Colpochila

Colpochila obesa, from Insects of Tasmania.

Belongs within: Melolonthinae.

Colpochila is a diverse genus of large chafers found in Australia.

See also: Colpochila: the chafing of a mega-genus.

Characters (from Britton 1986): Large (length 11-33 mm), of stout build; labrum not fused to clypeus; antennae 8- or 9-segmented, including a 3- to 7-lamellate club; eyes nearly circular though partly covered by prothorax; elytra not abbreviated (ratio of maximum length:width >2.3:1), covering propygidium and (in part) pygidium; elytral striae grouped in pairs so alternate intervals are obviously different in width; posterior and lateral margins of posterior coxae never thin and membranous; claws without a pulvillus beneath; abdomen with only terminal ventrite movable, separated from fifth ventrite by a pale membrane; propygidium and penultimate ventrite separated by suture on each side.

<==Colpochila Erichson 1843 [incl. Aplonycha Dejean 1833 (n. n.), Haplonycha Blanchard 1851] B86
    |--*C. gouldi (Hope 1842) [=Sericesthis gouldi, Haplonycha gouldi; incl. H. pulchripennis Lea 1924] B86
    |--C. abdita Britton 1986 HW92
    |--C. accepta (Blackburn 1906) [=Haplonycha accepta] HW92
    |--C. aequaliceps (Blackburn 1906) [=Haplonycha aequaliceps] HW92
    |--C. affinis Britton 1986 HW92
    |--C. amabilis (Blackburn 1906) [=Haplonycha amabilis] HW92
    |--C. andersoni Britton 1986 HW92
    |--C. antennalis (Blackburn 1906) [=Haplonycha antennalis] HW92
    |--C. aquila Britton 1986 HW92
    |--C. arida Britton 1986 HW92
    |--C. badia (Burmeister 1855) [=Haplonycha badia] HW92
    |--C. bella Blackburn 1890 HW92 (see below for synonymy)
    |--C. bicolor Blackburn 1890 [=Haplonycha bicolor] B86
    |--C. bidentipes (Lea 1926) [=Haplonycha bidentipes] HW92
    |--C. bimucronata (Lea 1917) [=Haplonycha bimucronata] HW92
    |--C. blackburni Britton 1986 HW92
    |--C. bogania Britton 1986 HW92
    |--C. boreas Britton 1986 HW92
    |--C. brevisetosa (Lea 1930) [=Haplonycha brevisetosa; incl. C. laciniata Britton 1959] HW92
    |--C. calabyi Britton 1986 HW92
    |--C. callida Britton 1986 HW92
    |--C. capta Britton 1986 HW92
    |--C. carinata Blackburn 1890 [=Haplonycha carinata] B86
    |--C. carnabyi Britton 1986 HW92
    |--C. castanea Britton 1986 HW92
    |--C. chinnicki Britton 1986 HW92
    |--C. clara (Blackburn 1906) [=Haplonycha clara] HW92
    |--C. clavipalpis Britton 1986 HW92
    |--C. clypealis Blackburn 1906 [=Haplonycha clypealis; incl. H. marginipennis Lea 1919] HW92
    |--C. comma Britton 1986 HW92
    |--C. crassiventris Blanchard 1850 HW92 (see below for synonymy)
    |--C. crepera Britton 1986 HW92
    |--C. crinita (Burmeister 1855) [=Haplonycha crinita] HW92
    |--C. deceptor Blackburn 1890 [=Haplonycha deceptor; incl. H. punctatissima Blackburn 1906] B86
    |--C. decolor Britton 1986 HW92
    |--C. dilatata (Lea 1917) [=Haplonycha dilatata] HW92
    |--C. efficax Britton 1986 HW92
    |--C. erythrocephala (Lea 1917) [=Haplonycha erythrocephala] HW92
    |--C. faceta (Blackburn 1906) [=Haplonycha faceta] HW92
    |--C. firma (Blackburn 1906) [=Haplonycha firma] HW92
    |--C. fortis Blackburn 1890 [=Haplonycha fortis; incl. C. campestris Blackburn 1892, H. campestris] B86
    |--C. freyi Britton 1959 HW92
    |--C. fulva Britton 1986 HW92
    |--C. funerea Blackburn 1890 HW92 [=Haplonycha funerea L19]
    |--C. gagatina (Burmeister 1855) [=Haplonycha gagatina; incl. H. pygidialis Lea 1930] HW92
    |--C. gibbosicollis Blackburn 1890 [=Haplonycha gibbosicollis, H. gibbicollis (l. c.)] B86
    |--C. gigantea (Burmeister 1855) [=Haplonycha gigantea; incl. H. fimbripennis Lea 1924] HW92
    |--C. goerlingi Britton 1986 HW92
    |--C. griffithi (Blackburn 1907) [=Haplonycha griffithi] HW92
    |--C. ignota Britton 1986 HW92
