Parepistenia pulchripes, from Bouček (1988).

Belongs within: Chalcidoidea.

Parepistenia is a primarily Southern Hemisphere genus of chalcidoid wasps. Specific hosts are as yet unknown, though some species have been raised from logs containing beetle borings (Bouček 1988).

Characters (from Bouček 1988): Eyes short, rounded-subtriangular, at most nearly 1.4 times as long as broad; antenna with eight distinct segments between pedicel and clava; occipital margin rounded; lower face rather short inc1uding the converging genae, regularly convex between antennal socket and lower end of eye; gena without strong lateral carina; notaular grooves complete; epipygium of gaster short, with cerci at base; ovipositor not produced.

<==Parepistenia Dodd in Girault 1915 [incl. Austrogerrhus Bouček 1958] B88
    |--*P. varicornis Dodd in Girault 1915 [incl. Epistenia aedicule Girault 1927] B88
    |--P. annulicornis (Girault 1913) [=Platygerrhus annulicornis] B88
    |--P. arenae (Girault 1932) [=Thaumasura arenae] B88
    |--P. australiensis (Girault 1913) [=Platygerrhus australiensis] B88
    |--P. camilli (Girault 1925) [=Epistenia camilli] B88
    |--P. corticis (Girault 1925) [=Epistenia corticis] B88
    |--P. devannyi (Girault 1937) [=Epistenia devannyi] B88
    |--P. froudei (Girault 1929) [=Platygerrhus froudei] B88
    |--P. fulviventris (Girault 1915) [=Platygerrhus fulviventris] B88
    |--P. incola (Girault 1929) [=Platygerrhus incola] B88
    |--P. inusitata (Girault 1937) [=Epistenia inusitata] B88
    |--P. miripes (Girault 1922) [=Epistenia miripes] B88
    |--P. nigriaenea (Girault 1915) [=Epistenia nigriaenea] B88
    |--P. pallidicoxa (Girault 1929) [=Platygerrhus pallidicoxa] B88
    |--P. pulchripes (Girault 1927) [=Thaumasura pulchripes; incl. *Austrogerrhus gloriosus Bouček 1958] B88
    |--P. resplendens (Gourlay 1928) [=Thaumasura resplendens] B88
    |--P. speciosissima (Girault 1927) [=Epistenia speciosissima] B88
    `--P. tasmaniensis (Girault 1913) [=Platygerrhus tasmaniensis] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

Chromeurytominae

Male Chromeurytoma, from John Heraty.

Belongs within: Chalcidoidea.

The Chromeurytominae are an Australian group of chalcidoid wasps associated with plant gall-makers and leaf-miners.

Characters (from Bouček 1988): Antennal flagellum with just six segments between pedicel and clava; the first of which may be anelliiform, short, but not a true anellus. Female clava at least slightly asymmetric, with more or less oblique sutures and a collapsing area of weaker sclerotisation and with special short trichoid sensilla. Scrobal cavity evident but not well delimited, ending far from ocellus; in lower part divided by an interantennal crest. Parascrobal areas convex. Antennae inserted below lower ocular line, at the convex but not well defined clypeus, the lower margin of which is arcuately produced and stout. Both mandibles 3-toothed. Malar groove ending at a finely margined and sometimes slightly depressed mouth border. Occipital carina present, of torymid form, but occasionally obsolete (in one species). Surface of body generally shiny, mostly shallowly cross-striate to smooth, mostly sparsely pilose (hairs on papillae), mostly with metallic gloss but in at least two species the gloss is absent. Pronotum fairly large and quadrangular, anteriorly often edged or carinate. Scutum rather short and broad, with deep complete notauli; frenal line distinct but weak, situated far before the apex of the scutellum. Mesopleuron bearing peculiar pit-like depressions. Propodeum bearing medially a large, depressed, subtriangular to cordiform area, which is often divided by a median carina; median area is delimited on sides by converging channels marking on inner side the convex smooth plical swellings which laterally merge with postspiracular sulci; posterior to the median area is a very short nuchal ring set off by a transverse groove. Mid tibia with a rather long spur, often longer than the basitarsus; hind tibia with two spurs. In the forewing the stigmal vein has a distinct uncus, the postmarginal vein is often at least as long as the marginal. Gaster on a mostly visible petiole, basal three tergites large, third usually larger than second; gastral body is strongly convex. Ovipositor sheaths are usually distinctly protruding, up to as long as gaster. Cerci always elongate.

<==Chromeurytominae B88
    |--Asaphoideus Girault 1913 B88
    |    `--*A. niger Girault 1913 B88
    `--Chromeurytoma Cameron 1912 (see below for synonymy) B88
         |--*C. clavicornis Cameron 1912 B88
         |--C. beerwahi (Girault 1926) [=Amonodontomerus beerwahi] B88
         |--C. emersoni (Girault 1915) [=Holasaphes emersoni] B88
         |--C. gregi (Girault 1915) [=*Holasaphes gregi] B88
         |--C. iucunda (Girault 1921) [=Amonodontomerus iucundus] B88
         |--C. megastigmoides (Girault 1926) [=Amonodontomerus megastigmoides] B88
         |--C. miltoni (Girault 1931) [=*Euryperilampus miltoni] B88
         |--C. montana (Girault 1929) [=Amonodontomerus montanus] B88
         |--C. mycon (Walker 1839) [=Miscogaster mycon] B88
         |--C. nelo (Walker 1839) [=Miscogaster nelo] B88
         |--C. noblei (Girault 1940) [=Amonodontomerus noblei] B88
         |--C. poeta (Girault 1939) [=Amonodontomerus poeta] B88
         |--C. silvae (Girault 1921) [=Amonodontomerus silvae] B88
         `--C. viridis (Girault 1913) [=*Amonodontomerus viridis] B88

Chromeurytoma Cameron 1912 [incl. Amodontomerus Girault 1913, Euryperilampus Girault 1931, Holasaphes Girault 1915] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

Agamerion

Male Agamerion, from CSIRO Australia.

