Carabinae

Forest caterpillar hunter Calosoma sycophanta, copyright Thomas Huntke.

Belongs within: Carabidae.
Contains: Pamborus, Carabus.

The Carabinae are a group of ground beetles that, in the sense used herein, include species with open procoxal and mesocoxal cavities (other classifications expand the 'Carabinae' to cover a much larger, probably non-monophyletic subgroup of the Carabidae). In members of the tribe Carabini, the mandibles lack teeth and have a rugose upper surface, and the clypeus possesses a setigerous puncture at each anterior corner. Species of Calosoma have antennae with the second segment globose and distinct carinae along the dorsal margin of the second and third segments, and mandibles with transversely corrugate surfaces. Members of this genus climb vegetation in search of tree-dwelling larvae of insects such as moths and sawflies, and may play a role in the control of defoliators (Ball & Bousquet 2001).

Members of the Cychrini have smooth and toothed mandibles, and lack prominent setae on the clypeus (Lawrence & Britton 1991). They are molluscivores, inhabiting cool, damp woods and ravines. Within Cychrini, Scaphinotus differs from the genera Sphaeroderus and Cychrus in possessing at least a partial suture separating the proepisternum and proepimeron (Ball & Bousquet 2001).

Carabinae [Carabitae]
    |  i. s.: Ceroglossus chilensis (Eschscholtz 1829) B14
    |         Damaster blaptoides Kollar 1836 B14
    |         Acoptolabrus B14
    |         Coptolabrus B14
    |--Cychrini [Cychrinae, Pamborinae] LB91
    |    |--Sphaeroderus Dejean 1826 BB01
    |    |--Cychropsis Boileau 1901 BB01
    |    |--Pamborus LB91
    |    |--Cychrus Fabricius 1794 BB01
    |    |    |--C. attenuatus B14
    |    |    |--C. caraboides Linnaeus 1758 B14
    |    |    |--C. cordicollis B14
    |    |    `--C. rostratus [=Carabus (Cychrus) rostratus] G20
    |    `--Scaphinotus Dejean 1826 BB01
    |         |--S. (Scaphinotus) [incl. Irichroa Newman 1838, Megaliridia Casey 1920] BB01
    |         |    |--S. (S.) elevatus Fabricius 1787) BB01
    |         |    |    |--S. e. elevatus BB01
    |         |    |    `--S. e. neomexicanus Van Dyke 1924 BB01
    |         |    `--S. (S.) petersi BB01
    |         |--S. (Brennus Motschulsky 1865) BB01
    |         |    `--S. (B.) marginatus (Fischer von Waldheim 1822) BB01
    |         |--S. (Maronetus Casey 1914) BB01
    |         |    |--S. (M.) debilis (LeConte 1853) BB01
    |         |    |--S. (M.) hubbardi (Schwarz 1895) BB01
    |         |    |--S. (M.) imperfectus (Horn 1860) BB01
    |         |    `--S. (M.) incompletus (Schwarz 1895) BB01
    |         |--S. (Neocychrus Roeschke 1907) BB01
    |         |    `--S. (N.) angulatus BB01
    |         |--S. (Nomaretus LeConte 1853) BB01
    |         |--S. (Pseudonomaretus Roeschke 1907) BB01
    |         |--S. (Steniridia Casey 1924) BB01
    |         |    `--S. (S.) andrewsii BB01