    |--C. immatura (Lea 1930) [=Haplonycha immatura] HW92
    |--C. infernalis Britton 1986 HW92
    |--C. ingens Britton 1986 HW92
    |--C. interocularis (Lea 1917) [=Haplonycha interocularis] HW92
    |--C. iota Britton 1986 HW92
    |--C. iricolor Britton 1986 HW92
    |--C. iridea (Lea 1930) [=Haplonycha iridea] HW92
    |--C. iris Britton 1986 HW92
    |--C. jungi (Blackburn 1906) [=Haplonycha jungi] HW92
    |--C. kalambi Britton 1986 HW92
    |--C. laminata Blackburn 1890 [incl. C. dubia Blackburn 1890] HW92
    |--C. latebricola (Blackburn 1906) [=Haplonycha latebricola] HW92
    |--C. leo Britton 1986 HW92
    |--C. limbata Britton 1986 HW92
    |--C. longiclava Britton 1986 HW92
    |--C. longior (Blackburn 1906) [=Haplonycha longior; incl. H. pallida Lea 1924] HW92
    |--C. longipalpis (Lea 1926) [=Haplonycha longipalpis] HW92
    |--C. lutea Britton 1986 HW92
    |--C. major Britton 1986 HW92
    |--C. marginata (Blackburn 1906) [=Haplonycha marginata] HW92
    |--C. maura Britton 1986 HW92
    |--C. melina Britton 1986 HW92
    |--C. minor Britton 1986 HW92
    |--C. mixta (Lea 1917) [=Haplonycha mixta] HW92
    |--C. nana Britton 1986 HW92
    |--C. nasuta Britton 1986 HW92
    |--C. nigra (Lea 1919) [=Haplonycha nigra] HW92
    |--C. nigromarginata Britton 1986 HW92
    |--C. nitens Britton 1986 HW92
    |--C. nitida Britton 1986 HW92
    |--‘Haplonycha’ nitidicollis Nonfried 1892 (n. d.) HW92
    |--C. nobilis (Blackburn 1906) [=Haplonycha nobilis; incl. H. colossa Lea 1919, H. lucifera Blackburn 1906] HW92
    |--C. obesa (Boisduval 1835) HW92 (see below for synonymy)
    |--C. opaca (Lea 1917) [=Haplonycha opaca] HW92
    |--C. pagana Britton 1986 HW92
    |--C. pallidula Britton 1986 HW92
    |--C. palpalis Blackburn 1895 HW92
    |--C. parva Britton 1986 HW92
    |--C. paula Britton 1986 HW92
    |--C. piceofulva Britton 1986 HW92
    |--C. picta Britton 1986 HW92
    |--C. pilosa (Blackburn 1906) [=Haplonycha pilosa] HW92
    |--C. pinguis (Macleay 1871) [=Haplonycha pinguis] HW92
    |--C. polita Britton 1986 HW92
    |--C. potens Britton 1986 HW92
    |--C. pruinosa (Lea 1930) [=Haplonycha pruinosa] HW92
    |--C. pubescens Britton 1986 HW92
    |--C. pulchella Blackburn 1890 [incl. Haplonycha novemarticulata Lea 1917, H. octoarticulata Lea 1917] HW92
    |--C. punctiventris Blackburn 1890 [incl. Haplonycha gymnopyga Lea 1917] HW92
    |--C. punctulata Blanchard 1851 B86
    |--C. pygmaea Blackburn 1890 HW92
    |--C. regia Britton 1986 HW92
    |--C. rubida Britton 1986 HW92
    |--C. rubiginosa Britton 1986 HW92
    |--C. ruficeps (Burmeister 1855) [=Haplonycha ruficeps; incl. H. fimbriata Lea 1924] HW92
    |--C. rufocastanea (Lea 1930) [=Haplonycha rufocastanea] HW92
    |--C. rutila Britton 1986 HW92
    |--C. satelles Britton 1986 HW92
    |--C. scutalis (Blanchard 1851) B86 (see below for synonymy)
    |--C. secreta Britton 1986 HW92
    |--C. setifera Britton 1986 HW92
    |--C. setosa Blackburn 1890 [=Haplonycha setosa] B86
    |--C. sinuaticollis Blackburn 1890 (see below for synonymy) B86
    |--C. sodalis Britton 1986 HW92
    |--C. solida Blackburn 1890 [=Haplonycha solida; incl. H. sloanei Blackburn 1906] B86
    |--C. sorella Britton 1986 HW92
    |--C. soror Britton 1986 HW92
    |--C. spadix (Blackburn 1906) [=Haplonycha spadix] HW92
    |--‘Haplonycha’ striatella Blanchard 1851 (n. d.) B86
    |--C. suavis (Lea 1919) [=Haplonycha suavis] HW92
    |--C. tarsalis (Lea 1924) [=Haplonycha tarsalis] HW92
    |--C. tindalei (Lea 1926) [=Haplonycha tindalei] HW92
    |--C. trichopyga (Blackburn 1906) [=Haplonycha trichopyga] HW92
    |--C. unidens Britton 1986 HW92
    |--C. vanga Britton 1986 HW92
    |--C. velata Britton 1986 HW92
    |--C. vesca Britton 1986 HW92
    |--C. vicina Britton 1986 HW92
    `--C. villosa (Lea 1917) [=Haplonycha villosa] B86