Belongs within: Chalcidoidea.

Agamerion is an Australasian genus of wasps found on smooth eucalypt trunks; larvae have been raised from cockroach eggs placed under eucalypt bark. Members of the genus have deep, carinate, inverted V-shaped scrobes, a short pronotum and a non-protruding ovipositor (Bouček 1988).

<==Agamerion Haliday 1844 [incl. Euchrysia Westwood 1874] B88
    |--*A. gelo (Walker 1839) [=Miscogaster gelo; incl. Euchrysia gemma Westwood 1874] B88
    |--A. cleptideum (Westwood 1874) [=*Euchrysia cleptidea] B88
    |--A. coeruleiventre Ashmead 1900 [=A. coeruleiventris] B88
    |--A. eupelmoides Girault 1925 B88
    |--A. io Girault 1935 B88
    |--A. metallicum Girault 1915 [=A. metallica] B88
    |--A. mirum Girault 1927 B88
    |--A. monodon Bouček 1988 B88
    |--A. nonstylatum (Girault 1922) [=Thaumasura nonstylata] B88
    |--A. omnicyaneum (Girault 1932) [=Thaumasura omnicyanea] B88
    |--A. prasinum (Westwood 1874) [=Euchrysia prasina] B88
    |--A. semialbicorne Girault 1927 [=A. semialbicornis] B88
    `--A. variflagellum Girault 1925 B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

Miscogasterini

Male Halticoptera laevigata, photographed by Thomas Hoffmeister.

Belongs within: Pteromalinae.

The Miscogasterini are a group of parasitic wasps whose known hosts are Diptera burrowing in or mining soft tissues of plants. The group is somewhat poorly defined, but its members generally have complete notauli and a petiolate gaster (Bouček 1988).

<==Miscogasterini [Sphegigasterini] MH11
    |--+--Diconocara petiolata MH11
    |  `--Sphaeripalpus MH11
    |--Cyrtogaster MH11
    |    |--C. liqueatus Ashmead 1894 GM79 (see below for synonymy)
    |    `--C. vulgaris MH11
    `--+--Lamprotatus splendens MH11
       `--+--Polstonia pelagocorypha MH11
          `--+--Thinodytes MH11
             `--Halticoptera Spinola 1811 MH11, B88 (see below for synonymy)
                  |--*H. flavicornis [=Diplolepis flavicornis] B88
                  |--H. laevigata MH11
                  `--H. stella Girault 1917 G17a

Miscogasterini incertae sedis:
  Toxeumella Girault 1913 G29, GM79
  Bairamlia B88
  Ammeia Delucchi 1962 B88
    `--*A. pulchella Delucchi 1962 B88
  Stictomischus Thomson 1876 B88
    |--*S. scaposus B88
    `--S. remotus Bouček 1988 B88
  Rhicnocoelia Graham 1956 B88
    |--*R. constans [=Pteromalus constans] B88
    `--R. incisa Bouček 1988 B88
  Miscogaster B88
    |--M. abnormicolor Girault 1917 G17a
    |--M. damia Walker 1839 (n. d.) [=Lamprotatus damia] B88
    |--M. discoloripes Girault 1917 G17a
    |--M. keatsi Girault 1917 G17b
    `--M. nicon Walker 1839 (n. d.) [=Lamprotatus nicon] B88
  Sphegigaster Spinola 1811 (see below for synonymy) B88
    |--*S. pallicornis [=Diplolepis pallicornis] B88
    |--S. agromyzae (Dodd 1917) [=Trigonogastra agromyzae] B88
    |--‘*Trigonogastra’ aurata B88
    |--S. elegantula [=*Basilewskyella elegantula] B88
    `--S. voltairei (Girault 1915) [=*Paratrigonogastra voltairei] B88

Cyrtogaster liqueatus Ashmead 1894 GM79 [incl. C. citripes Ashmead 1896 G17a, GM79, Polycyrtus floridanus Ashmead 1896 G17a, GM79, Polycystus foersteri Crawford 1913 G17a, GM79, C. occidentalis Ashmead 1896 G17a, GM79]

Halticoptera Spinola 1811 MH11, B88 [incl. Dicyclus Walker 1833 B88, Megorismus Walker 1846 B88, Pachylarthrus Westwood 1832 B88, Phacostomus Nees 1834 B88, Phagonia Curtis 1832 B88, Tityros Walker 1848 B88]

Sphegigaster Spinola 1811 [incl. Basilewskyella Risbec 1957, Paratrigonogastra Girault 1915, Trigonogastra Ashmead 1904] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[G17a] Girault, A. A. 1917a. Descriptiones hymenopterorum chalcidoidicarum variorum cum observationibus. III. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 102-111).

[G17b] Girault, A. A. 1917b. Chalcidoidea nova Marilandensis. III. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 132-137).

[G29] Girault, A. A. 1929. New pests from Australia. VI. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 266-269).

[GM79] Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 1-400.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.

Cerocephalinae

Cerocephala eccoptogastri, from here.

Belongs within: Chalcidoidea.

The Cerocephalinae are a group of wasps that parasitise small wood-boring beetles such as Scolytidae and Anobiidae (Bouček 1988).

