Hydrocenidae

Georissa wallaceiana, from Stanisic et al. (2010).

Belongs within: Neritoidea.

The Hydrocenidae are a group of small operculate terrestrial snails with a scattered distribution in warmer regions of Eurasia, Africa and Australasia. Members include the genus Georissa, found in northern and eastern Australia and the southwest Pacific, usually on isolated limestone outgroups, with shells ornamented with variously fine to bold spiral cords (Stanisic et al. 2010). The genus Hydrocena is widespread throughout the Old World and may be either smooth or spirally sculpted (Knight et al. 1960).

Characters (from Stanisic et al. 2010): Shell small to minute (less than five millimetres), operculate, globosely turbinate with just a few whorls. Outer lip generally simple, not thickened. Shell whorls usually rounded, less commonly shouldered with strongly impressed sutures. Inner whorls absent, being dissolved or resorbed as shell increases in size. Aperture ovate to semi-circular with prominent columellar callus. Operculum calcareous, D-shaped, bearing a digit-like peg or apophysis on inner surface; nucleus paucispiral. Shell surface either smooth or sculptured to varying degrees with spiral ribs or lines, often bearing calcified accretions. Colour usually translucent yellow, pink or orange. Apical whorls bulb-like, prominent. Animal lacking true tentacles, with broad tentacle-like processes on sides of head; eyes large, mounted on raised papillae; dioecious.

Hydrocenidae [Hydrocaenacea, Hydrocenina, Hydrocenoidea] SS10
    |--Acmella Blanford 1869 S88
    |--Monterissa Iredale 1944 SS10
    |    `--*M. gowerensis Iredale 1944 SS10
    |--Hydrocena Küster 1844 [=Hydrocaena (l. c.); Hydroceninae] BR05
    |    |  i. s.: H. nilotica B01
    |    |         H. noticola [incl. Assiminea tyttha Melvill & Ponsonby 1897] PB27
    |    |         H. saulae van Bentham Jutting 1966 BC01
    |    |         H. scalinella van Bentham Jutting 1966 BC01
    |    |         H. toili Schlickum 1978 TTE93
    |    |--H. (Hydrocena) BR17
    |    |    |--*H. (H.) sirkii (Küster 1844) [=Paludina sirkii] BR17
    |    |    |--H. (H.) cattaroensis (Pfeiffer 1841) [=Cyclostoma cattaroense] KC60
    |    |    `--H. (H.) dubrueilliana E99
    |    |--H. (Chondrella Pease 1871) KC60
    |    |    `--H. (*C.) parva (Pease 1864) [=Cyclostoma parvum] KC60
    |    `--H. (Georissopsis Pilsbry & Hirase 1908) KC60
    |         `--H. (*G.) heudei (Pilsbry & Hirase 1908) [=Georissa (*G.) heudei] KC60
    `--Georissa Blandford 1864 [incl. Omphalorissa Iredale 1937; Georissidae, Georissinae] SS10
         |--*G. pyxis (Benson 1856) [=Hydrocena pyxis] SS10
         |--G. angulata Stanisic in Stanisic, Shea et al. 2010 SS10
         |--G. atramontana Stanisic in Stanisic, Shea et al. 2010 SS10
         |--G. beerwah Stanisic in Stanisic, Shea et al. 2010 SS10
         |--G. chrysacme [=Hydrocena (Georissa) chrysacme] B01
         |--G. enacoucomae Stanisic in Stanisic, Shea et al. 2010 SS10
         |--G. isseliana (Tapparone-Canefri 1883) S88
         |--G. johnnetheryi Stanisic in Stanisic, Shea et al. 2010 SS10
         |--G. laseroni (Iredale 1937) SS10
         |--G. levicordata Stanisic in Stanisic, Shea et al. 2010 SS10
         |--G. minuta (Odhner 1917) SS10
         |--G. monterosatiana (Godwin-Austen & Nevill 1879) AA88
         |--G. multilirata Brazier 1875 SS10 [=*Omphalorissa multilirata P61]
         |--G. nodilirata Stanisic in Stanisic, Shea et al. 2010 SS10
         |--G. obesa Solem 1989 S97
         |--G. palmerensis Stanisic in Stanisic, Shea et al. 2010 SS10
         |--G. postulata (Iredale 1937) SS10
         |--‘Omphalorissa’ purchasi (Pfeiffer 1862) S88
         |--G. quadricarinata (van Benthem Jutting 1963) S88
         |--G. reticulata (van Benthem Jutting 1958) S88
         |--G. rufescens (Moellendorff 1887) AA88
         |--G. saritta [=Hydrocena (Georissa) saritta] B01
         |--G. walkunder Stanisic in Stanisic, Shea et al. 2010 SS10
         |--G. wallaceiana Stanisic in Stanisic, Shea et al. 2010 SS10
         `--G. williamsi (Godwin Austen 1889) [=Hydrocena (Georissa) williamsi] B01

*Type species of generic name indicated

REFERENCES

[AA88] Auffenberg, K., & T. Auffenberg. 1988. Density, spatial distribution, activity patterns, and biomass of the land snail, Geophorus bothropoma Moellendorff (Prosobranchia: Helicinidae). Nautilus 102 (1): 40–45.

[B01] Bandel, K. 2001. The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda). Mitteilungen des Geologisch-Paläontologischen Institutes der Universität Hamburg 85: 65–164.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[E99] Esu, D. 1999. Contribution to the knowledge of Neogene climate changes in western and central Europe by means of non-marine molluscs. In: J. Agustí, L. Rook & P. Andrews (eds) The Evolution of Neogene Terrestrial Ecosystems in Europe pp. 328–354. Cambridge University Press.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[PB27] Pilsbry, H. A., & J. Bequaert. 1927. The aquatic mollusks of the Belgian Congo, with a geographical and ecological account of Congo malacology. Bulletin of the American Museum of Natural History 53 (2): 69–602, pls 10–77.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4^th ed. Whitcombe and Tombs Limited: Christchurch.

[S88] Solem, A. 1988. Non-camaenid land snails of the Kimberley and Northern Territory, Australia. I. Systematics, affinities and ranges. Invertebrate Taxonomy 4: 455–604.

[S97] Solem, A. 1997. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). VII. Taxa from Dampierland through the Nullarbor. Records of the Western Australian Museum Supplement 50: 1461–1906.

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

Last updated: 16 October 2021.

Truncatelloidea

Caecum breve, copyright David Kirsh.

Belongs within: Littorinina.
Contains: Lithoglyphinae, Paleorissoininae, Bythinella, Pomatiopsinae, Anabathron (Scrobs), Potamopyrgus, Vitrinella, Bithynia, Amnicolinae, Baicaliinae, Truncatella, Semisalsinae, Garrettiinae, Assimineinae, Ctiloceratinae, Caecum (Caecum), Caecum (Brochina), Caecum (Fartulum), Iravadia, Macromphalina, Torninae, Teinostoma, Pyrgulinae, Belgrandiella, Horatia, Belgrandiinae, Hydrobiinae.