Colpochila bella Blackburn 1890 HW92 [incl. Haplonycha amoena Blackburn 1906 HW92, H. cara Lea 1917 HW92, Colpochila gracilis Blackburn 1890 HW92, H. gracilis L19, H. iridipennis Lea 1917 HW92, H. neglecta Blackburn 1906 HW92, H. paradoxa Blackburn 1906 HW92]

Colpochila crassiventris Blanchard 1850 HW92 [incl. Haplonycha fimbricollis Lea 1924 HW92, H. roei Burmeister 1855 HW92, Colpochila roei M86]

Colpochila obesa (Boisduval 1835) HW92 [=Melolontha obesa HW92, Aplonycha obesa B86, *Haplonycha obesa HW92; incl. Melolontha destructor Tepper 1878 HW92, H. pectoralis Blanchard 1851 B86, Colpochila tasmanica Blanchard 1853 B86, H. tasmanica M86, H. thoracica Blackburn 1906 HW92]

Colpochila scutalis (Blanchard 1851) B86 [=Haplonycha scutalis HW92; incl. C. testaceipennis Macleay 1888 HW92, H. testaceipennis L19]

Colpochila sinuaticollis Blackburn 1890 [=Haplonycha sinuaticollis; incl. H. arvicola Blackburn 1906, H. egregia Blackburn 1906, H. electa Blackburn 1906, Colpochila fraterna Blackburn 1890, H. fraterna, H. rustica Blackburn 1906, H. sabulicola Blackburn 1906, H. semiopaca Lea 1917] B86

*Type species of generic name indicated

REFERENCES

[B86] Britton, E. B. 1986. A revision of the Australian chafers (Coleoptera: Scarabaeidae: Melolonthinae) vol. 4. Tribe Liparetrini: genus Colpochila. Australian Journal of Zoology, Supplementary Series 118: 1–135.

