Agrilinae

Ethon latipennis, from Brisbane Insects.

Belongs within: Buprestidae.
Contains: Agrilus, Aphanisticus, Cisseis.

The Agrilinae are a group of jewel beetles characterised by a sternal cavity (receiving the prosternal process) formed in more or less equal parts by the mesosternum and metasternum. Members of the tribes Aphanisticini and Tracheini have mesocoxae more widely separated than the procoxae and legs with relatively short tarsi whereas Agrilini and Coraebini have procoxae and mesocoxae more or less equidistant and relatively long tarsi.

Agrilinae [Agrilites, Trachyinae, Trachysites] B14
    |--Tracheini [Brachyini, Pachyschelini] B14
    |    |--Habroloma Thomson 1864 B85
    |    |    |  i. s.: H. australis (Macleay 1888) G08
    |    |    |--H. (Malobroha Cobos 1979) B85
    |    |    `--H. (Parahabroloma Kurosawa 1959) B85
    |    |--Brachys Solier 1833 B14, B85
    |    |--Pachyschelus Solier 1833 B14, B85 [incl. Mentonius Say 1836 B85]
    |    |    |--P. laevigatus B14
    |    |    `--P. terminans (Fabricius 1801) B14
    |    |--Trachys Fabricius 1801 B85
    |    |    |--T. anthaxoides B88
    |    |    |--T. minutus B14
    |    |    |--T. phlyctaenoides Kolenati 1846 B14
    |    |    |--T. picta F37
    |    |    |--T. (Trachys) pipturi Fisher 1937 F37
    |    |    `--T. troglodytiformis B14
    |    |--Hylaeogena Obenberger 1923 B85
    |    |    `--H. (Hedwigiella Obenberger 1941) B85
    |    |--Lius Deyrolle 1864 B85
    |    |--Taphrocerus Solier 1833 B85
    |    |    `--T. (Parabrachys Cobos 1979) B85
    |    |--Leiopleura Deyrolle 1864 [incl. Enbrachys Fisher 1935, Leiopleurella Fisher 1922] B85
    |    |--Callimicra Deyrolle 1864 [incl. Bakerietta Obenberger 1932] B85
    |    `--Neotrachys Obenberger 1923 B85
    |--Agrilini B85
    |    |--Agrilus B14
    |    |--Agrilomorpha Thery 1909 B85
    |    |--Trypantius Waterhouse 1887 [incl. Trypanidius Waterhouse 1889] B85
    |    |--Dismorpha Gistel 1848 [incl. Stenogaster Solier 1833 (preoc.)] B85
    |    |--Wendleria Obenberger 1924 [incl. Wendleriana Thery 1925] B85
    |    |--Mychommatus Murray 1868 B85
    |    |--Maublancia Thery 1946 B85
    |    |--Cyphothorax Waterhouse 1887 B85
    |    |--Geralius Harold 1869 [incl. Acanthopygus Deyrolle 1864 (preoc.)] B85
    |    |--Omochyseus Waterhouse 1887 [=Omochryseus (l. c.)] B85
    |    |--Agrilodia Obenberger 1924 B85
    |    |--Sarawakita Obenberger 1924 B85
    |    |--Parakamosia Obenberger 1924 B85
    |    |--Sjoestedtius Thery 1931 [=Sjoestedtia Thery 1931; incl. Eukamosia Obenberger ms] B85
    |    |--Agriloides Kerremans 1903 [incl. Xenagrilus Obenberger 1924] B85
    |    |--Agrilacanthus Thery 1940 B85
    |    |--Agrilochyseus Thery 1935 B85
    |    |--Callchitones Obenberger 1931 B85
    |    |--Amyia Thomson 1878 B85
    |    |--Callipyndax Waterhouse 1887 B85
    |    |--Euamyia Kerremans 1903 [incl. Eumerus Gory & Laporte 1839 non Geoffroy Saint-Hailaire 1829] B85
    |    |--Pareumerus Deyrolle 1864 B85
    |    |--Rhaeboscelis Chevrolat 1837 B85
    |    |--Velutia Kerremans 1900 B85
    |    `--Paragrilus Saunders 1871 [incl. Clinocera Deryolle 1864 (preoc.)] B85
    |--Aphanisticini [Aphanisticites] B85
    |    |--Aphanisticus B85
    |    |--Anthaxomorphus Deyrolle 1864 [incl. Bolivarina Obenberger 1921] B85
    |    `--Endelus Deyrolle 1864 B85
    |--Germarica Blackburn 1887 [Germaricini] B85
    |--Cylindromorphoidini B85
    |    |--Cylindromorphoides Kerremans 1903 B85
    |    `--Eurynodes Thery 1934 B85
    `--Coraebini [Amorphosternae] B14
         |--Sibuyanella Obenberger 1942 B85
         |    |--S. bakeri (Fisher 1924) B14
         |    |--S. boudanti B14
         |    `--S. mimica B14
         |--Coraebus Laporte & Gory 1839 B85
         |    |  i. s.: C. chrysopygius M86
         |    |         C. elatus [incl. C. elatus var. repletus] C01
         |    |         C. fulgidicollis Lucas 1846 E12
         |    |         C. kamikochianus Kano 1929 [=Coraebuus (l. c.) kamikochianus] TYM08
         |    |         C. loochooensis Kano 1929 [=Coraebuus (l. c.) loochooensis] TYM08
         |    |         C. marmoratus Macleay 1872 M86
         |    |         C. undatus (Fabricius 1787) B14
         |    |         C. westwoodi M86
         |    |--C. (Katonia Thery 1941) B85
         |    `--C. (Negreia Cobos 1962) B85
         |--Synechocera Deyrolle 1864 B85
         |--Pseudosynechocera Obenberger 1943 B85
         |--Paracephala Thomson 1878 B85
         |    |--P. murina Thoms. 1878 M86
         |    `--P. pistacina (Hope 1846) [=Agrilus pistacinus; incl. Aphanisticus canaliculatus] M86
         |--Meliboeithon Obenberger 1920 B85
         |--Dinocephalia Obenberger 1923 B85
         |    `--D. browni (Carter 1933) G08
         |--Alcinous Deyrolle 1864 B85
         |--Clema Semenov-Tian-Shanskij 1900 B85
         |--Pseudoclema Thery 1938 B85
         |--Lepidoclema Bellamy & Holm 1985 B85
         |--Promeliboeus Obenberger 1924 B85
         |--Anodontodora Obenberger 1931 [incl. Discoderes Obenberger 1922 nec Chevrolat 1837 nec Thery 1905] B85
         |--Eudiadora Obenberger 1924 B85
         |--Diadora Kerremans 1900 B85
         |--Diadorina Cobos 1974 B85
         |--Bergidora Kerremans 1903 B85
         |--Helferina Cobos 1956 B85
         |--Amorphosternoides Cobos 1974 B85
         |--Amorphosternus Deyrolle 1864 B85
         |--Amorphosoma Laporte 1835 B85
         |--Anadora Kerremans 1898 B85
         |--Shimogia Obenberger 1942 B85