The Truncatelloidea are a diverse clade of mostly small gastropods united primarily by molecular phylogenetic analysis. Members include the Hydrobiidae which are highly diverse in fresh- and brackish-water environments including many subterranean species. The brackish-water and marine Iravadiidae resemble Hydrobiidae but possess a weak to strong anterior channel on the aperture (vs no such channel in most Hydrobiidae; Ponder 1984). The marine Caecidae have a coiled protoconch leading to an uncoiled teleoconch; in the genus Caecum, the protoconch is shed during growth so the adult shell is a simple tube. The Tornidae possess a low-spired shell, usually with a widely open umbilicus. The Vitrinellidae resemble Tornidae in overall shell form and have been synonymised with tornids by some authors but differ in the presence of a conspicuous, elongate osphradium that is not found Tornidae (Bieler & Mikkelsen 1988). The Assimineidae are terrestrial snails, most diverse in tropical rainforests, with conical to turbinate shells bearing an ovate, thin-lipped aperture (Stanisic et al. 2010).

Truncatelloidea BR17
    |--Calopia Ponder 1999 BR05 [Calopiidae BR17]
    |    `--*C. imitata Ponder 1999 BR17
    |--Clenchiella Abbott 1948 BR05 [Clenchiellidae BR17, Clenchiellinae, Clenchiellini]
    |    `--*C. victoriae Abbott 1948 BR17
    |--Falsicingula Habe 1958 BR05 [Falsicingulidae BR17]
    |    `--*F. kurilensis (Pilsbry 1905) [=Cingula kurilensis] BR17
    |--Helicostoa Lamy 1926 BR05 [Helicostoidae BR17]
    |    `--*H. sinensis Lamy 1926 BR17
    |--Mesocochliopa Yen & Reeside 1946 BR05 [Mesocochliopidae BR17]
    |    `--*M. assiminoides Yen & Reeside 1946 BR17
    |--Hydrococcus Thiele 1928 BR05 [Hydrococcidae BR17, Hydrococcinae]
    |    |--*H. graniformis Thiele 1928 [=Paludina granum Menke 1843 non Say 1822] BR17
    |    `--H. brazieri (Woods 1876) BM88 [=Assiminea brazieri P95]
    |--Lithoglyphidae BR17
    |    |--Lithoglyphinae BR05
    |    `--Benedictiinae [Benedictieae, Benedictiidae, Benedictiini] BR05
    |         |--Benedictia Dybowski 1875 BR05
    |         |    |--*B. (Benedictia) fragilis Dybowski 1875 BR17
    |         |    |--B. andreae [=Paludina andreae] B50
    |         |    |--B. (Baicalocochlea) baicalensis B50
    |         |    `--B. (Dalainoria) limaeoides B50
    |         `--Kobeltocochlea B50
    |              |--K. martensiana B50
    |              `--K. michnoi B50
    |--Palaeorissoinidae BR17
    |    |--Palaeorissoininae GK02
    |    `--Greveniella Harzhauser & Kowalke 2001 [Greveniellinae] GK02
    |         `--*G. mesohellenica Harzhauser & Kowalke 2001 GK02
    |--Elachisinidae BR17
    |    |--Dissochilus lineatus (Briart & Cornet 1887) TTE93
    |    `--Elachisina Dall 1918 BR05
    |         `--*E. grippi Dall 1918 O27
    |--Nematura Benson 1837 [=Stenothyra Benson 1856; Stenothyridae, Stenothyrinae] BR17
    |    |--*N. deltae Benson 1837 [=*Stenothyra deltae] BR17
    |    |--‘Stenothyra’ australis Hedley 1901 H09
    |    `--‘Stenothyra’ pupiformis (Briart & Cornet 1887) TTE93
    |--Bythinellidae BR17
    |    |--Bythinella BR17
    |    `--Terrestribythinella Sitnikova, Starobogatov & Anistratenko 1992 BR17, BR05 [Terrestribythinellidae]
    |         `--*T. baidashnikovi Sitnikova, Starobogatov & Anistratenko 1992 BR17
    |--Pomatiopsidae [Triculidae, Triculinae] BR17
    |    |  i. s.: Prosothenia TTE93
    |    |--Pomatiopsinae BR05
    |    `--Jullienia Crosse & Fischer 1876 BR05 [Jullieniidae, Jullieniinae BR17, Jullieniini]
    |         `--*J. flava (Deshayes 1876) [=Melania flava] BR17
    |--Tomichiidae BR17
    |    |--Tomichia Benson 1851 [Tomichiinae] BR05
    |    |    `--*T. ventricosa (Sowerby 1842) BR17 (see below for synonymy)
    |    `--Coxiella Smith 1894 BR17, BR05 [Coxiellidae]
    |         |--*C. striatula (Menke 1843) [=Truncatella striatula] BR17
    |         `--C. minima Macpherson 1954 M54
    |--Epigrus Hedley 1903 P27 [Epigridae BR17]
    |    |--*E. ischnus (Tate 1899) [=Rissoa ischna] BR17
    |    |--E. dissimilis (Watson 1883) [=Eulima dissimilis] H09
    |    |--E. fossilis P27
    |    |--E. striatus Powell 1927 P27
    |    |--E. verconis (Tate 1899) [=Rissoa verconis] H09
    |    `--E. xanthias (Watson 1886) [=Mucronalia xanthias] H09
    |--Anabathron Frauenfeld 1867 BR05 [Anabathridae BR17, Anabathrinae, Anabathroninae]
    |    |  i. s.: A. ascensum Hedley 1907 H09
    |    |         A. contortum Hedley 1907 H09
    |    |         A. excelsus HS01
    |    |         A. foliatum (Suter 1908) P61
    |    |         A. hedleyi HS01
    |    |         A. rugulosus HS01
    |    |--*A. (Anabathron) contabulata Frauenfeld 1867 BR17
    |    `--A. (Scrobs) TTE93
    |--Moitessieria Bourguignat 1863 BR05 [Moitessieridae, Moitessieriidae BR17, Moitessieriinae, Moitessierioidea]
    |    |--*M. simoniana (Saint-Simon 1848) [=Paludina simoniana] BR17
    |    |    |--M. s. simoniana BV86
    |    |    `--M. s. lescherae Boeters 1981 BV86
    |    |--M. juvenisanguis Boeters & Gittenberger 1980 BV86
    |    |--M. lineolata Coutagne 1881 BV86
    |    |--M. locardi Coutagne 1883 BV86
    |    |--M. olleri BV86
    |    |--M. rhodani Coutagne 1883 BV86
    |    `--M. rollandiana Bourguignat 1863 BV86
    |--Tateidae [Tateinae, Tateoidea] BR17
    |    |--Potamopyrgus BR05
    |    |--Tatea Tenison-Woods 1879 BR05
    |    |    |--*T. huonensis (Tenison-Woods 1876) BR17 [=Bythinia huonensis BR17, Bythinella huonensis L79]
    |    |    `--T. rufilabris H15
    |    |--Hemistomia Crose 1872 [Hemistomiidae, Hemistomiinae] BR05
    |    |    |--*H. caledonica Crosse 1872 BR17
    |    |    `--H. minutissima Ponder 1982 BV86
    |    `--Pyrgophorus Ancey 1888 PB27
    |         |--*P. spinosus [=Pyrgulopsis spinosa] PB27
    |         `--P. reevei (Frauenfeld 1863) BV86
    |--Vitrinellidae BR17
    |    |--Vitrinella BR05