Characters (from Bouček 1988): Antennae inserted always some distance above the mouth border, toruli separated by an interantennal crest which often forms a tooth; sometimes other ridges, often with teeth, are developed on either side of toruli; antennae rather shiny, with sparse pilosity and very low sensilla, often pale-coloured; eyes bare or nearly so; thorax frequently smooth, only rarely with rugosity, but propodeum always heavily sculptured, without smooth areas, except in dwarf forms in which all sculpture is obliterated. Forewing usually with strongly reduced discal pilosity but always with very conspicuous marginal fringe; short stigmal vein at a rather acute angle, postmarginal vein often shorter than stigmal. Gastral petiole sometimes long, sometimes strongly transverse; hind margin of first gastral terglte in females often emarginate or even deeply excised in the middle; ovipositor sheaths subexserted or longer, up to almost as long as the gaster. Short-winged forms common.

<==Cerocephalinae B88
    |--Acerocephala Gahan 1946 B88
    |    |--*A. atroviolacea [=Cerocephala atroviolacea] B88
    |    |--A. aenigma B88
    |    `--A. pacifica Bouček 1988 B88
    |--Theocolax Westwood 1832 [incl. Choetospila Westwood 1874, Spalangiomorpha Girault 1913] B88
    |    |--*T. formiciformis Westwood 1832 B88
    |    |--T. elegans (Westwood 1874) [=*Choetospila elegans; incl. *Spalangiomorpha fasciatipennis Girault 1913] B88
    |    `--T. frater (Girault 1913) [=Spalangiomorpha frater, Choetospila frater] B88
    |--Cerocephala Westwood 1832 (see below for synonymy) B88
    |    |--C. aquila (Girault 1920) [=*Proamotura aquila; incl. P. tostini Girault 1932] B88
    |    |--C. dinoderi Gahan 1925 [=C. (Parasciatheras) dinoderi; incl. Proamotura anselmi Girault 1932] B88
    |    `--C. eccoptogastri [=*Sciatherodes eccoptogastri] B88
    `--Neocalosoter Girault & Dodd in Girault 1915 [incl. Theocolaxia Girault 1924] B88
         |--*N. pulchripennis Girault 1915 B88
         |--N. grandis (Dodd 1927) [=Proamotura insularis grandis, Theocolaxia insularis grandis] B88
         |--N. insularis (Dodd 1924) [=Cratomus insularis, Proamotura insularis, Theocolaxia insularis] B88
         |--N. lessingi (Girault 1924) [=*Theocolaxia lessingi] B88
         |--N. perpulcher (Dodd 1927) (see below for synonymy) B88
         `--N. viridipronotum (Dodd 1924) (see below for synonymy) B88

Cerocephala Westwood 1832 [incl. Epimacrus Walker 1833, Proamotura Girault 1920, Prosciatheras Masi 1917, Sciatheras Ratzeburg 1848, Sciatherodes Masi 1917] B88

Neocalosoter perpulcher (Dodd 1927) [=Proamotura perpulchra, Theocolaxia perpulchra; incl. P. perpulchra metallica Dodd 1927, Theocolaxia perpulchra metallica] B88

Neocalosoter viridipronotum (Dodd 1924) [=Cratomus viridipronotum, Proamotura viridipronotum, Theocolaxia viridipronotum] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

Pteromalinae

Pachyneuron formosum, photographed by J. K. Lindsey.

Belongs within: Chalcidoidea.
Contains: Isoplatoides, Callitula, Mesopolobus, Miscogasterini, Sycoscapter, Philotrypesis, Otitesellinae, Pteromalus.

The Pteromalinae is a diverse (albeit somewhat poorly defined) group of chalcidoid wasps parasitising a wide range of host insects. The taxa listed below were treated as separate subfamilies by Bouček (1988), but these were not resolved as reciprocally monophyletic by the molecular analysis of Munro et al. (2011).

