Craspedophorus

Craspedophorus sublaevis, copyright Peter Geissler.

Belongs within: Panagaeitae.

Craspedophorus is a genus of ground beetles widespread in the Palaeotropics, species of which have a distinctive colour pattern of four bright spots on the elytra and are often flightless. This coloration reflects the beetle's glandular production of defensive chemicals that are held in large reservoirs in the abdomen.

Craspedophorus Hope 1838 [incl. Epicosmus Chaudoir 1846, Eudema Castelnau 1840, Isotarsus LaFerté-Sénectère 1851] LM87
    |--*C. reflexus (Fabricius 1781) [=Carabus reflexus] LM87
    |--C. alternans (Castelnau 1867) [=Eudema alternans] LM87
    |--C. angulatus (Fabricius 1781) B14
    |--C. angusticeps Sloane 1923 LM87
    |--C. australasiae (Chaudoir 1850) [=Epicosmus australasiae; incl. E. froggatti Sloane 1903 LM87
    |--C. australis (Dejean 1831) [=Panagaeus australis; incl. Epicosmus mastersi Sloane 1903] LM87
    |--C. banksi Sloane 1923 LM87
    |--C. clasispila B55
    |--C. comptus (LaFerté-Sénectère 1851) [=Isotarsus comptus; incl. Epicosmus corpulentus Chaudoir 1879] LM87
    |--C. elongatus (Castelnau 1867) [=Eudema elongata] LM87
    |--‘Epicosmus’ feae Bates 1889 B89
    |--C. hexagonus B14
    |--C. impictus B55
    |--C. insignis (Schaum 1853) [=Isotarsus insignis; incl. C. convexus Macleay 1864] LM87
    |--C. longicollis (Chaudoir 1879) [=Epicosmus longicollis] LM87
    |--C. macleayi (Sloane 1903) [=Epicosmus macleayi, E. nobile Macleay 1888 non Dejean 1826] LM87
    |--C. ornatus B55
    |--C. parvulus (Macleay 1888) [=Eudema parvulum] LM87
    |--C. pubiger B14
    |--‘Panageus’ regalis Gory 1833 [=*Eudema regale, *Isotarsus regalis] LM87
    |--C. rikatlensis B55
    |--C. rockhamptonensis (Castelnau 1867) [=Eudema rockhamptonensis; incl. Epicosmus obesulus Sloane 1903] LM87
    |--C. rufipalpis OM08
    |--C. sublaevis B14
    |--‘Panageus’ tormentosus Vigors 1825 [=*Epicosmus tormentosus] LM87
    `--‘Epicosmus’ transversalis B89

*Type species of generic name indicated

REFERENCES

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B89] Bates, H. W. 1889. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 100–111.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

Last updated: 3 November 2018.

Lacordairia

Male Lacordairia insulicola from Moore (1985).

Belongs within: Licinini.

Lacordairia is a genus of flightless ground beetles found in southeastern Australia, with a deeply emarginate labrum, short metepisterna, and hind tarsomeres that are sulcate on each side.

<==Lacordairia Castelnau 1867 [=Lacordairea (l. c.)] LM87
    |--*L. cychroides Castelnau 1867 LM87
    |--L. anchomenoides Castelnau 1867 LM87
    |--L. angustata Castelnau 1867 LM87
    |--L. argutoroides Castelnau 1867 LM87
    |--L. calathoides Castelnau 1867 LM87
    |--L. erichsoni Castelnau 1867 LM87
    |--L. fugax (Olliff 1889) [=Lestignathus fugax] LM87
    |--L. insulicola Moore 1985 LM87
    |--L. proxima Castelnau 1867 LM87
    `--L. terrena Olliff 1885 LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Licinini

Dicaelus purpuratus, copyright Betsy Betros.

Belongs within: Harpalinae.
Contains: Lacordairia.

The Licinini are a cosmopolitan group of ground beetles characterised by a deep notch on the labrum.