         |--Kerremansella Obenberger 1923 [incl. Kerremansia Obenberger 1920 non Peringuey 1908] B85
         |--Brachydora Obenberger 1923 B85
         |--Compsoglypta Fairmaire 1904 [=Compsoglyhpa (l. c.)] B85
         |--Discoderopsis Thery 1929 B85
         |--Chalcophlocteis Obenberger 1924 B85
         |--Suarezina Thery 1936 [incl. Suarezia Thery 1912 (preoc.)] B85
         |--Phlocteis Kerremans 1893 [incl. Discoderes Thery 1905 non Chevrolat 1837, Paradora Kerremans 1900] B85
         |--Speophora Thery 1936 B85
         |--Xenita Thery 1941 B85
         |--Seranambia Descarpentries 1974 B85
         |--Angatra Descarpentries 1969 B85
         |--Discoderes Chevrolat 1837 [incl. Paradorella Obenberger 1923] B85
         |    `--D. tasmanicus M86
         |--Discoderoides Thery 1936 B85
         |--Planidia Kerremans 1899 B85
         |--Parademostis Obenberger 1931 B85
         |--Holubia Obenberger 1924 B85
         |--Demostis Kerremans 1900 B85
         |--Helferia Obenberger 1932 B85
         |--Epimacha Kerremans 1900 B85
         |--Polyonychus Chevrolat 1837 B85
         |--Toxoscelus Deyrolle 1864 B85
         |--Dessumia Descarpentries & Villiers 1966 B85
         |--Melanodactylus Baudon 1963 B85
         |--Cryptodactylus Deyrolle 1864 [incl. Strandietta Obenberger 1931] B85
         |--Acanthocephalon Thery 1955 B85
         |--Neotoxoscelus Fisher 1921 B85
         |--Kerremansia Peringuey 1908 B85
         |--Pseudokerremansia Bellamy & Holm 1985 B85
         |--Pseudokerremansia Bellamy 1985 B85
         |--Cupriscobina Bellamy & Holm 1985 B85
         |--Peyrierasina Descarpentries 1975 B85
         |--Strigulia Kerremans 1893 B85
         |--Nastella Kerremans 1903 B85
         |--Paranastelle Obenberger 1931 B85
         |--Jaroslavia Obenberger 1942 B85
         |--Entomogaster Fairmaire 1899 B85
         |--Eumorphocerus Thery 1929 B85
         |--Coraebastus Fairmaire 1896 [incl. Annietta Obenberger 1931] B85
         |--Bourgoinia Obenberger 1926 B85
         |--Evimantius Kerremans 1891 B85
         |--Alissoderus Deyrolle 1864 [=Alyssoderus (l. c.)] B85
         |    `--A. (Angolia Obenberger 1920) B85
         |--Eulasiodora Obenberger 1924 B85
         |--Mundaria Kerremans 1894 B85
         |--Asymades Kerremans 1893 B85
         |--Belgaumia Kerremans 1903 B85
         |--Pseudocoraebus Thery 1905 B85
         |--Chloricala Kerremans 1893 B85
         |--Seyrigia Thery 1936 B85
         |--Neocoraebus Kerremans 1903 B85
         |--Euchroaria Obenberger 1924 B85
         |--Vanroonia Obenberger 1923 B85
         |--Dinocoraebus Obenberger 1924 B85
         |--Cisseicoraebus Kerremans 1903 B85
         |--Coraebosoma Obenberger 1923 B85
         |--Obenbergerula Strand 1932 [incl. Bottcheria Hoscheck 1931 (preoc.)] B85
         |--Melanocoraebus Baudon 1968 B85
         |--Autarcontes Waterhouse 1887 [=Autarchontes (l. c.)] B85
         |--Eupristocerus Deyrolle 1864 B85
         |--Zavattaria Obenberger 1940 B85
         |--Gigantocoraebus Obenberger 1942 B85
         |--Midongya Obenberger 1942 B85
         |--Antanambia Descarpentries 1975 B85
         |--Maroantsetra Thery 1936 B85
         |--Cisseis B85
         |--Cisseioides Kerremans 1893 [incl. Brachycisseis Thery 1931] B85
         |--Hypocisseis Thomson 1879 [incl. Maschalix Waterhouse 1887] B85
         |    `--H. ornata Carter 1923 G08
         |--Ethon Laporte & Gory 1839 B85
         |    |--E. affine G84 [incl. E. aurifluus M86, E. proximus Bohem. 1858 M86, E. purpurascens M86]
         |    |--E. corpulentus Bohem. 1858 M86
         |    |--E. fissiceps [incl. E. viridis] M86
         |    |--E. latipennis Macleay 1872 [incl. Cisseis cornuta] M86
         |    |--E. reichei M86
         |    |--E. roei Saund. 1868 M86
         |    `--E. subfasciatus Saund. 1868 M86
         |--Weyersiella Thery 1929 B85
         |--Brachycoraebus Kerremans 1903 B85
         |--Samboides Kerremans 1900 B85
         |--Metasambus Kerremans 1903 [incl. Pusilloderes Obenberger 1916] B85
         |--Neosambus Fisher 1921 B85
         |--Therysambus Descarpentries & Villiers 1968 B85
         |--Sambus Deyrolle 1864 [incl. Arrowia Obenberger 1923] B85
         |--Parasambus Descarpentries & Villiers 1966 B85
         |--Sambirania Obenberger 1942 B85
         |--Mandritsaria Obenberger 1942 B85
         |--Nickerleola Obenberger 1923 [incl. Achilleus Thery 1929] B85
         |--Eumerophilus Deyrolle 1864 [incl. Aubertia Thery 1922] B85
         |--Pseudagriloides Obenberger 1921 B85
         |--Pseudagrilus Laporte 1835 B85
         |--Tonkinula Obenberger 1923 [incl. Chakriia Tongyai 1935] B85
         |--Kamosia Kerremans 1898 [incl. Gassneria Obenberger 1919] B85
         |--Pseudokamosia Thery 1931 B85
         |--Therybuprestis Strand 1932 [incl. Maindronia Thery 1930 nec Bouvier 1897 nec Roewer 1910] B85
         |--Coraegrilus Fairmaire 1889 [=Coragrilus (l. c.)] B85
         |--Coraebina Obenberger 1923 B85
         |--Melicoraebus Thery 1932 B85
         |--Meliboeus Deyrolle 1864 [=Melybaeus Barthe 1902] B85
         |    |--M. (Melibaeopsis Kerremans 1903) B85
         |    |--M. (Meliboeoides Thery 1942) B85
         |    |--M. (Melixes Schaefer 1949) B85
         |    `--M. (Xenomerius Obenberger 1924) B85
         |--Nalanda Thery 1904 B85
         |--Lakhonia Descarpentries & Villiers 1968 [=Lactonia Thery ms] B85
         |--Sambomorpha Obenberger 1924 B85
         `--Deyrollius Obenberger 1922 [incl. Corydon Deyrolle 1864 (preoc.)] B85
              `--D. (Eucorydon Thery 1934) B85