[HW92] Houston, W. W. K., & T. A. Weir. 1992. Melolonthinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 174–358. AGPS Press: Canberra.

[L19] Lea, A. M. 1919. Notes on some miscellaneous Coleoptera with descriptions of new species.—Part V. Transactions of the Royal Society of South Australia 43: 166–261, pls 25–27.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.

Saprosites

Saprosites peregrinus, copyright P. Skelley.

Belongs within: Aphodiinae.

Saprosites is a cosmopolitan genus of aphodiine beetles with a cylindrical body, relatively large head and expanded mid and hind tibiae.

<==Saprosites Redtenbacher 1858 CW92
    |--*S. peregrinus Redtenbacher 1858 CW92
    |--S. australis Petrovitz 1958 CW92
    |--S. castaneus Lea 1923 CW92
    |--S. clypealis Lea 1923 CW92
    |--S. mansuetus Blackburn 1904 CW92
    |--S. mendax (Blackburn 1892) [=Ataenius mendax] CW92
    |--S. mesosternalis Lea 1923 CW92
    |--S. nitidicollis (Macleay 1871) [=Ammoecius nitidicollis] CW92
    |--S. pygidialis Lea 1923 CW92
    `--S. sternalis Blackburn 1904 CW92

*Type species of generic name indicated

REFERENCES

[CW92] Cassis, G., & T. A. Weir. 1992. Aphodiinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 81–105. AGPS Press: Canberra.

Ataenius

Ataenius spretulus, copyright Whitney Cranshaw.

Belongs within: Aphodiinae.

Ataenius is a cosmopolitan genus of fairly generalised aphodiine beetles with a basal bead on the elytra.