    |         `--S. (Stenocantharus Gistel 1857) [incl. Pemphus Motschulsky 1866] BB01
    |              `--S. (S.) angusticollis (Fischer von Waldheim 1823) BB01
    `--Carabini LB91
         |--Carabus BB01
         `--Calosomatina BB01
              |--Aplothorax Waterhouse 1843 BB01
              `--Calosoma Weber 1801 BB01 [=Callisoma Agassiz 1846 LM87]
                   |  i. s.: C. australe Hope 1845 LM87
                   |         C. cyaneoventre Mandl 1955 LM87
                   |         C. inquisitor L02
                   |--C. (Calosoma) [incl. Callipara Motschulsky 1865, Syncalosoma Breuning 1927] BB01
                   |    |--*C. (C.) sycophanta (Linnaeus 1758) [=Carabus sycophanta] LM87
                   |    `--C. (C.) frigidum Kirby 1837 BB01
                   |--C. (Australodrepa Lapouge 1929) LM87
                   |    |--C. (A.) oceanicum Perroud & Montrouzier 1864 (see below for synonymy) LM87
                   |    `--C. (A.) schayeri Erichson 1842 (see below for synonymy) LM87
                   |--C. (Blaptosoma Géhin 1876) [incl. Aulacopterum Géhin 1885] BB01
                   |    `--C. (B.) haydeni Horn 1870 BB01
                   |--C. (Callisthenes Fischer von Waldheim 1821) BB01
                   |--C. (Callitropa Motschulsky 1865) [incl. Paratropa Lapouge 1929] BB01
                   |    |--C. (C.) externum (Say 1823) BB01
                   |    `--C. (C.) macrum LeConte 1853 BB01
                   |--C. (Calodrepa Motschulsky 1865) BB01
                   |    |--C. (C.) aurocinctum Chaudoir 1850 BB01
                   |    |--C. (C.) scrutator (Fabriciys 1775) BB01
                   |    |--C. (C.) splendidum Dejean 1831 BB01
                   |    `--C. (C.) wilcoxi LeConte 1848 BB01
                   |--C. (Camedula Motschulsky 1865) [incl. Acamegonia Lapouge 1924] BB01
                   |    `--C. (C.) peregrinator Guérin-Méneville 1844 BB01
                   |--C. (Camegonia Lapouge 1924) BB01
                   |    |--C. (C.) marginale Casey 1897 BB01
                   |    `--C. (C.) prominens BB01
                   |--C. (Carabosoma Géhin 1885) BB01
                   |    `--C. (C.) angulatum Chevrolat 1834 BB01
                   |--C. (Castrida Motschulsky 1865) (see below for synonymy) BB01
                   |    `--C. (C.) sayi Dejean 1826 BB01
                   |--C. (Chrysostigma Kirby 1837) [incl. Lyperostenia Lapouge 1929] BB01
                   |    `--C. (C.) calidum (Fabricius 1775) BB01
                   |--C. (Microcallisthenes Apfelbeck 1918) [incl. Callistenia Lapouge 1929, Isotenia Lapouge 1929] BB01
                   |    |--C. (M.) luxatum Say 1823 BB01
                   |    `--C. (M.) moniliatum LeConte 1852 BB01
                   |--C. (Tapinosthenes Kolbe 1895) BB01
                   |    `--C. (T.) cancellatum Eschscholtz 1833 BB01
                   `--C. (Teratexis Semenov & Znojko 1933) BB01