*Type species of generic name indicated

REFERENCES

[B85] Bellamy, C. L. 1985. A catalogue of the higher taxa of the family Buprestidae (Coleoptera). Navorsinge van die Nasionale Museum Bloemfontein 4 (15): 405–472.

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[F37] Fisher, W. S. 1937. New species of Buprestidae from Java. II (Coleoptera). Proceedings of the Entomological Society of Washington 39 (8): 231–236.

[G84] Gauld, I. D. 1984. An Introduction to the Ichneumonidae of Australia. British Museum (Natural History).

[G08] Golding, M. R. 2008. A Pictorial Field Guide to the Beetles of Australia. Part 3: Chalcophorinae. Ocean Publishing.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.

[TYM08] Tennent, W. J., M. Yasuda & K. Morimoto. 2008. Lansania Journal of arachnology and zoology—a rare and obscure Japanese natural history journal. Archives of Natural History 35 (2): 252–280.

Last updated: 10 December 2020.

Buprestinae

Western cedar borer Trachykele blondeli, copyright Kildale.

Belongs within: Buprestidae.
Contains: Chrysobothris, Buprestis, Bubastini, Dicercini, Anthaxia, Neocuris, Melanophilini, Temognatha, Calodema, Stigmodera, Curis.

The Buprestinae are a group of jewel beetles characterised by the possession of antennae with arrays of pores concentrated in well defined, usually apical depressions on the antennal segments. In members of the tribes Anthaxiini, Melanophilini and Maoraxia, the metepimeron is at least partially covered by a broad anterolateral projection of the abdomen. The metepimeron is uncovered in Buprestini, Bubastini and Dicercini (Bellamy & Williams 1985).