    |    |--Cenomanella Cossmann 1918 TTTE93, KC60
    |    |    `--*C. archiaciana (Orbigny 1847) KC60, TTE93 [=Rotella archiaciana KC60]
    |    |--Circulus Jeffreys 1865 BR17, BR05 [Circulidae, Circulinae]
    |    |    |--C. striatus (Philippi 1836) (see below for synonymy) BM88
    |    |    |--C. cosmius Bartsch 1907 O27
    |    |    |--C. inornatus Marshall 1919 F26
    |    |    |--C. liratus J65
    |    |    |--C. philippii (Cantraine 1842) [=Solarium philippii] KC60
    |    |    |--C. rossellinus Dall 1919 O27
    |    |    `--C. texanus (Moore 1965) [=Vitrinella texana, Vitridomus texana] BM88
    |    `--Solariorbis M65
    |         |--S. blakei M65
    |         `--S. semipunctus Moore 1965 M65
    |--Bithyniidae [Bithiniadae, Bithynioidea, Bulimidae, Buliminae, Bythiniae] BR17
    |    |--Tylopoma G88
    |    |--Bithynia BR05
    |    |--Fossarulus Neumayr 1869 [Fossarulinae] BR05
    |    |    `--*F. stachei Neumayr 1869 BR17
    |    |--Parafossarulus Annandale 1924 [Parafossarulinae] BR05
    |    |    `--*P. striatulus (Benson 1842) [=Paludina striatula] BR17
    |    |--Alocinma Annandale & Prashad 1919 PB27
    |    |    `--*A. sistanica [=Amnicola sistanica] PB27
    |    `--Mysorella Godwin-Austen 1919 BR05 [=Mysoria Godwin-Austen 1919 non Watson 1893 BR17; Mysorellinae]
    |         |--*M. curta (Nevill 1884) BR17 (see below for synonymy)
    |         `--M. multisulcata (Bourguignat 1888) [=Bythinia multisulcata] PB27
    |--Amnicolidae [Amnicolae, Amnicoleae] BR17
    |    |  i. s.: Blanfordia Adams 1863 PB27, BC01
    |    |           |--B. formosana PB27
    |    |           |--B. nosophora (Robson 1915) [=Katayama nosophora] PB27
    |    |           `--B. viridescens (Pease 1861) [=Cyclostoma viridescens] BC01
    |    |         Peringia PB27
    |    |--Amnicolinae BR05
    |    `--Baicaliinae BR05
    |--Emmericiidae BR17
    |    |--Emmericiinae [Emmericieae] BR05
    |    |    |--Pyrgidium Tournouër 1869 non Nylander 1867 (ICBN) [Pyrgidiidae] BR05
    |    |    |    `--*P. nodotianum (Tournouër 1866) [=Pyrgula nodotiana] BR17
    |    |    `--Emmericia Brusina 1870 BR05
    |    |         `--E. ventricosa Brusina 1870 BV86
    |    `--Fontigens Pilsbry 1933 [=Stimpsonia Clessin 1878 non Bate & Westwood 1863; Fontigentinae] BR17
    |         |--*F. nickliniana (Lea 1838) [=Paludina nickliniana, *Stimpsonia nickliniana] BR17
    |         |--F. aldrichi (Call & Beecher 1886) BV86
    |         |--F. cryptica Hubricht 1962 BV86
    |         |--F. holsingeri Hubricht 1976 BV86
    |         |--F. orolibas Hubricht 1957 BV86
    |         |--F. tartarea Hubricht 1962 BV86
    |         `--F. turritella Hubricht 1976 BV86
    |--Truncatellidae BR17
    |    |--Truncatella BR05
    |    |--Blandiella Guppy 1871 BC01
    |    |    `--B. reclusa Guppy 1871 BC01
    |    |--Taheitia Adams & Adams 1863 BC01
    |    |    `--T. whitneyi Clench 1958 BC01
    |    `--Geomelania Pfeiffer 1845 [Geomelaniidae, Geomelaniinae] BR05
    |         |  i. s.: G. affinis Adams 1850 BC01
    |         |         G. minor Adams 1849 BC01
    |         |--G. (Geomelania) BC01
    |         |    |--*G. (G.) jamaicensis Pfeiffer 1845 BR17 (see below for synonymy)
    |         |    `--G. (G.) typica Adams 1850 [incl. G. vicina Adams 1850] BC01
    |         `--G. (Merrilliana Clench & Turner 1948) BC01
    |              |--G. (*M.) elegans Adams 1849 BC01
    |              |--G. (M.) lirata (Poey 1858) [=Truncatella lirata] BC01
    |              `--G. (M.) pygmaea (Adams 1845) [=Cylindrella pygmaea] BC01
    |--Cochliopidae BR17
    |    |--Semisalsinae BR05
    |    |--Littoridininae [Littoridineae, Littoridinidae, Littoridinini, Littoridinoidea] BR05
    |    |    |--Littoridina Souleyet 1852 BR05
    |    |    |    `--*L. gaudichaudii Souleyet 1852 BR17
    |    |    `--Cochliopina Morrison 1946 T84
    |    |         `--C. compacta (Pilsbry 1910) BV86
    |    `--Cochliopinae [Mexithaumatidae, Mexithaumatinae] BR05
    |         |--Potamolithoides GS75
    |         |--Lithococcus GS75
    |         |--Cochliopa Stimpson 1865 [Cochliopini] BR05
    |         |    `--*C. rowellii (Tryon 1863) [=Amnicola rowellii] BR17
    |         |--Mexithauma Taylor 1966 BR05
    |         |    `--*M. quadripaludium Taylor 1966 BR17
    |         `--Paludiscala Taylor 1966 [Paludiscalinae] BR05
    |              `--*P. caramba Taylor 1966 BR17
    |--Assimineidae [Assemaniidae, Assimineidae, Assimineoidea, Assiminiidae, Schistopelmata] BR17
    |    |  i. s.: Nesopoma BC01
    |    |           |--N. eyerdami Clench 1958 BC01
    |    |           `--N. galathea Clench 1958 BC01
    |    |         Setaepoma mayri Clench 1958 BC01
    |    |         Cryptassiminea PC08
    |    |         Ditropisena whitei (Brazier 1874) SS10
    |    |         Limborelia Iredale 1944 SS10
    |    |           |--*L. exquisita (Pfeiffer 1855) (see below for synonymy) SS10
    |    |           `--L. innesi Iredale 1944 [incl. L. innesi adjuncta Iredale 1944] SS10
    |    |         Opinorelia Iredale 1944 SS10
    |    |           `--*O. howeinsulae Iredale 1944 SS10
    |    |         Duritropis Iredale 1944 SS10
    |    |           `--D. albocarinata (Mousson 1873) (see below for synonymy) SS10
    |    |         Telmosena Iredale 1944 SS10
    |    |           `--*T. suteri (Sykes 1900) [=Omphalotropis suteri] SS10
    |    |         ‘Cyclostoma’ brazieri Cox 1870 G78
    |    |         Suterilla Thiele 1927 P61
    |    |           `--*S. neozelanica (Murdoch 1899) [=Cirsonella neozelanica] P61
    |    |         Turbacmella europaea Hrubesch 1965 TTE93
    |    |--Garrettiinae BR17
    |    |--Assimineinae BR05
    |    `--Ekadantinae BR05
    |         |--Ekadanta Rao 1928 BR05
    |         |    `--*E. shanensis Rao 1928 BR17
    |         |--Paludinella Pfeiffer 1841 [Paludinellidae] BR05
    |         |    `--*P. globularis (Hanley in Thorpe 1844) [=Cingula globularis] BR17