Calosoma (Australodrepa) oceanicum Perroud & Montrouzier 1864 [incl. C. (*A.) timorense Chaudoir 1869, C. walkeri Waterhouse 1898] LM87

Calosoma (Australodrepa) schayeri Erichson 1842 [incl. C. curtisi Hope 1845, C. grandipenne Castelnau 1867] LM87

Calosoma (Castrida Motschulsky 1865) [incl. Acampalita Lapouge 1929, Calamata Motschulsky 1865, Callistriga Motschulsky 1865, Catastriga Lapouge 1929] BB01

*Type species of generic name indicated

REFERENCES

[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2^nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Last updated: 19 December 2020.

Brachininae

Pheropsophus verticalis, copyright Peter Halasz.

Belongs within: Conjunctae.

The Brachininae, bombadier beetles, are a group of ground beetles capable of explosively emitting a defensive spray from the abdominal apex. They are most diverse in the tropics, particularly in Africa. Larvae of at least some species are ectoparasitoids of other insects with Brachinus attacking pupae of water beetles and Pheropsophus feeding on eggs of mole crickets (Ball & Bousquet 2001).

Characters (from Ball & Bousquet 2001): Size small to large (total length ca. 2–30 mm); head capsule ventrally without antennal grooves; antennal insertions laterad, each between base of mandible and anterior margin of eye; antenna filiform, number of antennomeres 11; antennomeres 2–4 not nodose, 2–6 without unusually long setae; labrum with six to eight setae; mandible with scrobe unisetose or plurisetose, occlusal margin with one terebral tooth, retinacular teeth about size of terebral tooth, not enlarged and molariform; terminal palpomeres about cylindrical not securiform; pterothorax with mesonotal scutellum dorsally visible, not concealed beneath elytra; front coxal cavities closed, with projection of proepimeron inserted each side into cavity of prosternum; middle coxal cavities conjunct-confluent, and hind coxal cavities lobate (disjunct)-confluent; elytra striate (intervals and striae broad) to smooth dorsally, extended to apex of abdominal tergum VI, and apical margins truncate; without flange of Coanda; metathoracic wings large to variously reduced; front tibia anisochaete, antenna cleaner emarginate, type C; abdomen with seven (II–VIII) or eight (II–IX) sterna normally exposed, sterna III and IV separated by suture; male genitalia with parameres glabrous, reduced, right minute, left larger, and in adults of most species baldric-like in position, across basal part of median lobe; ovipositor with gonocoxae rotated 90° in repose, without ramus, gonocoxa with two gonocoxites, gonocoxite 2 without accessory lobe; reproductive tract with spermatheca 1 only; secretion of pygidial glands quinones, delivered hot; discharge by crepitation.

<==Brachininae [Brachinides, Brachinini] B12
    |  i. s.: Mastax B55
    |           |--M. burgeoni B55
    |           `--M. ornatellus B55
    |         Brachynillus varendorffi Reitter 1904 BB01
    |--+--Aptinus OM08
    |  |    |--A. alpinus OM08
    |  |    `--A. displosor OM08
    |  `--+--Styphlodromus OM08
    |     `--Styphlomerus OM08
    |          |--S. ludicrus [incl. S. ludicrus var. grandis] B55
    |          `--S. vittaticollis OM08
    `--+--Crepidogaster [Crepidigastrini] OM08
       |    |--C. (Tyronia) ambreana OM08
       |    |--C. atrata OM08
       |    `--C. picipennis B55
       `--+--Brachinus Weber 1801 OM08, BB01
          |    |  i. s.: B. bimaculatus B35
          |    |         B. caligatus Bates 1889 B89
          |    |         B. frontalis B89
          |    |         B. oblongus G89
          |    |--B. (Brachinus) crepitans G20
          |    |--B. (Metabrachinus) armiger OM08 [incl. Metabrachinus armiger var. natalicus B55]
          |    `--B. (Neobrachinus Erwin 1970) BB01
          |         `--B. (N.) hirsutus OM08
          `--Pheropsophus Solier 1833 [Pherapsophina] B12
               |  i. s.: P. aequinoctialis (Linnaeus 1763) B14
               |         P. africanus [=Brachinus africanus] G89
               |         P. alexandrae Giachino 2003 B12
               |         P. arcanus B55
               |         P. darwini Giachino 2003 B12
               |         P. dimidiatus B55
               |         P. gregoryi Giachino 2003 B12
               |         P. jessoensis LB91
               |         P. livingstonei B55
               |         P. macleayi Sloane 1895 B12
               |         P. minor B55
               |         P. straneoi Giachino 2003 B12
               |         P. windjanae Baehr 2012 B12
               |--P. (Pheropsophus) B12
               |    |--*P. (P.) madagascariensis (Dejean 1831) [=Brachinus madagascariensis] B12
               |    `--P. (P.) complanatus (Fabricius 1775) LM87 [=Carabus complanatus LM87, Nebria complanata L02]
               `--P. (Stenaptinus Maindron 1906) [incl. Parapheropsophus Hubenthal 1914] LM87
                    |--P. (*S.) krichna (Maindron 1914) [=*Stenaptinus krichna] LM87
                    |--‘Stenaptinus’ insignis SBM11
                    `--P. (S.) verticalis (Dejean 1825) LM87 (see below for synonymy) LM87

Pheropsophus (Stenaptinus) verticalis (Dejean 1825) LM87 [=Brachinus verticalis LM87, P. (*Parapheropsophus) verticalis LM87; incl. P. australis Castelnau 1867 LM87, P. baliothorax Heller 1910 LM87, P. hypoxanthus Chaudoir 1876 LM87, P. (Parapheropsophus) intermedius Hubenthal 1914 LM87, Aptinus jamissoni B35, P. (Pa.) niger Hubenthal 1914 LM87, P. papuensis Macleay 1877 LM87]

*Type species of generic name indicated

REFERENCES

[B12] Baehr, M. 2012. A new species of the genus Pheropsophus (Coleoptera: Carabidae: Brachininae) from northern Australia. Records of the Western Australian Museum 27 (1): 62–67.