<==Licinini [Licininae] M80
    |--+--Licinus Fabricius 1802 OM08, BR05
    |  |    |--L. cassideus [=Carabus cassideus] L02
    |  |    `--L. punctatulus [=Carabus punctatulus] L02
    |  `--Dicaelina OM08
    |       |--Diplocheila striatopunctata OM08
    |       `--Dicaelus OM08
    |            |--D. ambiguus OM08
    |            `--D. purpuratus Bonelli 1813 B14
    |                 |--D. p. purpuratus B14
    |                 `--D. p. splendidus B14
    `--+--Dicrochile Guérin-Méneville 1846 OM08, LM87 (see below for synonymy)
       |    |--*D. goryi (Boisduval 1835) (see below for synonymy) LM87
       |    |--D. brevicollis Chaudoir 1852 [=Dicronochilus brevicollis] LM87
       |    |--D. gigas Castelnau 1867 LM87
       |    |--D. idae Moore 1985 LM87
       |    |--D. minuta Castelnau 1867 LM87
       |    |--D. punctatostriata Castelnau 1867 LM87
       |    |--D. punctipennis Castelnau 1867 LM87
       |    |--D. punctulata Sloane 1923 LM87
       |    |--D. quadricollis Castelnau 1867 LM87
       |    `--D. ventralis Blackburn 1891 LM87
       `--+--Eutogeneius fuscus OM08
          `--Lestignathina OM08
               |--Lacordairia OM08
               `--Lestignathus Erichson 1842 LM87
                    |--*L. cursor Erichson 1842 LM87
                    |--L. foveatus Sloane 1920 LM87
                    `--L. simsoni Bates 1878 LM87

Licinini incertae sedis:
  Microferonia Blackburn 1890 M80, LM87
    |--*M. adelaidae Blackburn 1890 LM87
    |--M. anchomenoides (Macleay 1871) [=Badister anchomenoides, M. macleayi Csiki 1931] LM87
    |--M. cinctipennis Sloane 1898 LM87
    `--M. marginata (Castelnau 1867) [=Lacordairia marginata] LM87
  Physoloesthus Chaudoir 1850 M80, LM87 [=Physolaesthus LM87]
    |--*P. australis Chaudoir 1850 [=Physolaesthus australis] LM87
    |--P. caviceps (Andrewes 1936) [=Badister caviceps] LM87
    |--P. grandipalpis Macleay 1871 LM87
    |--P. insularis Bates 1878 [=Physolaesthus insularis] LM87
    |--P. pallidus Blackburn 1890 [=Physolesthus pallidus] LM87
    |--P. ruficollis Sloane 1900 [=Physolesthus ruficollis] LM87
    `--P. suturalis Castelnau 1867 [=Physoloestus suturalis] LM87

Dicrochile Guérin-Méneville 1846 OM08, LM87 [=Dicronochilus (l. c.) LM87; incl. Stomatocoelus Macleay 1864 LM87; Dicrochilina]

*Dicrochile goryi (Boisduval 1835) [=Rembus goryi; incl. *Stomatocoelus licinoides Macleay 1864, Dicrochile montana Castelnau 1867] LM87

*Type species of generic name indicated

REFERENCES

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[M80] Matthews, E. G. 1980. A Guide to the Genera of Beetles of South Australia. Part 1. Archostemata and Adephaga. South Australian Museum: Australia.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

Chlaenius

Chlaenius festivus, copyright Siga.

Belongs within: Oodini.

Chlaenius is a cosmopolitan genus of often metallic ground beetles with pubescent elytra that have the epipleura interrupted by a twisted fold near the apex. Members of the genus have been divided between over fifty subgenera and monophyly of the genus perhaps warrants investigation.