<==Pteromalinae (see below for synonymy) B88
    |--+--Pachyneuron Walker 1833 MH11, B88 (see below for synonymy)
    |  |    |--*P. formosum B88
    |  |    |--P. aphidis (Bouché 1834) [=Diplolepis aphidis, P. (Serimus) aphidis; incl. P. siphonophorae] B88
    |  |    |--P. emersoni Girault 1916 B88
    |  |    |--P. eros (Girault 1917) [=*Nepachyneuron eros] B88
    |  |    |--P. kingsleyi Girault 1916 B88
    |  |    |--P. mucronatum Girault 1917 GM79
    |  |    `--P. nelsoni Girault 1928 [incl. P. aeneus Masi 1929, Atrichoptilus aeneus] B88
    |  `--Coruna clavata MH11
    `--+--+--Cratomus [Cratominae] MH11
       |  |    `--C. megacephalus MH11
       |  `--Habritys Thomson 1878 MH11, GM79
       |       `--H. brevicornis MH11
       `--+--Miscogasterini MH11
          `--+--+--+--Sycoscapter MH11
             |  |  `--Plutothrix MH11
             |  `--+--Pegopus MH11
             |     `--Pseudocatolaccus MH11
             `--+--+--Philotrypesis B88
                |  `--Otitesellinae MH11
                |--+--Stenoselma MH11
                |  `--+--Grasseiana Abdurahiman & Joseph 1967 MH11, B88
                |     |    |--*G. callosa B88
                |     |    `--G. boschmai (Wiebes 1964) [=Sycobiella boschmai] B88
                |     `--+--Platygerrhus MH11
                |        |    |--P. affinis MH11
                |        |    `--P. columbianus [=Panstenon columbianum] B88
                |        `--Cecidostiba MH11
                |             |--C. burkei Crawford 1912 GM79
                |             `--C. fungosa MH11
                `--+--Merisus MH11
                   |    |--M. febriculosus Girault 1917 G17b
                   |    |--M. flaviventris Girault 1915 GM79
                   |    |--M. mordellistenae Crawford 1910 GM79
                   |    |--M. splendidus MH11
                   |    `--M. texensis Girault 1917 G17b
                   `--+--+--Sycoecinae MH11
                      |  |    |--Crossogaster B88
                      |  |    |--Sycoecus B88
                      |  |    |--Seres B88
                      |  |    |--Diaziella Grandi 1928 B88
                      |  |    |    |--*D. bicolour B88
                      |  |    |    `--D. falcate B88
                      |  |    `--Robertsia Bouček 1988 B88
                      |  |         |--*R. mandibularis Bouček 1988 B88
                      |  |         `--R. xylosyciae Bouček 1988 B88
                      |  `--Cheiropachys Haliday 1833 MH11, GM79
                      |       `--C. quadrum MH11
                      |--+--Nasonia Ashmead 1904 MH11, B88 [incl. Mormoniella Ashmead 1904 B88]
                      |  |    |--N. vitripennis (Walker 1836) B88 (see below for synonymy)
                      |  |    |--‘*Mormoniella brevicornis’ non Nasonia brevicornis Ashmead 1904 B88
                      |  |    |--N. giraulti GJR97
                      |  |    |--N. longicornis GJR97
                      |  |    `--N. peratripes Girault 1917 G17c
                      |  |--Homoporus Thomson 1878 MH11, B88 (see below for synonymy)
                      |  |    |--*H. fulvicornis [=Pteromalus fulvicornis, Merisus (*Homoporus) fulvicornis] B88
                      |  |    |--H. chalcidiphagus (Walsh & Riley 1869) (see below for synonymy) B88
                      |  |    `--H. hallami (Girault 1924) [=Arthrolytus hallami] B88
                      |  `--Pteromalus MH11
                      |--+--Coelopisthia Forster 1856 MH11, GM79
                      |  |    `--C. extenta MH11
                      |  `--Meraporus graminicola MH11
                      `--Panstenon Walker 1846 MH11, B88 [incl. Caudonia Walker 1850 B88; Panstenoninae]
                           |--*P. oxylus [=Miscogaster oxylus] B88
                           |--‘*Caudonia’ agylla B88
                           |--P. australe Girault 1913 [=P. australis] B88
                           |--P. australiense Girault 1913 [=P. australiensis] B88
                           |--P. bellicosum Girault 1913 [=P. bellicosus] B88
                           |--P. clarum Girault 1915 [=P. clarus] B88
                           `--P. gracile Girault 1913 [=P. gracilis] B88