[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B89] Bates, H. W. 1889. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 100–111.

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso “Esploratore” dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 5–72.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2^nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

[SBM11] Solomon, E. P., L. R. Berg & D. W. Martin (eds) 2011. Biology 9^th ed. Brooks/Cole Cengage Learning.

Last updated: 19 December 2020.

Pamborus

Snail-eating carabid Pamborus alternans, from the Queensland Museum.

Belongs within: Carabinae.

Pamborus is a genus of flightless ground beetles found in eastern Australia, characterised by a large, almost heart-shaped pronotum and short, strongly curved mandibles.

<==Pamborus Latreille 1812 [incl. Callimosoma Castelnau 1834; Pamborini] LM87
    |--*P. alternans Latreille 1812 (see below for synonymy) LM87
    |--P. brisbanensis Castelnau 1867 LM87
    |--P. elegans Sloane 1915 LM87
    |--P. guerinii Gory 1831 [=*Callimosoma guerini] LM87
    |--P. macleayi Castelnau 1867 LM87
    |--P. opacus Géhin 1885 LM87
    |--P. pradieri Chaudoir 1869 LM87
    |    |--P. p. pradieri LM87
    |    `--P. p. darlingtoni Bänninger 1940 LM87
    |--P. punctatus Darlington 1961 LM87
    |--P. subtropicus Darlington 1961 LM87
    |--P. transitus Darlington 1961 LM87
    |--P. tropicus Darlington 1961 LM87
    `--P. viridis Gory 1836 LM87

*Pamborus alternans Latreille 1812 [incl. P. cunninghamii Castelnau 1834, P. elongatus Gory 1836, P. morbillosus Boisduval 1835, P. viridiaureus Macleay 1883] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Scopodes

Scopodes boops, copyright Lynne Forster.

Belongs within: Odacanthini.

Scopodes is a genus of ground beetles found in Indonesia and Australasia, characterised by large, strongly protruding eyes and a pentagonal pronotum.

<==Scopodes Erichson 1842 [incl. Molpus Newman 1842] LM87
    |--*S. boops Erichson 1842 [incl. *Molpus sexpunctus Newman 1842] LM87
    |--S. aeneus Macleay 1871 LM87
    |--S. angulicollis Macleay 1871 [incl. S. rimosicollis Sloane 1903] LM87
    |--S. aterrimus Chaudoir 1872 [incl. S. sydneyensis Sloane 1903] LM87
    |--S. denticollis Macleay 1864 [incl. S. sexfoveatus Macleay 1888] LM87
    |--S. elaphroides M83
    |--S. flavipes Blackburn 1894 [incl. S. lineatus Lea 1908] LM87
    |--S. griffithi Sloane 1903 LM87
    |--S. intermedius Blackburn 1894 LM87
    |--S. intricatus Blackburn 1895 LM87
    |--S. laevis Macleay 1871 LM87
    |--S. ocellatus Moore 1963 LM87
    |--S. rugatus Blackburn 1894 LM87
    |--S. sigillatus Germar 1848 [incl. S. sericeus Macleay 1871 LM87
    |--S. simplex Blackburn 1894 LM87
    |--S. splendens Moore 1963 LM87
    |--S. tasmanicus Bates 1878 LM87
    `--S. tripunctatus Chaudoir 1852 LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[M83] Martin, N. A. 1983. Miscellaneous observations on a pasture fauna: an annotated species list. DSIR Entomology Division Report 3: 1–98.

Odacanthini

Pentagonica vittipennis, copyright M. P. Navest.

Belongs within: Harpalinae.
Contains: Scopodes.

The Odacanthini are a cosmopolitan group of ground beetles characterised by a short first antennal segment, not forming a distinct scape, together with the head being posteriorly narrowed to a 'neck' and the prothorax being subcylindrical. Many Odacanthini produce formic acid for defense but some genera such as Pentagonica produce saturated acids instead.