Buprestinae [Chrysobothrinae, Chrysobothrites] B14
    |  i. s.: Agaeocera Waterhouse 1882 [incl. Anataxis Casey 1909; Agaeocerini] B85
    |           `--A. gigas G08
    |         Coomaniella Bourgoin 1924 [Coomaniellini] B85
    |           `--C. (Coomaniellina Bily 1974) B85
    |         Kisanthobia Marseul 1865 [=Cisanthobia (l. c.); Kisanthobiini] B85
    |         Trigonogenium Gemminger & Harold 1869 (see below for synonymy) B85
    |         Phrixiini B85
    |           |--Paraphrixia Saunders 1871 B85
    |           `--Phrixia Deyrolle 1864 B85
    |--Chrysobothrini B85
    |    |--Chrysobothris B14
    |    |--Colobogaster Solier 1833 B85
    |    |--Afrobothris Thery 1936 B85
    |    `--Hoscheckia Thery 1925 B85
    |--Actenodini B85
    |    |--Merimna Thomson 1878 B85
    |    |    `--M. atrata LB91
    |    |--Chalcangium Waterhouse 1887 B85
    |    |--Actenodes Dejean 1833 B85
    |    |--Pseudactenodes Kerremans 1890 [incl. Embrikiola Obenberger 1928] B85
    |    |--Paractenodes Thery 1933 B85
    |    |--Megactenodes Kerremans 1893 B85
    |    `--Belionota Eschscholtz 1829 B85
    |         `--B. (Pseudobelionota Thery 1928) B85
    |--Buprestes BW85
    |    |--Buprestini [Ancylocheirini] B85
    |    |    |--Buprestis B14
    |    |    |--Pterobothris Fairmaire & Germain 1858 B85
    |    |    |--Philandia Kerremans 1907 B85
    |    |    |--Nascio Laporte & Gory 1837 [incl. Geronia Dejean 1838] B85
    |    |    |    |--N. carissima M86
    |    |    |    |--N. chydaea Olliff 1886 O86
    |    |    |    |--N. multesima Olliff 1886 O86
    |    |    |    |--N. munda Olliff 1886 O86
    |    |    |    |--N. parryi Hope 1845 [incl. N. saundersi Hope 1846] M86
    |    |    |    |--N. simillima O86
    |    |    |    |--N. vetustus M86
    |    |    |    |--N. viridis Macleay 1872 M86
    |    |    |    `--N. xanthura M86
    |    |    |--Nascioides Kerremans 1903 B85
    |    |    |    `--N. parryi LB91
    |    |    |--Balthasarella Obenberger 1958 B85
    |    |    |--Trachykele Marseul 1865 B85
    |    |    |    |--T. blondeli Marseul 1865 B14
    |    |    |    |    |--T. b. blondeli B14
    |    |    |    |    |--T. b. cuperomarginata B14
    |    |    |    |    `--T. b. juniperi B14
    |    |    |    |--T. fattigi B14
    |    |    |    |--T. hartmani B14
    |    |    |    |--T. lecontei B14
    |    |    |    |--T. nimbosa B14
    |    |    |    `--T. opulenta B14
    |    |    |--Eurythyrea [=Eurythrea (l. c.)] B85
    |    |    |--Yamina Kerremans 1903 B85
    |    |    |--Stephansortia Thery 1925 B85
    |    |    |--Buprestina Obenberger 1923 B85
    |    |    |--Buprestodes Carter 1924 B85
    |    |    |--Melobasina Kerremans 1900 B85
    |    |    |--Pygicera Kerremans 1903 B85
    |    |    |--Mixochlorus Waterhouse 1887 B85
    |    |    |    `--M. suturalis G08
    |    |    |--Peronaemis Waterhouse 1887 B85
    |    |    |--Philanthaxia Deyrolle 1864 [incl. Engycera Saunders 1867] B85
    |    |    |--Pagdeniella Thery 1935 [incl. Pagdenia Thery 1934 (preoc.)] B85
    |    |    |--Thomassetia Thery 1928 B85
    |    |    |--Aristosoma Thomson 1879 B85
    |    |    `--Jakovleviola Obenberger 1924 B85
    |    |--Bubastini BW85
    |    `--Dicercini BW85
    |--Anthaxiae BW85
    |    |--Anthaxini [Anthaxites] B85
    |    |    |--Anthaxia BW85
    |    |    |--Brasilaxia Thery 1935 B85
    |    |    |--Brachanthaxia Thery 1930 B85
    |    |    |--Odettea Baudon 1966 B85
    |    |    |--Anthaxioides Cobos 1976 B85
    |    |    |--Pseudanilara Thery 1908 [incl. Neotorresita Obenberger 1923] B85
    |    |    |--Anilara Thomson 1879 B85
    |    |    |    `--A. adelaidae (Hope 1846) [=Anthaxia adelaidae] M86
    |    |    |--Notographus Thomson 1879 [incl. Notograptus MacLeay 1872 (preoc.)] B85
    |    |    |    |--‘Notograptus’ hieroglyphicus Macleay 1872 M86
    |    |    |    `--‘Notograptus’ sulcipennis Macleay 1872 M86
    |    |    |--Anthaxoschema Obenberger 1923 B85
    |    |    |--Tetragonoschema Thomson 1857 [incl. Pachypyga Steinheil 1872] B85
    |    |    |    `--T. (Anilaroides Thery 1934) B85
    |    |    |--Brachelytrium Obenberger 1923 B85
    |    |    |--Hesperorhipis Fall 1930 B85
    |    |    |--Xenorhipis LeConte 1866 [incl. Lamesis] B85
    |    |    |--Trichinorhipis Barr 1948 B85
    |    |    |--Isophaenus Waterhouse 1889 B85
    |    |    `--Neocuris B85
    |    |--Melanophilini B14
    |    `--Maoraxia Obenberger 1937 [=Maoriella Obenberger 1924 (preoc.); Maoraxiini] BW85
    |         |--M. eremita (White 1846) (see below for synonymy) BW85
    |         |--M. littoralis Bellamy & Williams 1985 BW85
    |         |--M. storeyi Bellamy & Williams 1985 BW85
    |         |--M. tongae Bellamy & Williams 1985 BW85
    |         `--M. viridis Bellamy & Williams 1985 BW85
    `--Stigmoderini [Stigmoderites] B14
         |--Temognatha G06
         |--Conognatha Eschscholtz 1829 [incl. Polydora Gistel 1834 non Bosc 1802] B85
         |    |  i. s.: C. navarchis Thoms. 1856 M86
         |    `--C. (Pithiscus Solier 1851) B85
         |--Calodema G06
         |--Stigmodera G06
         |--Julodimorpha Gemminger & Harold 1869 [Julodimorphini, Julodimorphites] G06
         |    |--J. bakewelli (White 1859) [=Stigmodera bakewelli] M86
         |    `--J. saundersii Thomson 1878 B14
         |--Castiarina Laporte & Gory 1838 B85
         |    |--C. carinata ZS10
         |    |--C. richardsi G09
         |    |--C. rufipennis ZS10
         |    `--‘Buprestis’ scalaris Boisduval 1835 P91 (see below for synonymy)
         |--Metaxymorpha Parry 1848 [incl. Semnopharus Van de Poll 1886] B85
         |    |--M. apiceruba Thery 1923 G06
         |    |--M. gloriosa Blackburn 1894 G06
         |    |--M. greyi (Parry 1848) G06
         |    |--M. hauseri Thery 1926 G06
         |    |--M. immitator Sainval 1994 G06
         |    |--M. nigrofasciata Nylander 2001 G06
         |    `--M. nigrosuturalis Nylander 2001 G06
         |--Hypostigmodera Blackburn 1892 B85
         |--Curis B85
         |--Selagis Dejean 1835 B85
         |    `--S. aurifera (Gory & Laporte 1838) G08
         |--Dactylozodes Chevrolat 1837 (see below for synonymy) B85
         |    |--D. (Agrilozodes Thery 1927) B85
         |    |--D. (Neodactylozodes Cobos 1958) B85
         |    |--D. (Parazodes Cobos 1958) B85
         |    `--D. (Stenozodes Thery 1935) B85
         `--Hiperantha Gistel 1834 [=Hyperantha; incl. Poecilonota Solier 1833 non Eschscholtz 1829] B85
              `--H. (Hyperanthella Hoscheck 1928) B85