<==Ataenius Harold 1867 CW92
    |--*A. scutellaris Harold 1867 CW92
    |--A. angulatus Petrovitz 1964 CW92
    |--A. aphodioides Lea 1923 CW92
    |--A. asper Petrovitz 1975 CW92
    |--A. australis Harold 1875 CW92
    |--A. basiceps Lea 1923 CW92
    |--A. brevitarsis Petrovitz 1964 CW92
    |--A. chapini Hinton 1937 H37
    |--A. circulusbrumalis Petrovitz 1964 CW92
    |--A. coloratus Blackburn 1904 CW92
    |--A. communis H37
    |--A. consobrinus Petrovitz 1961 CW92
    |--A. consors Blackburn 1904 CW92
    |--A. crenatipennis (Macleay 1871) [=Ammoecius crenatipennis] CW92
    |--A. cribrithorax H37
    |--A. demarzi Petrovitz 1961 CW92
    |--A. deserti Blackburn 1904 CW92
    |--A. egulyptus H37
    |--A. elongatulus (Macleay 1888) [=Ammaecius elongatulus] CW92
    |--A. erinaceus Petrovitz 1961 CW92
    |--A. eurynotus Lea 1923 CW92
    |--A. excisicollis Petrovitz 1961 CW92
    |--A. figurator H37
    |--A. freyi Petrovitz 1961 CW92
    |--A. gibbus Blackburn 1904 CW92
    |--A. goyderensis Blackburn 1896 CW92
    |--A. gracilis H37
    |--A. illaetabilis Lea 1923 CW92
    |--A. imparilis Blackburn 1904 CW92
    |--A. inciderus Petrovitz 1964 CW92
    |--A. insularis Lea 1923 CW92
    |--A. integricollis Lea 1923 CW92
    |--A. katherinei Petrovitz 1961 CW92
    |--A. koebelei Blackburn 1904 CW92
    |--A. laevifrons Lea 1923 CW92
    |--A. latericollis Lea 1926 CW92
    |--a. limbatus H37
    |--A. macilentus Blackburn 1904 CW92
    |--A. microtrichopterus Lea 1923 CW92
    |--A. molestus Petrovitz 1964 CW92
    |--A. moniliatus Blackburn 1904 CW92
    |--A. nudus Blackburn 1904 CW92
    |--A. obscurus (Macleay 1871) [=Ammoecius obscurus] CW92
    |--A. occidentalis (Macleay 1888) [=Ammaecius occidentalis] CW92
    |--A. palmerstoni Blackburn 1891 CW92
    |--A. panamensis H37
    |--A. parvus Lea 1923 CW92
    |--A. persimilis Lea 1923 CW92
    |--A. picinus Harold 1867 [incl. A. boucomonti Paulian 1937, A. duplopunctatus Lea 1923] CW92
    |--A. scutellaris H37
    |--A. semicoecus (Macleay 1888) [=Ammaecius semicoecus] CW92
    |--A. semicornutus (Macleay 1871) [=Ammoecius semicornutus] CW92
    |--A. setipennis Petrovitz 1961 CW92
    |--A. similis Petrovitz 1961 CW92
    |--A. simulator Harold 1868 CW92
    |--A. sparsicollis Blackburn 1904 CW92
    |--A. speculator Blackburn 1891 CW92
    |--A. spinipennis Lea 1923 CW92
    |--A. spissus Blackburn 1904 CW92
    |--A. spretulus T90
    |--A. strigifrons Schmidt 1921 CW92
    |--A. taylori Petrovitz 1975 CW92
    |--A. thoracicus Petrovitz 1971 CW92
    |--A. torridus Blackburn 1892 CW92
    |--A. tweedensis Blackburn 1904 CW92
    |--A. vulpes Petrovitz 1961 CW92
    `--A. walkeri Blackburn 1904 CW92

*Type species of generic name indicated

REFERENCES

[CW92] Cassis, G., & T. A. Weir. 1992. Aphodiinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 81–105. AGPS Press: Canberra.

[H37] Hinton, H. E. 1937. Ataenius chapini, sp. n., from Mexico (Coleoptera, Scarabaeidae). Proceedings of the Entomological Society of Washington 39 (1): 3–8.

[T90] Tashiro, H. 1990. Insecta: Coleoptera Scarabaeidae larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1191–1209. John Wiley & Sones: New York.

Aphodius (Podotenus)

Aphodius fulvientris, copyright Paul Skelley.

Belongs within: Aphodiinae.

The subgenus Podotenus of the genus Aphodius is an assemblage of aphodiine beetles found in Australia and South America that often have varicoloured elytra with alternately differentiated intervals.

<==Aphodius (Podotenus Schmidt 1914) [incl. Trogaphodius Balthasar 1965] CW92
    |--A. (P.) baldiensis Blackburn 1904 CW92
    |--A. (P.) callabonnensis Blackburn 1895 CW92
    |--A. (P.) erosus Erichson 1842 CW92
    |--A. (P.) fulvientris Fairmaire 1860 CW92
    |--A. (P.) gracilipes Harold 1867 CW92
    |--A. (P.) insignor Blackburn 1904 CW92
    |--A. (P.) lindensis Blackburn 1892 CW92
    |--A. (P.) suberosus Blackburn 1892 CW92
    |--A. (P.) subfuscus Petrovitz 1975 CW92
    |--A. (P.) tuberculatus (Balthasar 1965) [=*Trogaphodius tuberculatus] CW92
    `--A. (P.) victoriae Blackburn 1897 CW92

*Type species of generic name indicated

REFERENCES

[CW92] Cassis, G., & T. A. Weir. 1992. Aphodiinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 81–105. AGPS Press: Canberra.