<==Odacanthini [Pentagonicinae, Pentagonicini, Pentagonicitae]
    |--Pentagonica Schmidt-Goebel 1846 OM08, LM87 [incl. Wakefieldia Broun 1880 LM87]
    |    |--*P. ruficollis Schmidt-Goebel 1846 [incl. P. dichroa Sloane 1903] LM87
    |    |--P. atkinsoni Fauvel 1882 LM87
    |    |--P. blanda Andrewes 1929 LM87
    |    |--P. conradti B55
    |    |--P. elegans B55
    |    |--P. erichsoni Schmidt-Goebel 1846 LM87
    |    |--P. olivacea Chaudoir 1877 LM87
    |    |--P. pallipes (Nietner 1856) [=Elliotia pallipes] LM87
    |    |--P. roedingeri OM08
    |    `--P. vittipennis Chaudoir 1877 [incl. *Wakefieldia vittata Broun 1880] LM87
    `--+--+--+--Actenonyx bembidiodes OM08
       |  |  `--Scopodes OM08
       |  `--+--Colliuris Degeer 1774 OM08, LM87 [=Casnonia Latreille & Dejean 1823 LM87]
       |     |    |--*C. surinamensis (Linnaeus 1766) [=Attelabus surinamensis, *Casnonia surinamensis] LM87
       |     |    |--C. longicollis [=Collyris longicollis] L02
       |     |    |--C. par Darlington 1968 LM87
       |     |    `--C. pennsylvanica OM08
       |     `--Casnoidea Castelnau 1834 LM87 [incl. Ophionea Eschscholtz 1829 non Klug 1821 OM08, LM87]
       |           |--*C. cyanocephala (Fabricius 1798) [=Cicindela cyanocephala] LM87
       |           |--‘Attelabus’ indica Thunberg 1764 [=*Ophionea indica] LM87
       |           |--‘Ophionea’ ishii OM08
       |           |--‘Ophionea’ interstitialis B14
       |           |--‘Ophionea’ nigrofasciata B14
       |           |--C. puncticollis (Sloane 1923) [=Ophionea puncticollis] LM87
       |           `--C. thouzeti (Castelnau 1867) [=Ophionea thouzeti] LM87
       `--+--Celaenephes Schmidt-Goebel 1846 OM08, LM87 [incl. Taromorpha Blackburn 1894 LM87]
          |    |--*C. parallelus Schmidt-Goebel 1846 LM87
          |    `--C. linearis (Walker 1858) [=Leistus linearis; incl. *Taromorpha alternata Blackburn 1894] LM87
          `--+--Lasiocera OM08
             `--+--+--Clarencia Sloane 1917 OM08, LM87
                |  |    |--*C. aliena (Pascoe 1860) (see below for synonymy) LM87
                |  |    `--C. quadridens Darlington 1968 LM87
                |  `--Dicraspeda Chaudoir 1862 OM08, B12
                |       |--*D. brunnea Chaudoir 1862 [incl. Eudalia sublaevis Macleay 1888] LM87
                |       |--D. brunneipennis (Sloane 1917) [=Eudalia brunneipennis] LM87
                |       |--D. dubia (Gestro 1879) [=Odacantha dubia] LM87
                |       |--D. nitida (Sloane 1917) [=Eudalia nitida] LM87
                |       `--D. obscura (Castelnau 1867) [=Casnonia obscura] LM87
                `--Stenidia rugucollis OM08

Odacanthini incertae sedis:
  Odacantha melanura L02
  Porocara Sloane 1917 B12
    `--*P. punctata Sloane 1917 LM87
  Eudalia Castelnau 1867 B12
    |--*E. latipennis (Macleay 1864) [=Odacantha latipennis; incl. E. waterhousii Castelnau 1867] LM87
    |--E. castelnaui Sloane 1910 LM87
    |--E. froggatti Macleay 1888 LM87
    |--E. macleayi Bates 1871 LM87
    |--E. nigra Sloane 1900 LM87
    `--E. obliquiceps Sloane 1917 LM87
  Parascopodes Darlington 1968 LM87
    `--*P. cyaneus (Sloane 1907) [=Scopodes cyaneus] LM87

*Clarencia aliena (Pascoe 1860) [=Casnonia aliena; incl. Ca. angusticollis Macleay 1888, Ca. australis Chaudoir 1862, Ca. clarensii Castelnau 1867] LM87

*Type species of generic name indicated

REFERENCES

[B12] Baehr, M. 2012. A new species of the genus Pheropsophus (Coleoptera: Carabidae: Brachininae) from northern Australia. Records of the Western Australian Museum 27 (1): 62–67.