Pteromalinae incertae sedis:
  Isoplatoides B88
  Callitula B88
  Dinotiscus B88
  Manineura B88
  Mesopolobus B88
  Eurydinota Förster 1878 B88
  Goidanichium Bouček 1970 B88
  Euneura B88
  Platecrizotes soudanensis B88
  Asoka B88
  Merismus Walker 1833 G26, GM79
  Pycnetron Gahan 1925 B88
    `--*P. curculionidis Gahan 1925 B88
  Apycnetron Bouček 1988 B88
    `--*A. brevithorax Bouček 1988 B88
  Frena Bouček 1988 B88
    `--*F. oculata Bouček 1988 B88
  Halticopterella Girault & Dodd in Girault 1915 B88
    `--*H. nigriflagellum Girault 1915 B88
  Mokrzeckia Mokrzecki 1933 [incl. Beierina Deluchhi 1958] B88
    `--*M. pini [=Pteromalus pini] B88
  Canberrana Bouček 1988 B88
    `--*C. hypera Bouček 1988 B88
  Nuchata Bouček 1988 B88
    `--*N. conica Bouček 1988 B88
  Fijita Bouček 1988 B88
    `--*F. leveri Bouček 1988 B88
  Pestra Bouček 1988 B88
    `--*P. splendida Bouček 1988 B88
  Neocylus Bouček 1988 B88
    `--*N. nigraeneus (Girault 1913) (see below for synonymy) B88
  Delislea Girault 1936 B88
    `--*D. pattersoni Girault 1936 B88
  Habritella Girault & Dodd in Girault 1915 B88
    `--*H. graciliventris Girault & Dodd in Girault 1915 B88
  Cairnsia Bouček 1988 B88
    `--*C. stylifera Bouček 1988 B88
  Trinotiscus Bouček 1988 B88
    `--*T. susuanus Bouček 1988 B88
  Briania Bouček 1988 B88
    `--*B. kukensis Bouček 1988 B88
  Buloloa Bouček 1988 B88
    `--*B. bidens Bouček 1988 B88
  Kratinka Bouček 1988 B88
    `--*K. brevis Bouček 1988 B88
  Chlorocytus Graham 1956 B88
    `--*C. pulchripes [=Pteromalus pulchripes] B88
  Mirekia Bouček 1988 B88
    `--*M. planiventris Bouček 1988 B88
  Inkaka Girault 1939 [incl. Leptospermophila Girault 1939] B88
    `--*I. quadridentata Girault 1939 [incl. *Leptospermophila fauna Girault 1939] B88
  Pachycrepoideus Ashmead 1904 [incl. Anisopteromalia Bouček 1954, Toxeumopsis Girault 1915] B88
    `--P. vindemmiae (Rondani 1875) (see below for synonymy) B88
  Laticlypa Bouček 1988 B88
    `--*L. sedlaceki Bouček 1988 B88
  Yanchepia Bouček 1988 B88
    `--*Y. rugosa Bouček 1988 B88
  Genangula Bouček 1988 B88
    `--*G. silvestris Bouček 1988 B88
  Sphegipterosema Girault 1913 B88
    `--*S. ferale Girault 1913 [=S. feralis] B88
  Acroclypa Bouček 1988 B88
    `--*A. pallipes Bouček 1988 B88
  Sphegipterosemella Girault 1913 B88
    `--*S. unicolor Girault 1913 [=Pterosema unicolor Girault 1914] B88
  Megadicylus Girault 1929 B88
    `--M. dubius (Girault 1917) [=Sphegipterosema dubia; incl. *M. montanus Girault 1929] B88
  Longinucha Bouček 1988 B88
    `--*L. petiolaris Bouček 1988 B88
  Paracarotomus Ashmead 1894 [=Paracaratomus (l. c.)] B88
    `--*P. cephalotes Ashmead 1894 B88
  Tsela Bouček 1988 B88
    `--*T. robusta Bouček 1988 B88
  Acanthometopon Ashmead 1904 [=Acanthometapon] B88
    `--*A. clavicorne Ashmead 1904 [=A. clavicornis] B88
  Acroclisella Girault 1915 [incl. Asyntomopus Girault & Dodd in Girault 1915] B88
    `--*A. perplexa Girault 1915 [incl. *Asyntomopus flaviscapus Girault & Dodd in Girault 1915] B88
  Hillerita Bouček 1988 B88
    `--*H. buccata Bouček 1988 B88
  Acroclisissa Girault 1933 B88
    `--*A. clypeata (Girault 1926) [=Acroclisella clypeata] B88
  Lomonosoffiella Girault 1913 B88
    `--*L. albipes Girault 1913 B88
  Maorita Bouček 1988 B88
    `--*M. reticulata Bouček 1988 B88
  Paracroclisis Girault 1913 B88
    `--*P. washingtoni Girault 1913 B88
  Notoglyptus Masi 1917 B88
    `--N. scutellaris (Dodd & Girault in Girault 1915) (see below for synonymy) B88
  Muscidifurax Girault & Sanders 1910 B88
    |--*M. raptor Girault & Sanders 1910 B88
    `--M. uniraptor GJR97
  Oxysychus Delucchi 1956 B88
    |--*O. silvestrii [=Dinarmus silvestrii] B88
    `--O. ferus (Girault 1927) [=Metastenoides ferus] B88
  Uniclypea Bouček 1976 B88
    |--*U. conica B88
    `--U. deporai Bouček 1988 B88
  Propicroscytus Szelényi 1941 [incl. Obtusiclava Subba Rao 1973] B88
    |--P. mirificus (Girault 1915) (see below for synonymy) B88
    `--P. oryzae (Subba Rao 1973) [=*Obtusiclava oryzae] B88
  Anisopteromalus Ruschka 1912 [incl. Aplastomorpha Crawford 1913] B88
    |--*A. mollis B88
    `--A. calandrae (Howard 1881) (see below for synonymy) B88
  Lariophagus Crawford 1909 [incl. Uriellomyia Girault 1915] B88
    |--*L. texanus B88
    `--L. distinguendus (Förster 1841) (see below for synonymy) B88
  Pterosemigastra Girault & Dodd in Girault 1915 B88
    |--*P. oenone Girault & Dodd in Girault 1915 B88
    `--P. nigriflagellum Girault & Dodd in Girault 1915 B88
  Roptrocerus Ratzeburg 1848 [=Pachyceras Ratzeburg 1844 (nom. rej.); incl. Roptroceroidea Ishii 1939] B88
    |--*R. xylophagorum (Ratzeburg 1844) [=*Pachyceras xylophagorum] B88
    `--‘*Roptroceroidea’ karafutoensis B88
  Rhaphitelus Walker 1834 (see below for synonymy) B88
    |--*R. maculatus Walker 1834 [incl. Eucerchysius scolytii] B88
    `--R. ladenbergi B88
  Stinoplus Thomson 1878 B88
    |--*S. militaris [=Etroxys (*Stinoplus) militaris] B88
    `--S. etearchus (Walker 1848) (see below for synonymy) B88
  Lampoterma Graham 1956 B88
    |--*L. viridis [=Metastenus viridis] B88
    `--L. gympicum Bouček 1988 B88
  Cyclogastrella Bukowski 1938 B88
    |--C. deplanata [incl. *C. quercina] B88
    `--C. shorti Bouček 1988 B88
  Toxeumorpha Girault 1915 [incl. Nigricolana Bouček 1976] B88
    |--*T. nigra Girault 1915 B88
    `--T. nigricola [=Trigonogastra nigricola, *Nigricolana nigricola] B88
  Pterosemopsis Girault 1917 B88
    |--*P. javensis Girault 1917 G17a
    `--P. eilogoa Bouček 1988 B88
  Neopolycystus Girault 1915 B88
    |--*N. insectifurax Girault 1915 B88
    `--N. abdominalis (Girault & Dodd in Girault 1915) [=Polycystella abdominalis] B88
  Oxyharma Bouček 1988 B88
    |--*O. subaenea (Dodd 1917) [=Pterosema subaenea] B88
    `--O. pulchra (Girault & Dodd in Girault 1915) [=Eurydinota pulchra, Trichomalopsis pulchra] B88
  Trigonogastrella Girault 1915 [incl. Cryptoprymnoides Girault 1915] B88
    |--*T. parasitica Girault 1915 B88
    `--T. rabiosa (Girault 1915) [=*Cryptoprymnoides rabiosus] B88
  Syntomopus Walker 1833 [incl. Merismorella Girault 1926] B88
    |--*S. thoracicus B88
    `--S. shakespearei (Girault 1926) [=*Merismorella shakespearei] B88
  Neocatolaccus Ashmead 1904 [incl. Paradicylus Dodd in Girault 1915] B88
    |--*N. tylodermae [=Catolaccus tylodermae] B88
    |--N. moneilemae B88
    `--N. varicornis (Dodd in Girault 1915) [=*Paradicylus varicornis] B88
  Eurydinoteloides Girault 1913 [incl. Aeronea Cameron 1913, Protolaccus Burks 1954] B88
    |--*E. americanus B88
    |--E. albiclavus Girault 1917 G17e
    `--E. longiventris B88