<==Chlaenius Bonelli 1810 [Chlaeniina] B12
    |--C. (Chlaenius) (see below for synonymy) BB01
    |    |--*C. (C.) festivus (Fabricius 1810) [=Carabus festivus] LM87
    |    |--C. (C.) aestivus Say 1823 BB01
    |    |--C. (C.) augustus Newman 1838 BB01
    |    |--C. (C.) cordicollis Kirby 1837 BB1
    |    |--C. (C.) cursor BB01
    |    |--C. (C.) laticollis BB01
    |    |--C. (C.) platyderus Chaudoir 1856 BB01
    |    |--C. (C.) ruficauda BB01
    |    |--C. (C.) sericeus (Forster 1771) BB01
    |    |--C. (C.) solitarius BB01
    |    `--C. (C.) viduus Horn 1871 BB01
    |--C. (Agostenus Motschulsky 1850) (see below for synonymy) BB01
    |    |--‘Agostenus’ glaucus BB01
    |    |--C. (A.) impunctifrons Say 1823 BB01
    |    |--‘Agostenus’ lithophilus BB01
    |    |--‘Agostenus’ nemoralis BB01
    |    |--‘Agostenus’ pennsylvanicus BB01
    |    |--C. (A.) purpuricollis Randall 1838 BB01
    |    |--C. (A.) tricolor Dejean 1826 BB01
    |    |--‘Agostenus’ vafer BB01
    |    `--‘Agostenus’ variabilipes BB01
    |--C. (Anomoglossus Chaudoir 1856) BB01
    |--C. (Diapheropsophus) B66
    |    |--C. (D.) mellyi B66
    |    `--C. (D.) swinhoei Bates 1866 B66
    |--C. (Epomis) B14
    |    |--C. (E.) circumscriptus (Duftschmid 1812) B14
    |    `--C. (E.) dejeanii B14
    |--C. (Eurydactylus LaFerté Sénectère 1851) [incl. Glyptoderus LaFerté Sénectère 1851] BB01
    |    |--C. (E.) pimalicus Casey 1914 BB01
    |    `--C. (E.) tomentosus (Say 1823) BB01
    |--C. (Lissauchenius) rufifemoralis OM08
    |    |--C. r. rufifemoralis OM08
    |    `--C. r. bimaculatus Dejean 1826 OM08, LM87
    |--C. (Pelasmomimus Grundmann 1955) LM87
    |    `--C. (*P.) greyanus White 1841 [=C. grayianus] LM87
    `--C. (Pseudanomoglossus Bell 1960) BB01
         `--C. (P.) maxillosus Horn 1876 BB01

Chlaenius incertae sedis:
  C. australis Dejean 1831 LM87
  C. darlingensis Castelnau 1867 LM87
  C. flaviguttatus Macleay 1825 (see below for synonymy) LM87
  C. hamifer Chaudoir 1856 [incl. C. bihamatus Chaudoir 1856, C. queenslandicus Sloane 1910] LM87
  C. maculiger Castelnau 1867 LM87
  C. nigricornis [incl. C. nigricornis var. melanocornis] C01
  C. nigripes Macleay 1886 M86
  C. ophonoides Fairmaire 1843 (see below for synonymy) LM87
  C. planicornis B66
  C. rudicollis Chaudoir 1876 [incl. C. bimaculatus Macleay 1864 non Dejean 1826] LM87
  C. spoliatus C01
  C. subcostatus Macleay 1864 LM87
  C. tenellus G89
  C. virgulifer C01

Chlaenius flaviguttatus Macleay 1825 [incl. Lissauchenius biguttatus Montrouzier 1860, C. binotatus Dejean 1826, C. guttatus Eschscholtz 1833, C. flaviguttatus immaculipennis Jedlicka 1951, C. maculifer Castelnau 1867, C. punctatus Chaudoir 1856, C. puncticeps Gemminger & Harold 1868] LM87

Chlaenius ophonoides Fairmaire 1843 [incl. C. peregrinus Chaudoir 1856, C. viridescens Gemminger & Harold 1868, C. viridis Montrouzier 1860 non Ménétries 1832] LM87