Corynophyllus

Corynophyllus fortnumi, copyright Ralph Foster.

Belongs within: Dynastinae.

Corynophyllus is a genus of dynastine beetles found in eastern Australia.

Characters (from Carne 1957): Reddish brown, bicolorous, or black, 12-20 mm in length. Mentum narrow, tapering, not compressed, sometimes slightly reflexed at ligula, the latter rounded. Maxillae with galeae small, triangular, untoothed; palps of moderate size, their terminal segments cylindrical. Mandibles broadly ovate, usually surpassed by clypeus, but visible at sides. Clypeus usually elongate and narrowed anteriorly, subtruncate; sometimes rounded. Labrum concealed. Antennae 10-segmented, both faces of club and at least first 2 segments setose. Clypeofrontal ridge transverse; ocular canthi coarse, often angulate, setose. Elytra with sutural striae linear-punctate, rarely linear, disc usually with conspicuous punctate striae, intervals punctate. Epipleurae well defined, bearing abundant lateral setae, these often continuous to apices. Legs slender: fore tibiae tridentate; tarsi with first and fifth segments subequal in length; claws symmetrical. Hind femora unisulcate, abundantly clothed; tibiae bicarinate or with basal carina subobsolete, distal ciliae sharp, stout, closely set; spurs sharp, curved, and unequal. Tarsi slender, all segments subequal in length. Body with ventral surface clothed in abundant white to yellowish brown setae. Postcoxal prosternal process setose, slender and tapering. Male with antennal club greatly enlarged, of characteristic shape. Clypeofrontal ridge giving rise to simple elevation or horn, sometimes to bisinuate ridge or bifurcate process, rarely unarmed. Pronotum with sharp anteromedian tubercle and subcircular anterior impression, or with tubercle reduced to a small node and disc with a large retuse excavation, rarely evenly convex; basal convexity often interrupted by a shallow depression. Pygidium strongly convex, genitalia with gonostyli bearing distolateral brushes of fine setae. Female with antennal club shorter than shaft; head and pronotum unarmed; pygidium more or less flat.

<==Corynophyllus Hope 1845 CW92
    |--*C. fortnumi Hope 1845 CW92
    |--C. andersoni Blackburn 1888 CW92
    |--C. arrowi Carne 1957 CW92
    |--C. curvicornis Lea 1919 CW92
    |--C. debilis Fairmaire 1877 CW92
    |--C. deserti Carne 1957 CW92
    |--C. dorrigoensis Carne 1957 CW92
    |--C. elricus Carne 1957 CW92
    |--C. haroldi Sharp 1875 [incl. C. melas Fairmaire 1877] CW92
    |--C. interocularis Lea 1919 CW92
    |--C. macdougalli Carne 1957 CW92
    |--C. major Sharp 1873 CW92
    |--C. modestus Blackburn 1888 CW92
    |--C. pygidialis Carne 1957 CW92
    |--C. teinogenoides Carne 1957 CW92
    `--C. tulanus Carne 1957 CW92

*Type species of generic name indicated

REFERENCES

Carne, P. B. 1957. A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Division of Entomology, Commonwealth Scientific and Industrial Research Organization, Australia: Melbourne.

[CW92] Cassis, G., & T. A. Weir. 1992. Dynastinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 383–425. AGPS Press: Canberra.

Cryptodus

Cryptodus sp., copyright Bill and Mark Bell.

Belongs within: Dynastinae.

Cryptodus is a genus of dynastine beetles, usually associated with ant or termite nests, found in Australia and New Caledonia.