‘Buprestis’ scalaris Boisduval 1835 P91 [=Polychroma scalaris Dejean 1833 (n. n.) P91, Stigmodera scalaris M86; incl. S. crucigera Hope 1836 (nom. inv.) P91, B. octosignata B35]

Dactylozodes Chevrolat 1837 [incl. Lasionota Dejean 1836 (n. n.), Zemina Laporte & Gory 1839, Zemima (l. c.)] B85

Maoraxia eremita (White 1846) [=Buprestis eremita, Maoriella eremita, Nascio eremita, Neocuris eremita; incl. *Maoriella novaezeelandiae Obenberger 1924, Maoraxia novaezeelandiae, Maoriella strandi Obenberger 1936, Maoraxia strandi] BW85

Trigonogenium Gemminger & Harold 1869 [incl. Trigonophorus Solier 1851 non Stephens 1829; Trigonogenini] B85

*Type species of generic name indicated

REFERENCES

[B85] Bellamy, C. L. 1985. A catalogue of the higher taxa of the family Buprestidae (Coleoptera). Navorsinge van die Nasionale Museum Bloemfontein 4 (15): 405–472.

[BW85] Bellamy, C. L., & G. A. Williams. 1985. A revision of the genus Maoraxia with a new synonym in Acmaeodera (Coleoptera: Buprestidae). International Journal of Entomology 27 (1–2): 147–161.

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[G06] Golding, M. R. 2006. A Pictorial Field Guide to the Beetles of Australia. Part 1: Stigmoderini. Ocean Publishing.

[G08] Golding, M. R. 2008. A Pictorial Field Guide to the Beetles of Australia. Part 3: Chalcophorinae. Ocean Publishing.

[G09] Golding, M. R. 2009. A Pictorial Field Guide to the Beetles of Australia. Part 4: Bolboceratinae. Privately published.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2^nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part III. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 21–126.

[O86] Olliff, A. S. 1886. Contributions towards a knowledge of the Coleoptera of Australia. No. III.—On the genus Nascio (fam. Buprestidae). Proceedings of the Linnean Society of New South Wales, series 2, 1 (3): 861–864.

[P91] Peterson, M. 1991. A replacement name for Buprestis scalaris Gory & Laporte (Coleoptera: Buprestidae). Records of the Western Australian Museum 15 (1): 13–15.

[ZS10] Zborowski, P., & R. Storey. 2010. A Field Guide to Insects in Australia 3^rd ed. Reed New Holland: Sydney.

Last updated: 10 April 2022.

Cardiophorinae

Paracardiophorus musculus, copyright Hungarian Natural History Museum.

Belongs within: Elateridae.

The Cardiophorinae are a group of click beetles united by the presence of two sublateral carinae and incisions on the base of the pronotum, a truncate prosternal spine, a cordate scutellum, and larvae with exodont mandibles (Calder 1996).

Characters (from Calder 1996): Head more or less convex anteriorly. Frontoclypeal region raised above plane of labrum forming wide transverse region between basal articulations of antennae. Frontal carina complete across front between eyes. Pronotum usually wider than long, occasionally as wide as long. Base of pronotum with two sublateral carinae and incisions. Prosternal spine (posterior to procoxae) about same length as coxal diameter, truncate apically. Scutellum cordate with strongly pointed apex. Meso-trochantin not visible. Mesocoxal cavity closed to mesepisternum and mesepimeron. Hind wing with short radial cell; vein MP₄ without apparent cross vein to CuA₂; wedge cell absent; apex of wing membrane (without membrane) fairly long occupying at least one-quarter of length of wing membrane; apical sclerotisations various. Tarsomeres usually simple, tarsomere 4 strongly expanded laterally and cordiform in Cardiotarsus. Tarsal claws without basal setae on outer flat portion. Female genitalia with large glandular reservoir entering bursa copulatrix on mid-ventral surface.