    |         `--Cyclotropis Tapparone-Canefri 1883 [Cyclotropidae] BR05
    |              `--*C. papuensis Tapparone Canefri 1883 BR17
    |--Caecidae [Caecoidea] BR17
    |    |--Ctiloceratinae BR05
    |    |--Strebloceras Carpenter 1859 [Strebloceratinae, Streblocerinae] BR05
    |    |    `--*S. cornuoides Carpenter 1859 BR17
    |    `--Caecum Fleming 1813 BR05 [incl. Odontidium Philippi 1836 APS-P86; Caecinae]
    |         |  i. s.: C. angustum de Folin 1886 H09
    |         |         C. californicum Dall 1885 [=C. cooperi Carpenter 1864 non Smith 1862] O27
    |         |         C. clathratum C64
    |         |         C. cooperi Smith 1862 O27
    |         |         C. dalli Bartsch 1920 O27
    |         |         C. diegense Bartsch 1920 O27
    |         |         C. ‘eburneum’ de Folin 1886 non Adams 1852 H09
    |         |         C. grippi Bartsch 1920 O27
    |         |         C. imbricatum HC79
    |         |         C. insularum Moore 1970 BC01
    |         |         C. licalum Bartsch 1920 O27
    |         |         C. lilianum Hedley 1903 H09
    |         |         C. liratocinctum C64
    |         |         C. microcyclus de Folin 1879 H09
    |         |         C. pulchellum WE63
    |         |         C. semilaeve C64
    |         |         C. subflavum de Folin 1879 H09
    |         |         C. subimpressum C64
    |         |         C. succineum de Folin 1879 H09
    |         |         C. undatum C64
    |         |--C. (Caecum) APS-P86
    |         |--C. (Brochina) APS-P86
    |         |--C. (Elephantulum Carpenter 1857) [incl. Elephantanellum Bartsch 1920, Quadrulata Folin 1867] APS-P86
    |         |    |--C. (*E.) hexagonum O27
    |         |    |--C. (E.) abbreviatum Folin in Folin & Périer 1870 APS-P86
    |         |    |--C. (E.) bicinctum Folin in Folin & Périer 1870 APS-P86
    |         |    |--C. (E.) breve Folin 1867 APS-P86
    |         |    |--‘Elephantulum’ carpenteri Bartsch 1920 O27
    |         |    `--C. (E.) clava Folin 1867 APS-P86
    |         |--C. (Fartulum) APS-P86
    |         `--C. (Meioceras Carpenter 1857) APS-P86
    |              |--C. (M.) cubitatum Folin in Folin & Périer 1868 [incl. C. tenerum Folin 1869] APS-P86
    |              |--C. (M.) sandwichensis Folin 1881 APS-P86
    |              `--C. (M.) subinflexum Folin in Folin & Périer 1869 APS-P86
    |--Iravadiidae [Fairbankiidae, Fairbankiinae] BR17
    |    |--Iravadia P84
    |    |--Rehderiella Brandt 1974 [Rehderiellidae, Rehderiellinae, Rehderielloidea] BR05
    |    |    `--*R. parva (Lea 1856) [=Pachychilus parvus] BR17
    |    |--Hyala Adams & Adams 1852 BR05
    |    |    |--*H. vitrea (Montagu 1803) [=Turbo vitreus; incl. H. mediterranea Nordsieck 1972] P84
    |    |    |--H. adamsi Golikov & Kussakin 1971 P84
    |    |    `--H. rotundata C64
    |    |--Liroceratia Ponder 1984 P84
    |    |    |--*L. sulcata (Boettger 1893) [=Cingula sulcata; incl. Pellamora minatura Laseron 1956] P84
    |    |    `--‘Rissoa’ truncata Garrett 1873 P84
    |    |--Acliceratia Ponder 1984 P84
    |    |    |--*A. beddomei (Dautzenberg 1912) [=Aclis beddomei] P84
    |    |    `--A. carinata (Smith 1871) P84 [=Aclis carinata P84, A. walleri var. carinata N79]
    |    |--Rhombostoma Seguenza 1876 [=Eulimopsis Brugnone 1881] P84
    |    |    |--*R. carmelae (Brugnone 1873) [=Eulima carmelae, *Eulimopsis carmelae] P84
    |    |    |--R. imperforatum (Sacco 1892) [=Ondina imperforata; incl. O. pliobliqua Sacco 1892] P84
    |    |    `--R. striatum (Hörnes 1856) [=Chemnitzia striata] P84
    |    |--Ceratia Adams & Adams 1852 [incl. Hebetaclis Pilsbry 1922] P84
    |    |    |--*C. proxima Alder in Forbes & Hanley 1850 [incl. Rissoa striatuls Jeffreys 1847 (preoc.)] P84
    |    |    |--C. alta (Gabb 1873) [=Auriculina alta, *Hebetaclis alta] P84
    |    |    `--C. minutissima Cossmann 1888 P84
    |    |--Rissopsis Garrett 1873 P84
    |    |    |--*R. typica Garrett 1873 P84
    |    |    |--R. prolongata (Turton 1932) [=Rissopsis prolongata; incl. R. ligula Kilburn 1975] P84
    |    |    `--R. tuba Kilburn 1977 P84
    |    |--Chevallieria Cossmann 1888 [incl. Nanadoma Laseron 1956] P84
    |    |    |--*C. labrosa Cossmann 1888 P84
    |    |    |--C. australis Ponder 1984 P84
    |    |    |--C. balcombensis Ponder 1984 P84
    |    |    |--C. columen (Melvill 1904) [=Rissoina columen; incl. R. (Scrobs) elspethae Melvill 1910] P84
    |    |    |--C. cylindroides Cossmann 1907 P84
    |    |    |--C. gippslandica Ponder 1984 P84
    |    |    `--C. imitoris (Laseron 1956) [=*Nanadoma imitoris] P84
    |    `--Nozeba Iredale 1915 (see below for synonymy) P84
    |         |--*N. emarginata (Hutton 1885) P84 (see below for synonymy)
    |         |--‘*Pasithea’ claibornensis Lea 1833 [=*Pasitheola claibornensis] P84
    |         |--N. eulimoides (Cossmann 1888) [=Balanocochlis eulimoides] P84
    |         |--N. gatliffiana (Chapman & Gabriel 1914) [=Rissoa gatliffiana] P84
    |         |--N. globulus (Grateloup 1827) [=Bulimus globulus, *Antinodulus globulus] P84
    |         |--N. guttula (Lea 1833) [=Pasithea guttula] P84
    |         |--N. lucida (Cossmann 1886) [=Amphimelania lucida] P84
    |         |--N. striata Ponder 1984 P84
    |         `--N. topaziaca (Hedley 1908) (see below for synonymy) P84
    |--Tornidae [Tornoidea, Tornoidei] BR17
    |    |  i. s.: ‘Cyclostrema’ bushi Dautzenberg & Fischer 1907 BM88
    |    |         *Pleuromalaxis balesi (Pilsbry & McGinty 1945) BM88
    |    |         Parviturboides interruptus (Adams 1850) BM88
    |    |         Vitridomus Pilsbry & Olsson 1945 BM88
    |    |         Anticlimax decorata Rolán, Fernández-Garcés & Rubio 1997 BC01
    |    |         Vitrinorbis arabscripta Rolán & Rubio 1996 BC01
    |    |         Scissilabra Bartsch 1907 O27
    |    |           `--*S. dalli Bartsch 1907 O27