Chlaenius (Agostenus Motschulsky 1850) [inc. Brachylobus Chaudoir 1876, Chlaeniellus Reitter 1908, Pelasmus Motschulsky 1850] BB01

Chlaenius (Chlaenius) [incl. Callistometus Grundmann 1956, Hemichlaenius Lutschnik 1933 (preoc.), Lithochlaenius Kryzhanovskij 1976] BB01

*Type species of generic name indicated

REFERENCES

[B12] Baehr, M. 2012. A new species of the genus Pheropsophus (Coleoptera: Carabidae: Brachininae) from northern Australia. Records of the Western Australian Museum 27 (1): 62–67.

[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[B66] Bates, H. W. 1866. On a collection of Coleoptera from Formosa sent home by R. Swinhoe, Esq., H. B. M. Consul, Formosa. Proceedings of the Zoological Society of London 1866: 339–355.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2^a, 7: 5–72.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[M86] Macleay, W. 1886. The insects of the Fly River, New Guinea, "Coleoptera". Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 136–157.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

Last updated: 3 November 2018.

Oodini

Male Oodes helopioides, copyright U. Schmidt.

Belongs within: Harpalinae.
Contains: Panagaeitae, Chlaenius.

The Oodini are a group of ground beetles characterised by a twisted fold near the apex of the elytral epipleura (Bell 1990).

Characters (from Bell 1990): Mandibles usually pointed, not strongly curved; scrobal seta absent. Eye usually large; one orbital seta over each eye. Pronotum more or less flattened, broadest at or near base, basal impressions scarcely evident. Elytra glabrous; stria 8 much deeper than stria 7; interval 9 narrow, careniform; epipleura interrupted by twisted fold near apex. Metepimeron visible as scale-like structure overlapping abdominal sternum I at side.

<==Oodini [Oodinae] B90
    |  i. s.: Evolenes exarata B90
    |         Dercylinus impressus B90
    |         Stenomorphus B90
    |         Systolocranius validus B55
    |         Paroodes similatus B55
    |         Lonchosternus B55
    |           |--L. deceptor B55
    |           `--L. politus B55
    |--+--+--Anatrichis LeConte 1853 OM08, LM87
    |  |  |    |--*A. minutus (Dejean 1831) [=Oodes minutus] LM87
    |  |  |    |--A. australasiae Chaudoir 1882 [incl. Oodes pygmaeus Macleay 1888] LM87
    |  |  |    |--A. lilliputana (Macleay 1888) [=Oodes lilliputanus] LM87
    |  |  |    |--A. pusilla Sloane 1910 LM87
    |  |  |    `--A. sexstriata Sloane 1910 LM87
    |  |  `--Dercylus [Dercylini] OM08
    |  `--+--Adelopomorpha glabra OM08
    |     `--Panagaeitae OM08
    `--+--Stenocrepis elegans OM08
       `--+--Oodes Bonelli 1810 OM08, LM87
          |    |--*O. helopioides (Fabricius 1792) [=Carabus helopioides] LM87
          |    |--O. abaxoides Lucas 1846 E12
          |    |--O. amaroides OM08
          |    |--O. bostockii Castelnau 1867 LM87
          |    |--O. denisonensis Castelnau 1867 LM87
          |    |--O. fitzroyensis Macleay 1888 LM87
          |    |--O. froggatti Macleay 1888 LM87
          |    |--O. impressus Chaudoir 1882 LM87
          |    |--O. inornatus Castelnau 1867 [incl. O. proximus Castelnau 1867] LM87
          |    |--O. latior Csiki 1931 [=O. latus Castelnau 1867 non LaFerté 1851] LM87
          |    |--O. mauritanicus Lucas 1846 E12
          |    |--O. modestus Castelnau 1867 LM87
          |    |--O. oblongus Castelnau 1867 (see below for synonymy) LM87
          |    |--O. paroensis Castelnau 1867 LM87
          |    |--O. parviceps Sloane 1896 LM87
          |    |--O. trisulcatus Castelnau 1867 LM87
          |    `--O. waterhousei Castelnau 1867 [incl. O. interioris Castelnau 1867] LM87
          `--Chlaeniini [Chlaeniides, Chlaeniinae] OM08
               |--Chlaenius OM08
               |--Rhysotrachelus quadrimaculatus B55
               |--Ocybatus deyrollei B55
               |--Stenodinodes nepos B55
               |--Pachydinodes conformis B55
               |--Chlaenites capicola B55
               |--Pseudochlaeniellus paenulatus B55
               |--Callistoides B55
               |    |--C. hessei Basilewsky 1949 B55
               |    `--C. pulchellus B55
               |--Leptochlaenius varians B55
               |    |--L. v. varians B55
               |    |--L. v. angolanus Basilewsky 1955 B55
               |    `--L. v. verecundus B55
               |--Trachychlaenites B55
               |    |--T. gonioderus B55
               |    |    |--T. g. gonioderus B55
               |    |    `--T. g. nigrofemoralis B55
               |    `--T. obesus [incl. T. ovalipennis] B55
               |--Homalolachnus B55
               |    |--H. goossensi [incl. H. goossensi var. nigrocyanescens] B55
               |    `--H. lineatus B55
               |         |--H. l. lineatus B55
               |         `--H. l. tristis B55
               `--Chlaeniostenus B55
                    |--C. denticulatus B55
                    |    |--C. d. denticulatus B55
                    |    `--C. d. elatus B55
                    |--C. euryscopus B55
                    `--C. sulcipennis B55
                         |--C. s. sulcipennis B55
                         `--C. s. sulcatulus B55