<==Cardiophorinae [Cardiophorini, Esthesopinae]
    |--Cardiohypnus C96
    |--Agrypnella C96
    |--Patriciella Van Zwaluwenburg 1953 C96
    |    `--*P. australica (Van Zwaluwenburg 1953) [=*Patricia australica] C96
    |--Dicronychus [Dicronychidae] KB11
    |    |--D. cinereus KB11
    |    `--D. rubripes KB11
    |--Cardiotarsus Eschscholtz 1836 C96
    |    |--*C. capensis C96
    |    |--C. housaianus Kishii 1994 PS07
    |    |--C. mjobergi Carter 1939 C96
    |    `--C. taiwanus Kishii 1994 PS07
    |--Cardiophorus B14
    |    |--C. cinereus C01
    |    |--C. dilutus G89
    |    |--C. equiseti C01
    |    |--C. erichsoni KB11
    |    |--C. funebris G89
    |    |--C. hedenborgii G89
    |    |--C. jocularius G89
    |    |--C. notatus (Fabricius 1781) [=Elater notatus] B14
    |    |--C. rubripes C01
    |    |--C. rufipes C01
    |    `--C. sexmaculatus Lucas 1846 E12
    `--Paracardiophorus Schwarz 1895 C96
         |--*P. musculus [=Cardiotarsus musculus] C96
         |--P. alternatus Carter 1939 C96
         |--P. amabilis Carter 1939 C96
         |--P. antennalis Schwarz 1907 C96
         |--P. assimilis Carter 1939 C96
         |--P. atronotatus Carter 1939 C96
         |--P. attenuatipennis Elston 1930 C96
         |--P. australis (Candèze 1860) [=Horistonotus australis; incl. H. bicolor Rainbow 1904] C96
         |--P. bicolor (Candèze 1878) [=Cardiophorus bicolor] C96
         |--P. carissimus Carter 1939 C96
         |--P. compactus (Candèze 1882) [=Cardiophorus compactus] C96
         |--P. consobrinus (Candèze 1878) [=Horistonotus consobrinus] C96
         |--P. consputus (Candèze 1878) [=Cardiophorus consputus] C96
         |--P. cooki Carter 1939 C96
         |--P. despectus (Candèze 1882) [=Cardiophorus despectus] C96
         |--P. dimidiatus Schwarz 1902 C96
         |--P. dissimilis Schwarz 1903 C96
         |--P. divisus (Candèze 1865) [=Cardiophorus divisus] C96
         |--P. dulcis Carter 1939 C96
         |--P. elevatus (Van Zwaluwenburg 1947) [=Cardiophorus elevatus] C96
         |--P. elisus (Candèze 1865) [=Horistonotus elisus] C96
         |--P. eucalypti (Blackburn 1892) [=Cardiophorus eucalypti] C96
         |--P. flavipennis (Candèze 1878) [=Cardiophorus flavipennis] C96
         |--P. flavopictus (Carter 1939) [=Hypnoidus flavopictus] C96
         |--P. fulvosignatus (Candèze 1878) [=Cardiophorus fulvosignatus] C96
         |--P. hamatus (Candèze 1878) [=Cardiophorus hamatus] C96
         |--P. humilis (Candèze 1865) [=Cardiophorus humilis] C96
         |--P. jugulus Elston 1930 C96
         |--P. lenis (Candèze 1865) [=Horistonotus lenis] C96
         |--P. litoralis Carter 1939 C96
         |--P. longicornis (Candèze 1878) [=Horistonotus longicornis] C96
         |--P. macleayi (Schwarz 1907) (see below for synonymy) C96
         |--P. malkini (Van Zwaluwenburg 1947) [=Cardiophorus malkini] C96
         |--P. mastersii (Macleay 1872) [=Elater mastersii] C96
         |--P. minimus (Candèze 1878) [=Cardiophorus minimus] C96
         |--P. mjobergi Elston 1930 C96
         |--P. moseri Schwarz 1902 C96
         |--P. nigrosuffusus Carter 1939 C96
         |--P. occidentalis Carter 1939 C96
         |--P. octavus (Candèze 1878) [=Cardiophorus octavus] C96
         |--P. octosignatus Carter 1939 C96
         |--P. pallidipennis (Candèze 1878) [=Cardiophorus pallidipennis] C96
         |--P. quadripunctatus (Blanchard 1853) [=Agriotes quadipunctatus] C96
         |--P. quadristellatus Carter 1939 C96
         |--P. rufopictus Carter 1939 C96
         |--P. sexnotatus Carter 1939 C96
         |--P. stellatus Carter 1939 C96
         |--P. subcruciatus Carter 1939 C96
         |--P. subfasciatus Carter 1939 C96
         |--P. tumidithorax (Schwarz 1907) [=Cardiophorus tumidithorax, C. tumidicollis Schwarz 1907] C96
         |--P. vagus Schwarz 1907 C96
         |--P. varians Carter 1939 C96
         |--P. variegatus Schwarz 1902 C96
         |--P. venustus (Candèze 1860) [=Cardiophorus venustus] C96
         |--P. victoriensis (Blackburn 1892) [=Cardiophorus victoriensis] C96
         |--P. vittipennis Carter 1939 C96
         `--P. xanthomus (Candèze 1865) [=Horistonotus xanthomus] C96

Paracardiophorus macleayi (Schwarz 1907) [=Cardiophorus macleayi, C. quadrimaculatus Macleay 1888 non Motschulsky 1861] C96

*Type species of generic name indicated

REFERENCES

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C96] Calder, A. A. 1996. Click beetles: genera of the Australian Elateridae (Coleoptera). CSIRO Australia: Collingwood.

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 5–72.

[KB11] Kundrata, R., & L. Bocak. 2011. The phylogeny and limits of Elateridae (Insecta, Coleoptera): is there a common tendency of click beetles to soft-bodiedness and neoteny? Zoologica Scripta 40: 364–378.

[PS07] Platia, G., & R. Schimmel. 2007. Click beetles of Taiwan collected by the expeditions of the Hungarian Natural History Museum in the years 1995 to 2003 (Coleoptera: Elateridae). Annales Historico-Naturales Musei Nationalis Hungarici 99: 49–91.

Eucnemidae

Flightless eucnemids, photographed by S. E. Thorpe.

Belongs within: Elateroidea.

The Eucnemidae, false click beetles, are a group of beetles resembling the click beetles of the Elateridae in possessing a ventral click process between the prothorax and mesothorax but differing in having all five abdominal ventrites connate. Larvae are found in dead wood but their exact habits are uncertain (Lawrence & Britton 1991).

Characters (from Lawrence & Britton 1991): Elongate, cylindrical to slightly flattened, compact, usually finely pubescent and with acute posterior angles on pronotum; labrum concealed beneath clypeus; antennae eleven-segmented, sometimes received in grooves just beneath lateral pronotal carinae or ventrally crossing or bordering notosternal suture; fore coxae globular, trochantins concealed, fore coxal cavities externally and internally open; clicking device present involving prosternal process and mesosternal cavity; mid coxal cavities narrowly separated, laterally open; tarsi pentamerous, usually simple, rarely bearing membranous appendages on segments 2–4; abdomen with five ventrites, all connate. Larvae usually flattened, with no legs and reduced head with minute antennae and non-opposable or immovable mandibles.