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

Sarothrocrepis

Sarothrocrepis inquinata, from Insects of Tasmania.

Belongs within: Tetragonoderus.

Sarothrocrepis is a genus of arboreal carabid beetles found in insular south-east Asia and Australasia, characterised by long, serrulate tibial spurs and bilobate fourth tarsomeres.

Sarothrocrepis Chaudoir 1876 (see below for synonymy) LM87
    |--*S. corticalis (Fabricius 1801) LM87 (see below for synonymy)
    |--S. benefica (Newman 1842) (see below for synonymy) LM87
    |--S. civica (Newman 1840) [=Lebia civica, Ectroma civicum, Lebiomorpha civica] LM87
    |--S. dimidiata Macleay 1888 LM87
    |--S. elegans (Blackburn 1901) [=Ectroma elegans] LM87
    |--S. fasciata Macleay 1871 LM87
    |--S. fragilis (Blackburn 1901) [=Ectroma fragile] LM87
    |--S. gravis (Blackburn 1901) [=Ectroma grave] LM87
    |--S. humerata Sloane 1900 [=S. humeratus] LM87
    |--S. inquinata (Erichson 1842) [=Cymindis inquinata, Ectroma inquinatum] LM87
    |--S. liturata Macleay 1888 LM87
    |--S. luctuosa (Newman 1842) [=Lebia luctuosa] lM87
    |--S. mastersii Macleay 1871 LM87
    |--S. mucronata Sloane 1907 LM87
    |--S. notabilis Macleay 1888 LM87
    |--S. notata Macleay 1888 LM87
    |--S. obsoleta (Blackburn 1892) [=Ectroma obsoletum; incl. S. blackburni Sloane 1911] LM87
    |--S. obtusa Sloane 1917 LM87
    |--S. pallida Macleay 1871 LM87
    |--S. parvicollis (Blackburn 1894) [=Ectroma parvicolle] LM87
    |--S. posticalis (Guérin-Méneville 1830) [=Lebia posticalis] LM87
    |--S. setulosa Sloane 1911 LM87
    |--S. suavis Blackburn 1890 LM87
    `--S. tridens (Newman 1840) [=Dromius tridens, Ectroma tridens] LM87

Sarothrocrepis Chaudoir 1876 [incl. Ectroma Blackburn 1890 non Westwood 1833, Lebiomorpha Chaudoir 1876] LM87

Sarothrocrepis benefica (Newman 1842) [=Lebia benefica, Lebiomorpha benefica; incl. Lebia duponti Putzeys 1845] LM87

*Sarothrocrepis corticalis (Fabricius 1801) LM87 [=Carabus corticalis LM87, Lebia corticalis B35; incl. L. callida Newman 1842 LM87, Sarothrocrepis corticalis infuscata Sloane 1916 LM87]

*Type species of generic name indicated

REFERENCES

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Coptocarpus

Coptocarpus philipi, copyright David R. Madison.

Belongs within: Harpalinae.

Coptocarpus is a genus of ground beetles found in Australia and New Guinea, characterised by asymmetrical fore tarsi in the male and a lack of ambulatory setae on the abdominal sterna.

<==Coptocarpus Chaudoir 1857 LM87
    |--*C. australis (Dejean 1831) (see below for synonymy) LM87
    |--C. championensis Chaudoir 1882 LM87
    |--C. chaudoiri Macleay 1873 LM87
    |--C. doddi Sloane 1910 LM87
    |--C. fuscitarsis (Blanchard 1853) [=Oodes fuscitarsis] LM87
    |--C. gibbus Chaudoir 1882 LM87
    |--C. grossus Erwin 1974 LM87
    |--C. impar Sloane 1910 LM87
    |--C. nitidus Macleay 1873 [incl. C. oviformis Chaudoir 1882] LM87
    |--C. philipi Erwin 1974 LM87
    |--C. thoracicus (Castelnau 1867) [=Oodes thoracicus; incl. C. parvus Sloane 1915] LM87
    `--C. yorkensis Erwin 1974 LM87

*Coptocarpus australis (Dejean 1831) [=Oodes australis; incl. O. convexus Castelnau 1867, O. reichei LaFerté-Sénectère 1851] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.