Characters (from Carne 1957): Mentum very broad, enclosing all mouthparts; base of mentum produced, rarely with conspicuous vestiture. Maxillary palps with terminal segment tapering, as long as first 3 segments combined; labial palps with terminal segment elongate, truncate. Mandibles flattened, molar regions somewhat membranous, scissorial area bidentate. Antennae 10-, 9-, or, rarely, 8-segmented; club shorter than shaft. Clypeus transverse, usually slightly narrowed, more or less truncate. Pronotum weakly convex, with abundant punctation, especially along median line; anterolateral angles acute; anterior margin corneous. Scutellum punctate. Elytra with or without very narrow linear sutural striae, disc with abundant annulate punctation, the intervals commonly costate; epipleurae glabrous. Legs slender: fore tibiae 3- or 4-dentate; tarsi with fifth segment longer than first; claws often asymmetrical in male. Hind femora usually without a distinct sulcus near inner margin; tibiae with distal margin and carinae digitate, very rarely with distinct ciliae; spurs long, acute, curved; hind tarsi with first and fifth segments subequal in length, the former produced to form a conspicuous spine. Pygidium glabrous or bearing a uniform covering of very small fine setae, apical ridge glabrous. Male with pygidium usually very highly convex, occasionally obliquely flattened and convex just before apical ridge; fore tarsi often thickened and bearing asymmetrical claws. Female with pygidium flat, rarely with a median fovea; claws simple.

<==Cryptodus Macleay 1819 [incl. Cryptodellus Westwood 1856, Cryptodinus Westwood 1856] CW92
    |--*C. paradoxus Macleay 1819 (see below for synonymy) CW92
    |--C. bilobus Lea 1917 CW92
    |--C. caviceps Westwood 1856 [=C. (*Cryptodellus) caviceps] CW92
    |--C. concentricus Lea 1920 [incl. C. reginae Arrow 1937] CW92
    |--C. costulipennis Fairmaire 1877 CW92
    |--C. cycnorum Fairmaire 1877 [incl. C. explanatus Carne 1957] CW92
    |--C. debilis Blackburn 1890 [incl. C. aberrans Lea 1919] CW92
    |--C. foveatus Lea 1919 [incl. C. quadridentatus Lea 1920] CW92
    |--C. fraternus Fairmaire 1877 (see below for synonymy) CW92
    |--C. gigas Lea 1917 CW92
    |--C. goerlingi Carne 1957 CW92
    |--C. grossipes Fairmaire 1877 CW92
    |--C. matthewsi Endrödi 1977 CW92
    |--C. oblongoporus Fairmaire 1877 CW92
    |--C. obscurus Macleay 1871 (see below for synonymy) CW92
    |--C. passaloides Germar 1848 [=C. (Cryptodinus) passaloides; incl. C. decipiens Fairmaire 1877] CW92
    |--C. piceus Germar 1848 [incl. C. convexicollis Westwood 1856] CW92
    |--C. relictus Carne 1957 CW92
    |--C. rotundicollis Fairmaire 1877 [incl. C. platyceroides Fairmaire 1879] CW92
    |--C. tasmannianus Westwood 1841 (see below for synonymy) CW92
    `--C. variolosus White 1841 CW92

Cryptodus fraternus Fairmaire 1877 [incl. C. angustus Lea 1919, C. fairmairei Macleay 1888, C. protensus Fairmaire 1879] CW92

Cryptodus obscurus Macleay 1871 [incl. C. antennalis Lea 1919, C. neuter Fairmaire 1877, C. platessa Fairmaire 1877, C. subcostatus Macleay 1871] CW92

*Cryptodus paradoxus Macleay 1819 [incl. C. creberrimus Fairmaire 1877, C. diffinis Fairmaire 1877, C. incornutus Macleay 1871] CW92

Cryptodus tasmannianus Westwood 1841 [=C. (Cryptodinus) tasmannianus; incl. C. anthracinus Erichson 1842, C. ater Lea 1920, C. elongatus Arrow 1937, C. (Cryptodinus) politus Westwood 1856] CW92

*Type species of generic name indicated

REFERENCES

Carne, P. B. 1957. A Systematic Revision of the Australian Dynastinae (Coleoptera: Scarabaeidae). Division of Entomology, Commonwealth Scientific and Industrial Research Organization, Australia: Melbourne.