Eucnemidae [Melasidae]
    |--Idiotarsus KB11
    `--+--+--Nematodes KB11
       |  `--Fornax additus Bon. 1872 LB91, Mas86
       `--+--Arrhipis KB11
          `--+--Protofarsus KB11
             `--Entomophthalmus americanus KB11

Eucnemidae incertae sedis:
  Neocharis E02
  Dromoeolus Mas86
    |--D. australasiae Bonv. 1871 Mas86
    |--D. dignoscendus Bonv. 1871 Mas86
    `--D. lugubris Bonv. 1871 Mas86
  Trigonopleurus rugolosus Bonv. 1875 Mas86
  Phyllocerus LB91
  Hemiopsida mastersi Macleay 1872 LB91, Mas86
  Euryptychus porosus LB91
  Palaeoxenus [Palaeoxeninae] B14
    `--P. dorhni (Horn 1878) B14
  Galbites [Eucneminae, Galbitini] B14
    |--G. albiventris B14
    |--G. auricolor (Bonvouloir 1875) B14
    |--G. australiae B14
    |--G. bicolor B14
    |--G. chrysocoma B14
    |--G. modiglianii B14
    |--G. sericata B14
    `--G. tigrina B14
  Galba Mac86
    |--G. auricolor Mac86
    |--G. marmorata [=Pterotarsus marmoratus] B35
    `--G. wallacei Perroud 1864 Mac86
  Cafolus moestus Mac86
  Arisus depressus Bonv. 1871 Mas86
  Anabolus mirus Bonv. 1871 Mas86
  Phoenocerus subclavatus Bonv. 1871 Mas86
  Galbodema Mas86
    |--G. mannerheimi Mas86
    `--G. ruficeps Bonv. 1872 Mas86
  Orodotes jansoni Bonv. 1875 Mas86
  Lycaon Mas86
    |--L. expulsus Bonv. 1875 Mas86
    |--L. kraatzi Bonv. 1875 Mas86
    |--L. nanus Bonv. 1875 Mas86
    |--L. nigricans Bonv. 1875 Mas86
    `--L. novus Bonv. 1875 Mas86
  Melasis RD77
    |--M. buprestoides RD77
    `--M. flabellicornis L02

*Type species of generic name indicated

REFERENCES

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[E02] Emberson, R. M. 2002. The beetle (Coleoptera) fauna of the Chatham Islands: additions and corrections. New Zealand Entomologist 25: 69–77.

[KB11] Kundrata, R., & L. Bocak. 2011. The phylogeny and limits of Elateridae (Insecta, Coleoptera): is there a common tendency of click beetles to soft-bodiedness and neoteny? Zoologica Scripta 40: 364–378.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2^nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[Mac86] Macleay, W. 1886. The insects of the Fly River, New Guinea, "Coleoptera". Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 136–157.

[Mas86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part IV. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 259–380.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms' General Textbook of Entomology 10^th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

Last updated: 10 December 2020.

Cantharidae

Plague soldier beetle Chauliognathus lugubris, copyright Fir0002/Flagstaffotos.

Belongs within: Elateroidea.
Contains: Heteromastix, Telephorus, Cantharis.

The Cantharidae are a group of soft-bodied beetles that are usually coloured in contrasting patterns of yellow and black, reflecting their production of distasteful defensive chemicals. Larvae are found in soil and leaf litter; most are predators though some are phytophagous (Lawrence & Britton 1991).

Characters (from Lawrence & Britton 1991): Elongate, parallel-sided, soft-bodied, usually in various combinations of yellow and black or bluish black; head relatively large and deflexed; labrum membranous, often inconspicuous; antennal insertions well separated; antenna eleven-segmented, filiform; pronotum oval or subquadrate and flattened; fore coxae projecting, trochantins exposed, fore coxal cavities externally and internally open; mid coxal cavities medially contiguous, laterally open; tarsi pentamerous, fourth tarsal segment bilobed; abdomen with seven or eight ventrites, none connate; male genitalia usually large, asymmetrical, exposed. Larva elongate, lightly sclerotised (although often darkly pigmented or patterned), with characteristic clothing of fine, velvety pile; stemmata well developed; mandibles more or less falcate; gular region well developed, without or with fused sutures; paired glandular openings on all thoracic and abdominal segments; segment 10 reduced and terminal.

Cantharidae [Cantharini, Telephorinae]
    |--+--Malthodes ruficollis MW15, C01
    |  `--Silis [Silinae] MW15
    |       |--S. bidentata (Say 1825) B14
    |       `--S. latiloba B14
    `--Chauliognathinae B14
         |--Trypherus B14
         |    |--T. babai B14
         |    `--T. rossicus (Barovsky 1922) B14
         `--Chauliognathus MW15
              |--C. apterus LB91
              |--C. lecontei B14
              |--C. lugubris LB91
              |--C. pennsylvanicus AT01
              |--C. profundus LeConte 1858 B14
              `--C. pulchellus B74