    |    |         Pseudoliotia Tate 1898 BM88
    |    |           `--P. clathrata M54
    |    |         Tomura BM88
    |    |           |--*T. bicaudata (Pilsbry & McGinty 1946) BM88
    |    |           `--T. abscondita Rolán & Rubio 1999 BC01
    |    |         Cochliolepis BM88
    |    |           |--*C. parasitica Stimpson 1858 BM88
    |    |           |--C. albicerata Ponder 1966 BM88
    |    |           `--C. reductus Rolán & Rubio 1990 BC01
    |    |         Scrupus Finlay 1927 BM88
    |    |           |--*S. hyalinus (Odhner 1924) [=Fossarus hyalinus] P61
    |    |           |--S. marmoratus (Hedley 1907) [=Cithna marmorata] F26
    |    |           |--S. minutus (Michaud 1827) F26, F27 [=Fossarus minutus F27]
    |    |           `--S. uniliratus Powell 1931 P61
    |    |         Naricava Hedley 1913 P61, F26 [incl. Micreschara F26]
    |    |           |  i. s.: N. huttoni (Marwick 1924) (see below for synonymy) F27
    |    |           |--*N. (Naricava) angasi [=Adeorbis angasi] F27
    |    |           `--N. (Tropidorbis Iredale 1936) P61
    |    |                |--N. (*T.) mendica [=*Tropidorbis mendicus] P61
    |    |                `--N. (T.) neozelanica Powell 1940 P61
    |    |         Cyclostremiscus Pilsbry & Olsson 1945 [incl. Ponocyclus Pilsbry 1953] BM88
    |    |           |--*C. panamensis (Adams 1852) [=Vitrinella panamensis] BM88
    |    |           |--‘Cyclostrema’ angulata Adams 1850 BM88
    |    |           |--C. beauii (Fischer 1857) (see below for synonymy) BM88
    |    |           |--C. gordanus (Hertlein & Strong 1951) [=Cyclostrema gordana] BM88
    |    |           |--C. major Olsson & Smith 1951 BM88
    |    |           `--C. pentagonus (Gabb 1837) [incl. Circulus trilix (Bush 1885)] BM88
    |    |         Macromphalina BC01
    |    |         Pygmaeorota P08
    |    |         Sigaretornus BR17
    |    |--Torninae BR17
    |    `--Teinostoma BR17
    `--Hydrobiidae [Hydrobiae] BR17
         |  i. s.: Fluvidona Iredale 1937 PC96
         |           |--*F. petterdi (Smith 1882) [=Hydrobia petterdi] PC96
         |           `--F. centralia Ponder, Colgan et al. 1996 PC96
         |         0--Jardinella PC96
         |         `--+--Fonscochlea Ponder et al. 1989 PC96
         |            `--Dalhousia Ponder, Colgan et al. 1996 PC96
         |                 |--*D. globosa Ponder, Colgan et al. 1996 PC96
         |                 `--D. harrisi Ponder, Colgan et al. 1996 PC96
         |         Islamia Radoman 1973 [Islamiidae, Islamiinae] BR05
         |           |--*I. servaini (Bourguignat 1887) [=Horatia servaini] BR17
         |           `--I. pusilla (Piersanti 1952) [incl. Valvata troglobia Piersanti 1952] BV86
         |         Pseudamnicola Paulucci 1878 [Pseudamnicolinae] BR05
         |           |--*P. macrostoma (Küster 1852) [=Paludina macrostoma] BR17
         |           |--P. acuta Pan 1982 TTE93
         |           |--P. klemmi Boeters 1969 BV86
         |           |--P. likharevi Izzatulaev 1973 BV86
         |           |--P. lindbergi Boettger 1957 BV86
         |           `--P. pallida (Martens 1874) BV86
         |         Pyrgulinae BR05
         |         Mexipyrgus HH88
         |           |--M. carranzae HH88
         |           |--M. churinceanus HH88
         |           |--M. lugoi HH88
         |           `--M. mojarralis HH88
         |         Pyrgulopsis ozarkensis Hinkley 1915 BC01
         |         Soapitia Binder 1961 BC01
         |           `--*S. dageti Binder 1961 BC01
         |         Subcochliopa Morrison 1946 BC01
         |           `--*S. trochus Morrison 1946 BC01
         |         Austropyrgus PC08
         |         Ascorhis PC08
         |         Clessinia variabilis (Eichwald 1841) [=Paludina variabilis] S01
         |         Belgrandiella BR17
         |         Caspia Clessin & Dybowski 1887 [Caspiidae, Caspiinae] BR17
         |           `--*C. baerii Dybowski 1887 BR17
         |         Mercuria Boeters 1971 [Mercuriinae] BR17
         |           `--*M. confusa (Frauenfeld 1863) [=Amnicola confusa] BR17
         |         Horatiinae [Horatiidae] BR17
         |           |--Horatia BR05
         |           |--Tanousia Bourguignat in Servain 1881 (see below for synonymy) BR05
         |           |    |--*T. zrmanjae (Brusina 1866) [=Lymnaea zrmanjae] BR17
         |           |    `--‘*Lithoglyphulus’ tedanicus Schlickum & Schütt 1971 BR17
         |           |--‘Orientalina’ Radoman 1978 non Kolosnitsyna 1973 (see below for synonymy) BR05
         |           |    |--*O. curta (Kürster 1852) [=Paludina curta, *Orientalia curta] BR17
         |           |    `--O. troglobia (Bole 1961) BV86
         |           `--Sadleriana Clessin 1887 BR17, BR05 [Sadlerianidae, Sadlerianinae]
         |                |--*S. fluminensis (Kuster 1853) BR17 (see below for synonymy)
         |                |--S. bavarica Boeters 1989 SW04
         |                |--S. cavernosa Radoman 1978 SW04
         |                `--S. schmidti (Menke 1849) BV86
         |         Trochidrobia BW04
         |         Cylindrica Clessin 1882 BK20
         |         Antroselates Hubricht 1963 H63a
         |           `--*A. spiralis Hubricht 1963 H63a
         |--Belgrandiinae SW04
         `--+--Hydrobiinae SW04
            `--Nymphophilinae BR05
                 |--Notogillia wetherbyi BR05, H63b
                 |--Marstonia decepta (Baker 1928) DPS84, P88
                 |--Nymphophilus Taylor 1966 [Nymphophilini] BR05
                 |    `--*N. minckleyi Taylor 1966 BR17
                 |--Birgella Baker 1926 T84
                 |    `--*B. subglobosa (Say 1825) (see below for synonymy) T84
                 `--Cincinnatia SW04
                      |--C. cincinnatiensis (Anthony 1840) P88
                      |--C. integra B63
                      `--C. winkleyi (Pilsbry 1912) SW04