[CW92] Cassis, G., & T. A. Weir. 1992. Dynastinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 383–425. AGPS Press: Canberra.

Cheiroplatys

Cheiroplatys latipes, from Insects of Tasmania.

Belongs within: Dynastinae.

Cheiroplatys is a genus of dynastine beetles found in Australia and the Americas (Cassis & Weir 1992).

Characters (from Carne 1957): Deep reddish brown to black, length 15–28 mm. Mentum narrow at base, only slightly narrowed towards bilobed ligula; maxillae with galeae strongly developed, 3- to 5-dentate; palps short, with fourth segment elongate, cylindrical or (occasionally) dilated, and abruptly truncate. Mandibles obtusely rounded, slightly bisinuate, surpassed by clypeus, but visible at sides. Labrum completely concealed. Antennae 10-segmented, club usually shorter than shaft, the first 2 segments setose. Clypeus in form of a truncated triangle, the apex obliquely truncate, occasionally serrate. Clypeofrontal ridge transverse, usually forming a conspicuous ridge. Head rugulose, ocular canthi short, setose, continuing side of clypeus without angularity. Pronotum with anterior margin membranous, anterolateral angles acute. Epipleurae with lateral setae confined to anterior half of elytra, the latter with disc usually bearing punctate striae. Postcoxal prosternal process short, columnar, setose. Fore tibiae bi- or trisinuate, never acutely dentate; tarsi with first segment longer than fifth; claws symmetrical. Hind legs stout: femora with sparse vestiture; tibiae bicarinate, ciliae sharp; spurs unequal, blunt, and curved; tarsi short, with first segment longer than fifth, the former moderately to strongly dilated. Pygidium glabrous or sparsely setose across base; apical ridge glabrous or setose. Abdominal sternites other than last commonly glabrous across middle. Last sternite with transverse sulcus, especially in female. Male with pronotum with anteromedian tubercle preceded by an impression or deep excavation; antennal club as long as shaft in those species with serrate clypeal apices, shorter in others. Female with pronotal tubercle and impression reduced or subobsolete.

<==Cheiroplatys Hope 1837 CW92
    |--*C. latipes (Guérin-Méneville in Duperrey 1831) (see below for synonymy) CW92
    |--C. amphioxus Arrow 1937 CW92
    |--C. bifossus Blackburn 1896 [incl. C. accedens Blackburn 1896] CW92
    |--C. borealis Carne 1957 CW92
    |--C. castaneus Lea 1917 CW92
    |--C. curtus Guér. 1830 [incl. C. ixion] M86
    |--C. excavatus Lea 1917 CW92
    |--C. inconspicuus Macleay 1887 CW92
    |--C. nitidus Carne 1957 CW92
    |--C. solidus Carne 1957 CW92
    |--C. subcostatus M86
    |--C. truncatus (Fabricius 1775) [=Geotrupes truncatus] CW92
    `--C. volsellus Carne 1976 CW92

*Cheiroplatys latipes (Guérin-Méneville in Duperrey 1831) [=Scarabaeus latipes; incl. C. compactus Blackburn 1896, C. inermis Blackburn 1900, C. juvencus Burmeister 1847, C. laevipes Burmeister 1847, C. maelius Erichson 1842, Scarabaeus porcellus Boisduval 1835, C. pygmaeus Blackburn 1896] CW92

*Type species of generic name indicated

REFERENCES

[CW92] Cassis, G., & T. A. Weir. 1992. Dynastinae. In: Houston, W. W. K. (ed.) Zoological Catalogue of Australia vol. 9. Coleoptera: Scarabaeoidea pp. 383–425. AGPS Press: Canberra.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.