  Pterisemoppa Girault 1933 [incl. Lincolnanna Girault 1939] B88
    |--*P. australiensis (Girault & Dodd in Girault 1915) [=Sphegipterosema australiensis] B88
    |--P. malpighii (Girault 1939) [=*Lincolnanna malpighii] B88
    `--P. poeta Girault 1933 B88
  Epanogmus Girault 1920 B88
    |--*E. breviventris Girault 1920 B88
    |--E. bebius (Walker 1839) [=Pteromalus bebius] B88
    `--E. sanguiniventris Girault 1938 B88
  Apsilocera Bouček 1956 B88
    |--*A. verticillata B88
    |--A. australis Bouček 1988 B88
    `--A. bramleyi B88
  Klabonosa Bouček 1976 B88
    |--*K. ferox B88
    |--K. austini Bouček 1988 B88
    `--K. cabra Bouček 1988 B88
  Pseudanogmus Dodd & Girault in Girault 1915 [incl. Pseudomicromelus Gahan & Fagan 1923] B88
    |--*P. fasciipennis B88
    |--P. australia (Girault 1917) [=Dibrachys australia, Pseudomicromelus australia] B88
    `--P. silanus (Walker 1843) (see below for synonymy) B88
  Metastenus Walker 1834 [incl. Scymnophagus Ashmead 1904, Tripolycystus Dodd in Girault 1915] B88
    |--*M. concinnus B88
    |--M. sulcatus (Dodd in Girault 1915) [=*Tripolycystus sulcatus] B88
    `--‘*Scymnophagus’ townsendi B88
  Aiemea Bouček 1988 B88
    |--*A. albicrus Bouček 1988 B88
    |--A. flavipes Bouček 1988 B88
    `--A. monticola Bouček 1988 B88
  Psilocera Walker 1833 (see below for synonymy) B88
    |--*P. obscura B88
    |--P. pulchricornis (Girault 1915) [=*Parapolycystus pulchricornis] B88
    `--P. tennysoni (Girault 1913) (see below for synonymy) B88
  Acroclisoides Girault & Dodd in Girault 1915 B88
    |--*A. megacephalus Girault & Dodd in Girault 1915 B88
    |--A. laticeps Dodd & Girault in Girault 1915 B88
    |--A. major Dodd & Girault in Girault 1915 B88
    `--A. tectacorisi (Girault 1924) [=Pachycrepis tectacorisi] B88
  Agiommatus Crawford 1911 [incl. Polycystella Girault 1913, Pterosemella Girault 1913] B88
    |--*A. sumatraensis B88
    |--A. javensis (Girault 1917) [=Cyrtogaster javensis] B88
    |--A. paria B88
    `--A. viridis (Girault 1913) (see below for synonymy) B88
  Cryptoprymna Förster 1856 (see below for synonymy) B88
    |--*C. atra [=*Prosodes ater] B88
    |--C. africana B88
    |--C. australiensis (Girault 1913) (see below for synonymy) B88
    `--C. crucigera Bouček 1988 B88
  Dinarmus Thomson 1878 (see below for synonymy) B88
    |--*D. acutus [=Dimachus (*Dinarmus) acutus] B88
    |--D. colemani B88
    |--‘*Bruchobius’ laticeps B88
    |--D. simus (Girault 1915) [=*Metastenoides simus; incl. Toxeumella pulchripes Girault & Dodd 1915] B88
    `--‘*Oedaule’ stringifrons B88
  Eurydinotomorpha Girault 1913 B88
    |--*E. pax Girault 1913 B88
    |--E. basalis Girault 1915 B88
    |--E. fusciventris Girault 1913 B88
    |--E. grandis Girault 1915 B88
    |--E. incerta Girault 1915 B88
    `--E. monteithi Bouček 1988 B88
  Trichomalopsis Crawford 1913 (see below for synonymy) B88
    |--*T. shirakii B88
    |--T. australiensis (Girault 1926) [=Metadicylus australiensis] B88
    |--T. braconis (Dodd 1917) [=Eurydinota braconis] B88
    |--T. braconophaga (Cameron 1912) [=Trichoglenes braconophagus] B88
    |--T. iambe (Walker 1839) [=Pteromalus iambe] B88
    |--‘Pteromalus’ nidulans [=*Eupteromalus nidulans] B88
    `--‘Merisus’ subapterus Riley 1885 G17d, GM79 (see below for synonymy)
  Norbanus Walker 1843 (see below for synonymy) B88
    |--*N. dysaulus B88
    |--N. aureicornis (Dodd & Girault in Girault 1915) [=Neocaudonia aureicornis] B88
    |--N. cyaneus (Girault 1913) [=*Amicromelus cyaneus] B88
    |--N. grandaevus (Girault 1925) [=Amicromelus grandaevus] B88
    |--N. longifasciatus (Girault 1914) [=*Apirene longifasciata] B88
    |--N. scelestus (Girault 1914) [=*Epicaudonia scelestus] B88
    |--N. spinosiclava (Girault & Dodd in Girault 1915) [=*Neocaudonia spinosiclava] B88
    `--N. tenuicornis B88
  Merismomorpha Girault 1913 (see below for synonymy) B88
    |--*M. acutiventris Girault 1913 B88
    |--M. asilus (Girault 1915) [=Epipolycystus asilus] B88
    |--M. faunus Girault 1933 B88
    |--M. flavipetiole (Girault 1933) [=*Giorgionia flavipetiole] B88
    |--M. fulvicoxa Girault 1913 B88
    |--M. nigra Girault 1913 B88
    |--M. petiolata (Girault & Dodd in Girault 1915) [=Polycystella petiolata] B88
    `--M. sicarius (Girault 1915) [=Neopolycystella sicarius] B88
  Dibrachys Förster 1856 B88
    |  i. s.: D. apatelae (Ashmead 1893) GM79
    |         D. cavus B88
    |         D. daphne Girault 1917 G17c
    |         ‘Coelopisthia’ diacrisiae Crawford 1910 G17f, GM79
    |         ‘Tritneptis’ hemerocampae Girault 1908 G17f, GM79
    |--D. (Dibrachys) [incl. Coelopisthoidea Gahan 1913] B88
    |    |--*D. (D.) boucheanus [=Pteromalus boucheanus] B88
    |    `--D. (D.) boarmiae (Walker in Newman 1863) [=Pteromalus boarmiae] B88
    `--D. (Allodibrachys Bouček 1965) B88
  Trigonoderus irvingi Girault 1917 G17d
  Trigonoderoides Kamijo 1963 B88
    |--*T. nigricans [=Glyptosticha nigricans] B88
    `--T. niger Bouček 1988 B88
  Miscogasteriella Girault 1915 [incl. Glyptosticha Masi 1926] B88
    |--*M. longiventris Girault 1915 B88
    |--M. flavipes [=*Glyptosticha flavipes] B88
    `--M. sulcata (Kamijo 1963) [=Glyptosticha sulcata] B88
  Apocrypta Coquerel 1855 (see below for synonymy) B88
    |--*A. perplexa B88
    |--A. acaeta Ulenberg in Ulenberg & Pelt 1985 B88
    |--A. caudata (Girault 1915) [=*Goniogastrella caudatus] B88
    |--A. centrocampta Ulenberg in Ulenberg & Pelt 1985 B88
    |--A. channingi (Girault 1913) [=*Idarnoides channingi] B88
    |--A. macroseta Ulenberg in Ulenberg & Pelt 1985 B88
    |--A. mega Ulenberg in Ulenberg & Pelt 1985 B88
    |--A. megischia Ulenberg in Ulenberg & Pelt 1985 B88
    `--A. meromassa Ulenberg in Ulenberg & Pelt 1985 B88
  Critogaster B88
  Dobunabaa Bouček 1988 B88
    `--*D. flava Bouček 1988 B88
  Tenka Bouček 1988 B88
    `--*T. percaudata Bouček 1988 B88
  Watshamiella Wiebes in Bouček, Watsham & Wiebes 1981 [=Metidarnes Riek 1970 (n. n.)] B88
    |--*W. alata B88
    `--W. aurea (Girault 1919) [=Philotrypesis aurea] B88
  Calolelaps Timberlake 1925 D07
    |--*C. basalis Timberlake 1925 D07
    `--C. coeruleus Timberlake 1925 D07
  Mesolelaps Ashmead 1901 D07 [=Mesolaelaps Schulz 1906 B88; Mesolelapini]
    `--*M. cyaneiventris Ashmead 1901 D07
  Neolelaps Ashmead 1901 D07
    |--*N. hawaiiensis Ashmead 1901 D07
    `--N. flavipes Ashmead 1901 D07
  Stictolelaps Timberlake 1925 D07
    |--*S. flaviventris Timberlake 1925 D07
    `--S. stigmatus Timberlake 1925 D07
  Eutelisca D07
  Dinarmolaelaps Masi 1917 D07
    |--*D. protus Masi 1917 D07
    `--D. vatomandryi Risbec 1952 D07