Nomen nudum: Littoridina jumandi Bole & Velkovrh 1986 BV86

*Birgella subglobosa (Say 1825) [=Paludina subglobosa, Somatogyrus subglobosus; incl. Melania isogona Say 1829, Birgella subglobosa isogona, Somatogyrus isogona, Paludina pallida Lea 1839] T84

Circulus striatus (Philippi 1836) [=Valvata striata; incl. Delphinula duminyi Requien 1848, *Circulus duminyi, Trochus duminyi] BM88

Cyclostremiscus beauii (Fischer 1857) [=Adeorbis beauii, Cyclostrema beani (l. c.), Cyclostremiscus (*Ponocyclus) beauii, Vitrinella (Solariorbis) beaui; incl. Cyclostrema bicarinata Guppy 1866, Adeorbis beaui var. bicarinata, Circulus bicarinatus, Cyclostremiscus (Ponocyclus) beaui bicarinatus, Ci. stirophorus Smith 1937, Skenea sulcata Bush in Simpson 1887 (n. n.)] BM88

Duritropis albocarinata (Mousson 1873) [incl. Omphalotropis brenchleyi Sykes 1900, *D. brenchleyi, D. fortilirata Iredale 1945] SS10

*Geomelania (Geomelania) jamaicensis Pfeiffer 1845 BR17 [incl. G. expansa Adams 1849 BC01, G. fortis Adams 1850 BC01, G. gracilis Adams 1850 BC01, G. magna Adams 1850 BC01, G. media Adams 1850 BC01]

*Limborelia exquisita (Pfeiffer 1855) [=Hydrocena exquisitor; incl. Omphalotropis pfeifferi Crosse 1868] SS10

*Mysorella curta (Nevill 1884) BR17 [=Bythinia curta BR17, Bithynia costigera var. curta PB27, *Mysoria curta BR17]

Naricava huttoni (Marwick 1924) [=Micreschara (Macromphalina) huttoni, Sigaretus carinatus Hutton 1877 non Münster 1841] F27

Nozeba Iredale 1915 [incl. Neozeba (l. c.); incl. Antinodulus Cossmann in Cossmann & Peyrot 1919, Pasithea Lea 1833 (n. d.), Pasitheola Cossmann 1896, Syntharella Laseron 1955] P84

*Nozeba emarginata (Hutton 1885) P84 [=Rissoa emarginata P84; incl. Rissoina coulthardi Webster 1908 P84, Nozeba coulthardi P61, N. mica Finlay 1930 P84]

Nozeba topaziaca (Hedley 1908) [=Eulima topaziaca, *Syntharella topaziaca; incl. Estea amblycorymba Cotton 1944] P84

‘Orientalina’ Radoman 1978 non Kolosnitsyna 1973 [incl. Orientalia Radoman 1972 non Bykova 1947; Orientaliidae, Orientalinidae] BR05

*Sadleriana fluminensis (Kuster 1853) BR17 [=Paludina fluminensis BR17, Lithoglyphus (Sadleriana) fluminensis SW04]

Tanousia Bourguignat in Servain 1881 [=Lithoglyphulus Schlickum & Schütt 1971; Lithoglyphulidae, Tanousiidae] BR05

*Tomichia ventricosa (Sowerby 1842) BR17 [=Truncatella ventricosa BR17; incl. Tr. ventricosa var. brevis Krauss 1848 PB27, Tomichia ventricosa var. brevis PB27, Hydrobia caledonensis Chaper 1885 PB27, Truncatella capensis Krauss in Pfeiffer 1846 PB27]

*Type species of generic name indicated

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[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4^th ed. Whitcombe and Tombs Limited: Christchurch.

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

[S01] Sturany, R. 1901. Csigák és kagylók [Mollusken]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 69–74. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[SW04] Szarowska, M., & T. Wilke. 2004. Sadleriana pannonica (Frauenfeld, 1865): a lithoglyphid, hydrobiid or amnicolid taxon? Journal of Molluscan Studies 70: 49–57.

[T84] Thompson, F. G. 1984. North American freshwater snail genera of the hydrobiid subfamily Lithoglyphinae. Malacologia 25 (1): 109–141.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[WE63] Warmke, G. L., & D. S. Erdman. 1963. Records of marine mollusks eaten by bonefish in Puerto Rican waters. Nautilus 76 (4): 115–120.