Cantharidae incertae sedis:
  Lygistopterus sanguineus C01
  Charopus plumbeomicans C01
  Dolichosoma lineare C01
  Telephorides [Telephoridae] L09
    |--Heteromastix LB91
    |--Telephorus L09
    `--Selenurus Fairm. 1879 L09
         |--S. annulatus Macl. 1887 L09
         |--S. apicalis Macl. 1887 L09
         |--S. appendiculatus Lea 1909 L09
         |--S. depressicollis (Macleay 1872) L09, M86 (see below for synonymy)
         |--S. granulatus Lea 1895 L09
         |--S. sydneyanus Blackb. 1892 L09
         |--S. tricolor Lea 1895 [incl. S. fernshawensis Blackburn 1900] L09
         |--S. variegatus Blackburn 1892 L09
         `--S. v-flavus Lea 1909 L09
  Sphaerarthrum LB91
  Malthinus KB11
    |--M. flaveolus KB11
    |--M. longipennis Lucas 1846 E12
    |--M. marginatus [=Thelephorus (Malthinus) marginatus] G20
    `--M. pulchellus Lucas 1846 E12
  Maltypus KB11
  Inmalthodes KB11
  0--Ichtyurus KB11
  `--+--+--Asiosilis KB11
     |  `--Laemoglyptus KB11
     `--+--Cantharis KB11
        `--Rhagonycha KB11
             |--R. atra C01
             |--R. femoralis C01
             |--R. limbata RD77
             `--R. testacea C01
  Palestrida bicolor M86
  Goetymes flavicornis Pascoe 1863 M86

Selenurus depressicollis (Macleay 1872) L09, M86 [=Ichthyurus depressicollis L09; incl. S. depressicollis var. luteopictus Fairm. 1877 L09]

*Type species of generic name indicated

REFERENCES

[AT01] Arnett, R. H., Jr & M. C. Thomas (eds) 2001. American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press: Boca Raton.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[B74] Britton, E. B. 1974. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers. Supplement 1974 pp. 62–89. Melbourne University Press.

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[KB11] Kundrata, R., & L. Bocak. 2011. The phylogeny and limits of Elateridae (Insecta, Coleoptera): is there a common tendency of click beetles to soft-bodiedness and neoteny? Zoologica Scripta 40: 364–378.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2^nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[L09] Lea, A. M. 1909. Revision of the Australian and Tasmanian Malacodermidae. Trans. Ent. Soc. Lond. 1909 (1): 45–251, pls 2–6.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part IV. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 259–380.

[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms' General Textbook of Entomology 10^th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

Last updated: 10 December 2020.

Lycidae

Macrolycus flabellatus, copyright Y. J. Chang.

Belongs within: Elateroidea.
Contains: Trichalus, Lycini, Metriorrhynchus.

The Lycidae are a group of soft-bodied beetles with narrow, dehiscent elytra that are generally coloured in contrasting aposematic patterns of reddish yellow and black. The distasteful lycids often function as models in mimicry complexes.

Lycidae [Lycides] B14
    |--Lyropaeinae B14
    |    |  i. s.: Platerodrilus korinchianus Blair 1928 B14
    |    |         Leptolycus B14
    |    |--+--Duliticola KB11
    |    |  `--+--Horakiella emasensis KB11
    |    |     `--+--Pendola KB11
    |    |        `--Macrolibnetis KB11
    |    `--+--+--Antennolycus constrictus KB11
    |       |  `--Microlyropaeus dembickyi KB11
    |       `--+--Alyculus kurbatovi KB11
    |          `--Lyropaeus KB11
    `--+--+--Thonalmus hubbardi KB11
       |  `--+--Lycoprogenthes KB11
       |     `--+--Lyponia quadricollis KB11
       |        `--Libnetis granicollis KB11
       `--Lycinae B14
            |--+--Dihammatus KB11
            |  `--Macrolycus KB11 [Macrolycini B14]
            |       `--M. flabellatus (Motschulsky 1860) B14
            |--+--+--Eurrhacus KB11
            |  |  `--Platycis minutus KB11
            |  `--+--Idiopteron biplagiatum KB11
            |     `--Lycini KB11
            `--+--Metriorrhynchus KB11
               `--+--+--Dilophotes KB11
                  |  `--Scarelus KB11
                  `--+--+--Benibotarus nigripennis KB11
                     |  `--Pyropterus nigroruber KB11
                     `--+--Taphes brevicollis KB11
                        `--Dictyoptera KB11
                             |--D. dichroma Boisduval 1835 B35
                             `--D. elegans KB11

Lycidae incertae sedis:
  Dumbrellia Lea 1909 L09
    |--D. brevicornis (Lea 1898) [=Calochromus brevicornis] L09
    `--D. pilosicornis (Lea 1898) [=Calochromus pilosicornis] L09
  Trichalus L09
  Xylobanus L09
    |--X. costifer L09
    |--X. gratiosus L09
    |--X. obscurus Macleay 1886 M86
    `--X. regularis M86
  Melampyrus L09
  Calopteron flavicans M86
  Cladophorus M86
    |--C. apicalis Macleay 1886 M86
    |--C. ingenuus M86
    |--C. longicornis Macleay 1886 M86
    |--C. nigrescens Macleay 1886 M86
    `--C. semirufus Macleay 1886 M86

*Type species of generic name indicated

REFERENCES

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[KB11] Kundrata, R., & L. Bocak. 2011. The phylogeny and limits of Elateridae (Insecta, Coleoptera): is there a common tendency of click beetles to soft-bodiedness and neoteny? Zoologica Scripta 40: 364–378.

[L09] Lea, A. M. 1909. Revision of the Australian and Tasmanian Malacodermidae. Trans. Ent. Soc. Lond. 1909 (1): 45–251, pls 2–6.

[M86] Macleay, W. 1886. The insects of the Fly River, New Guinea, "Coleoptera". Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 136–157.

Artematopodidae

Eurypogon niger, copyright Katja Schulz.

Belongs within: Elateroidea.

The Artematopodidae are a small but widespread group of beetles with an unusual tongue-like structure on the ventral side of the elytra near the apex (Bouchard 2014).

Artematopodidae [Artematopodinae, Eurypogonidae] B14
    |--Electribius GE05
    |--Carcinognathus B14
    |--Macropogon B14
    |--Proartematopus B14
    |--Electrapate B14
    `--Eurypogon B14
         |--E. brevipennis B14
         |--E. californicus B14
         |--E. harrisii B14
         |--E. japonicus B14
         `--E. niger (Melsheimer 1846) B14

*Type species of generic name indicated

REFERENCES

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.