Agiommatus viridis (Girault 1913) [=*Pterosemella viridis; incl. *Polycystella fasciativentris Girault 1913] B88

Anisopteromalus calandrae (Howard 1881) [=Pteromalus calandrae; incl. Neocatolaccus australiensis Girault 1913, Aplastomorpha australiensis, *Ap. pratti, Meraporus vandinei, Ap. vandinei] B88

Apocrypta Coquerel 1855 [incl. Agaonella Baker 1913, Goniogaster Mayr 1885, Goniogastrella Girault & Dodd in Girault 1915, Idarnoides Girault 1913; Apocryptini] B88

Cryptoprymna Förster 1856 [=Prosodes Walker 1833 non Eschscholz 1829; incl. Polycystelomorpha Girault 1915] B88

Cryptoprymna australiensis (Girault 1913) [=Syntomopus australiensis; incl. *Polycystelomorpha flavifemur Girault 1915] B88

Dinarmus Thomson 1878 [incl. Bruchobius Ashmead 1904, Metastenoides Girault 1915, Oedaule Waterston 1922, Sphaerakis Masi 1924] B88

Homoporus Thomson 1878 MH11, B88 [incl. Merisoporus Masi 1924 B88, Parapteromalus Ashmead 1904 B88, Phaenacra Förster 1878 B88, Pseudomerisus Erdös & Novitzky 1953 B88]