Last updated: 24 February 2022.

Helicininae

Pleuropoma extincta, copyright Bronwen Scott.

Belongs within: Helicinoidea.

The Helicininae are a group of operculate terrestrial snails in which the radula possesses five lateral teeth, including a comb-lateral and an accessory plate (Knight et al. 1960).

<==Helicininae BR05
    |--Bourciera Pfeiffer 1852 BR05 [incl. Pseudhelicina Sykes 1907 KC60; Bourcierinae]
    |    `--*B. helicinaeformis (Pfeiffer 1852) [=Cyclostoma helicinaeforme] BR17
    |--Ceratopoma Moellendorff 1893 KC60, S88 (see below for synonymy)
    |    `--*C. caroli (Kobelt 1886) [=Helicina caroli] KC60
    |--Hemipoma Wagner 1905 KC60
    |    `--*H. hakodadiensis (Hartmann 1890) [=Helicina hakodadiensis] KC60
    |--Orobophana Wagner 1905 KC60
    |    `--*O. uberta (Gould 1847) [=Helicina uberta] KC60
    |--Sturanya Wagner 1905 [incl. Sturanyella Pilsbry & Cooke 1934, Sturyanella] KC60
    |    `--*S. plicatilis (Mousson 1865) [=Helicina plicatilis] KC60
    |--Palaeohelicina Wagner 1905 (Palaeohelicina) KC60
    |    |--*P. (P.) fischeriana (Montrouzier 1863) [=Helicina fischeriana] KC60
    |    `--P. (P.) mayri Clench 1958 BC01
    |--Olygyra Say 1818 BR05 (see below for synonymy)
    |    |--*O. orbiculata Say 1818 [=Helicina (*Olygyra) orbiculata] KC60
    |    `--O. sprucei [=Helicina (Oligyra) sprucei] A66
    |--Sulfurina Möllendorff 1893 [incl. Hypostrongyla Tomlin 1930, Pestomena Iredale 1941 (n. n.)] KC60
    |    |--*S. (Sulfurina) citrina (Grateloup 1840) [=Helicina citrina] KC60
    |    `--S. (Kosmetopoma Wagner 1905) KC60
    |         `--S. (*K.) amaliae (Kobelt 1886) [=Helicina amaliae] KC60
    |--Schasicheila Shuttleworth 1852 [=Atoyac Baker 1928, Schasichila Fischer 1885] KC60
    |    |--*S. (Schasicheila) alata (Pfeiffer 1848) [=Helicina alata, *Atoyac alata] KC60
    |    |--S. (Misantla Baker 1928) KC60
    |    |    `--S. (*M.) misantlensis Fischer & crosse 1893 KC60
    |    `--S. (Necaxa Baker 1928) KC60
    |         `--S. (*N.) minuscula (Pfeiffer 1859) [=Helicina minuscula] KC60
    |--Alcadia Gray 1840 (see below for synonymy) KC60
    |    |  i. s.: A. delmonteana Pilsbry 1932 BC01
    |    |         A. pompholyx (Pilsbry 1933) [=Helicina pompholyx] BC01
    |    |--*A. (Alcadia) major (Gray 1824) [=Helicina major] KC60
    |    |--A. (Analcadia Wagner 1907) KC60
    |    |    `--A. (*A.) dysoni (Pfeiffer 1849) [=Helicina dysoni] KC60
    |    |--A. (Emoda Adams & Adams 1856) [incl. Glyptemoda Clench & Aguayo 1950) KC60
    |    |    `--A. (*E.) silacea (Morelet 1849) [=Helicina silacea] KC60
    |    |--A. (Idesa Adams & Adams 1856) (see below for synonymy) KC60
    |    |    `--A. (*I.) rotunda (d’Orbigny in Sagra 1842) [=Helicina rotunda] KC60
    |    |--A. (Penisoltia Baker 1954) KC60
    |    |    `--A. (*P.) hispida (Pfeiffer 1839) [=Helicina hispida] KC60
    |    `--A. (Striatemoda Baker 1940) [incl. Diaphana Guppy 1895 non Brown 1827] KC60
    |         `--A. (S.) subfusca (Menke 1828) (see below for synonymy) KC60
    |--Helicina Lamarck 1799 BR05 (see below for synonymy)
    |    |  i. s.: H. chionea Pilsbry 1949 BC01
    |    |         H. clappi BP90
    |    |         H. hollandi Adams 1849 BC01
    |    |         H. laciniosa Mighels 1845 [=Pleuropoma laciniosa; incl. Orobophana berniceia] J49
    |    |         H. rotelloidea Mighels 1845 [incl. Pleuropoma hawaiiensis] J49
    |    |--*H. (Helicina) neritella Lamarck 1801 (see below for synonymy) KC60
    |    |--H. (Oxyrhombus Fischer & Cross 1893) [incl. Angulata Wagner 1905, Tamsiana Wagner 1905] KC60
    |    |    `--H. (*O.) amoena Pfeiffer 1845 KC60
    |    |--H. (Pseudoligyra Baker 1954) KC60
    |    |    `--H. (*P.) tenuis Pfeiffer 1849 KC60
    |    `--H. (Tristramia Crosse 1863) [incl. Caloplisma Fischer & Crosse 1893, Retorquata Wagner 1905] KC60
    |         `--H. (*T.) salvini Tristram 1861 KC60
    `--Pleuropoma Moellendorff 1893 KC60, KKW12 (see below for synonymy)
         |--*P. dichroa (Möllendorff 1890) [=Helicina dichroa] SS10
         |--P. brecanrio Stanisic in Stanisic, Shea et al. 2010 SS10
         |--'Helicina (Pachystoma)’ concentrica A66
         |--P. draytonensis (Pfeiffer 1857) [incl. Helicina diversicolor Cox 1866] SS10
         |--P. extincta (Odhner 1917) SS10
         |--P. gladstonensis (Cox 1864) SS10
         |--P. gouldiana (Forbes 1851) (see below for synonymy) SS10
         |--P. jana (Cox 1872) SS10
         |--P. louisiadensis (Forbes 1851) S88
         |--P. macleayi (Brazier 1876) SS10
         |--P. queenslandica Stanisic in Stanisic, Shea et al. 2010 SS10
         |--P. reflexilabiata Stanisic in Stanisic, Shea et al. 2010 SS10
         |--P. richardson Stanisic in Stanisic, Shea et al. 2010 SS10
         |--P. solomonensis (Smith 1885) S88
         |--P. spatei Stanisic in Stanisic, Shea et al. 2010 SS10
         |--P. talusata Stanisic in Stanisic, Shea et al. 2010 SS10
         |--P. verecunda (Gould 1859) [=Helicina verecunda, P. (*Aphanoconia) verecunda, *Sphaeroconia verecunda] KC60
         `--P. walkeri (Smith 1894) KKW12 (see below for synonymy)