Homoporus chalcidiphagus (Walsh & Riley 1869) [=Semiotellus chalcidiphagus, Merisoporus chalcidiphagus] B88

Lariophagus distinguendus (Förster 1841) [=Pteromalus distinguendus; incl. Nasonia miltoni Girault 1929, *Uriellomyia resoluta Girault 1915] B88

Merismomorpha Girault 1913 [incl. Epipolycystus Girault 1915, Giorgionia Girault 1933, Neopolycystella Girault 1915] B88

‘Merisus’ subapterus Riley 1885 G17d, GM79 [=*Nemicromelus subapterus G17d; incl. M. fulvipes (Forbes 1885) G17d, GM79]

Nasonia vitripennis (Walker 1836) B88 [=Pteromalus vitripennis B88, Mormoniella vitripennis A71; incl. *Nasonia brevicornis Ashmead 1904 B88]

*Neocylus nigraeneus (Girault 1913) [=Polycysteloides nigraeneus; incl. Aplastomorpha reticulata Dodd in Girault 1915] B88

Norbanus Walker 1843 [incl. Amicromelus Girault 1913, Apirene Girault 1914, Arthrolysis Förster 1856, Epicaudonia Girault 1914, Neocaudonia Dodd in Girault 1915, Picroscytus Thomson 1878, Stylophorella Ashmead 1904] B88

Notoglyptus scutellaris (Dodd & Girault in Girault 1915) [=Merismus scutellaris; incl. *N. niger Masi 1917, M. squamosus Girault 1915, N. virescens Masi 1917] B88

Pachycrepoideus vindemmiae (Rondani 1875) [=Pteromalus vindemmiae, Pa. vindemiae (l. c.); incl. *Anisopteromalia crassinervis, Toxeumella dissimilis Dodd & Girault in Girault 1915, Pterosemoidea drosophilae Dodd 1917, *Pachycrepoideus dubius Ashmead 1904, *Toxeumopsis nigra Girault 1915] B88

Pachyneuron Walker 1833 MH11, B88 [incl. Atrichoptilus Delucchi 1955 B88, Eupachyneuron Blanchard 1948 B88, Nepachyneuron Girault 1917 B88, Propachyneuronia Girault 1917 B88, Serimus Brèthes 1913 B88]

Propicroscytus mirificus (Girault 1915) [=Arthrolysis mirificus; incl. A. flaviventris Girault & Dodd in Girault 1915, Propicroscytus flaviventris, P. indicus Subba Rao 1981, Arthrolysis trilongifasciatus Girault 1915, *P. trilongifasciatus] B88

Pseudanogmus silanus (Walker 1843) [=Micromelus silanus, M. silvanus (l. c.), *Pseudomicromelus silanus, *Pseudomicromelus silvanus (l. c.); incl. Pseudanogmus fasciipennis Dodd in Girault 1915, Pseudanogmus fuscipes Girault 1915, Pteromalus stironotus Cameron 1912] B88

Psilocera Walker 1833 [incl. Dichalysis Förster 1856, Eupsilocera Westwood 1839, Metopon Walker 1834, Parapolycystus Girault & Dodd in Girault 1915, Polycystoides Girault 1913] B88

Psilocera tennysoni (Girault 1913) [=*Polycystoides tennysoni; incl. Parapolycystus clavicornis Dodd in Girault 1915] B88

Pteromalinae [Bruchobiini, Dinarmini, Merisinae, Mischogastrides, Miscogasteridae, Miscogasterinae, Miscogasteroidae, Pteromalina, Pteromalini, Sphegigasterinae, Sycoryctinae, Sycoryctini, Trigonoderini] B88

Rhaphitelus Walker 1834 [incl. Eucerchysius Brèthes 1913, Storthygocerus Ratzeburg 1848, Styloceras Ratzeburg 1844; Rhaphitelini] B88

Stinoplus etearchus (Walker 1848) [=Pteromalus etearchus; incl. Etroxys (Stinoplus) aureolus (Thomson 1878)] B88

Trichomalopsis Crawford 1913 [incl. Eupteromalus Kurdjumov 1913, Metadicylus Girault 1926, Nemicromelus Girault 1917] B88

*Type species of generic name indicated

REFERENCES

[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[D07] Desjardins, C. A. 2007. Phylogenetics and classification of the world genera of Diparinae (Hymenoptera: Pteromalidae). Zootaxa 1647: 1-88.

[GJR97] Giordano, R., J. J. Jackson & H. M. Robertson. 1997. The role of Wolbachia bacteria in reproductive incompatibilities and hybrid zones of Diabrotica beetles and Gryllus crickets. Proceedings of the National Academy of Sciences of the USA 94: 11439-11444.

[G17a] Girault, A. A. 1917a. New Javanese Hymenoptera. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 59-71).

[G17b] Girault, A. A. 1917b. Descriptiones stellarum novarum. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 80-101).

[G17c] Girault, A. A. 1917c. Descriptiones hymenopterorum chalcidoidicarum variorum cum observationibus. III. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 102-111).

[G17d] Girault, A. A. 1917d. Descriptiones hymenopterorum chalcidoidicarum variorum cum observationibus. V. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 116-131).

[G17e] Girault, A. A. 1917e. Chalcidoidea nova Marilandensis. III. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 132-137).

[G17f] Girault, A. A. 1917f. Descriptiones hymenopterorum chalcidoidicarum variorum cum observationibus. V. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 116-131).

[G26] Girault, A. A. 1926. New pests from Australia. IV. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 205).

[GM79] Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 1-400.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.