Alcadia Gray 1840 [incl. Eualcadia Wagner 1907, Incrustata Wagner 1907, Intusplicata Wagner 1907, Isoltia Guppy 1859, Palliata Wagner 1907, Sericea Wagner 1907] KC60

Alcadia (Idesa Adams & Adams 1856) [incl. Hjalmarsona Baker 1940, Leialcadia Wagner 1907, Mammilla Wagner 1907 non Schumacher 1817, Schrammia Guppy 1895, Weinlandella Baker 1954] KC60

Alcadia (Striatemoda) subfusca (Menke 1828) [=Helicina subfusca; incl. H. striata Lamarck 1822 non Defrance 1821, Alcadia (*Striatemoda) striata] KC60

Ceratopoma Moellendorff 1893 KC60, S88 [incl. Diversicolor Wagner 1905 KC60, Negopenia Iredale 1941 (n. n.) KC60]

Helicina Lamarck 1799 BR05 [=Euneritella Wagner 1905 KC60, Pitonnellus deMontfort 1810 KC60, Pitonnillus deMontfort 1810 KC60; incl. Ampliata Wagner 1907 KC60, Colyma Rafinesque 1815, Festiva Wagner 1910 KC60, Pachytoma Swainson 1840 KC60]

*Helicina (Helicina) neritella Lamarck 1801 [=Helix neritella, *Euneritella neritella, *Pitonnellus neritella] KC60

Olygyra Say 1818 BR05 [=Oligyra BR05; incl. Subglobulosa Wagner 1905 KC60, Succincta Wagner 1905 KC60; Oligyradae, Olygyridae]

Pleuropoma Moellendorff 1893 KC60, KKW12 [incl. Albocincta Wagner 1909 SS10, Aphanoconia Wagner 1905 SS10, Dichroa Wagner 1905 SS10, Pachystoma Wagner 1905 non Guilding 1828 KC60, Reticulata Wagner 1905 SS10, Sculpta Wagner 1905 SS10, Sphaeroconia Wagner 1909 SS10]

Pleuropoma gouldiana (Forbes 1851) [incl. Helicina fumigata Sowerby 1866, H. lizardensis Cox 1868, H. reticulata Pfeiffer 1863, Aphanoconia (Sphaeroconia) subreticulata Wagner 1909] SS10

Pleuropoma walkeri (Smith 1894) KKW12 [=Helicina walkeri KKW12, Orobophana walkeri S88, Aphanoconia (Sphaeroconia) baudinensis Wagner 1909 KKW12]

*Type species of generic name indicated

REFERENCES

[A66] Adams, H. 1866. List of land and freshwater shells collected by Mr. E. Bartlett on the Upper Amazons, and on the River Ucayali, eastern Peru, with descriptions of new species. Proceedings of the Zoological Society of London 1866: 440–445.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[BP90] Burch, J. B., & T. A. Pearce. 1990. Terrestrial Gastropoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 201–309. John Wiley & Sones: New York.

[J49] Johnson, R. I. 1949. Jesse Wedgwood Mighels with a bibliography and a catalogue of his species. Occasional Papers on Mollusks 1 (14): 213–231.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[KKW12] Köhler, F., V. Kessner & C. Whisson. 2012. New records of non-marine, non-camaenid gastropods (Mollusca: Gastropoda) from islands off the Kimberley coast, Western Australia. Records of the Western Australian Museum 27 (1): 21–39.

[S88] Solem, A. 1988. Non-camaenid land snails of the Kimberley and Northern Territory, Australia. I. Systematics, affinities and ranges. Invertebrate Taxonomy 4: 455–604.

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

Signepupina

Signepupina worsfoldi, copyright Femorale.

Belongs within: Cyclophoroidea.

Signepupina is a genus of small operculate snails with a smooth, glossy shell found in northeastern Australia (Stanisic et al. 2010).

Characters (from Stanisic et al. 2010): Shell very small to small in size (up to 12 mm in height); highly polished and glassy, generally pupiform in shape but turbinate as juveniles; whorls rounded to flat in adults, angulate in juveniles, sutures weakly impressed to flat; imperforate as adults but juveniles generally possess a narrow umbilicus; aperture circular in shape, lip thickened and reflected; anterior and posterior breathing canals present, often with accompanying ridges and flanges (auricles) and in some cases considerably elaborated; operculum circular, corneous, thin and multispiral. Colour varies from creamy white to bright yellow, and pale pink to reddish brown.

<==Signepupina Iredale 1937 (see below for synonymy) SS10
    |--S. meridionalis (Pfeiffer 1864) (see below for synonymy) SS10
    |--S. attenuata Stanisic in Stanisic, Shea et al. 2010 SS10
    |--S. babinda Stanisic in Stanisic, Shea et al. 2010 SS10
    |--S. bilinguis (Pfeiffer 1851) SS10
    |--S. coxeni (Brazier 1875) SS10
    |--S. crossei (Brazier 1876) SS10
    |--S. davidsoni Stanisic in Stanisic, Shea et al. 2010 SS10
    |--S. dunkensis Stanisic in Stanisic, Shea et al. 2010 SS10
    |--S. glenugie Stanisic in Stanisic, Shea et al. 2010 SS10
    |--S. masoni Stanisic in Stanisic, Shea et al. 2010 SS10
    |--S. pfeifferi (Dohrn 1862) [incl. Pupina carlottae Cox 1864] SS10
    |--S. pineticola (Cox 1866) [incl. Pupina pineticola modesta Fulton 1906] SS10
    |--S. robusta (Cox 1868) SS10
    |--S. rubiflava Stanisic in Stanisic, Shea et al. 2010 SS10
    |--S. strangei (Pfeiffer 1854) SS10
    |--S. tenuis (Hedley 1912) SS10
    |--S. thomsoni (Forbes 1851) [incl. Pupina bidentata Beddome 1897] SS10
    |--S. ventrosa (Dohrn 1862) [incl. Pupina nitida Brazier 1876] SS10
    |--S. wilcoxi (Cox 1864) [incl. Dolopupina wilcoxi edna Iredale 1940] SS10
    |--S. worsfoldi Stanisic in Stanisic, Shea et al. 2010 SS10
    `--S. yaamba Stanisic in Stanisic, Shea et al. 2010 SS10

Signepupina Iredale 1937 [incl. Diplopupina Iredale 1937, Dolopupina Iredale 1937, Lopupina Iredale 1937, Parapupina Iredale 1937] SS10

Signepupina meridionalis (Pfeiffer 1864) [incl. Pupina clara Fulton 1906, Pupinella macgillivrayi Cox 1864, *Signepupina macgillivrayi] SS10

*Type species of generic name indicated

REFERENCES

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.