Oodes oblongus Castelnau 1867 [incl. Coptocarpus planipennis Macleay 1878, C. riverinae Macleay 1873] LM87

*Type species of generic name indicated

REFERENCES

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B90] Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[OM08] Ober, K. A., & D. R. Maddison. 2008. Phylogenetic relationships of tribes within Harpalinae (Coleoptera: Carabidae) as inferred from 28S ribosomal DNA and the wingless gene. Journal of Insect Science 8: 63.

Agonocheila

Agonocheila curtula, copyright Simon Grove.

Belongs within: Harpalinae.

Agonocheila is an Australasian genus of ground beetles with a toothed mentum, two pairs of lateral setae on the pronotum, a conspicuously short-pubescent and roughened elytral surface, and simple tarsi with several teeth on the claws.

<==Agonocheila Chaudoir 1848 [=Agonochila (l. c.)] LM87
    |--*A. guttata Chaudoir 1848 LM87
    |--A. anomala Chaudoir 1869 [=Agonochila anomala] LM87
    |--A. biguttata Chaudoir 1869 [=Agonochila biguttata] LM87
    |--A. bimaculata Sloane 1920 [=Agonochila bimaculata] LM87
    |--A. binotata (White 1846) [=Lebia binotata] LM87
    |--A. chaudoiri Sloane 1898 [=Agonochila chaudoiri] LM87
    |--A. cribripennis Chaudoir 1869 [=Agonochila cribripennis] LM87
    |--A. curtula (Erichson 1842) [=Cymindis curtula; incl. Agonochila corticalis Chaudoir 1869] LM87
    |--A. fasciata Sloane 1898 [=Agonochila fasciata] LM87
    |--A. fenestrata Blackburn 1892 LM87
    |--A. flindersi Sloane 1920 [=Agonochila flindersi] LM87
    |--A. froggatti (Macleay 1888) [=Philophlaeus (l. c. for Philophloeus) froggatti] LM87
    |--A. irrita (Newman 1842) [=Lebia irrita] LM87
    |--A. koebelei Blackburn 1895 LM87
    |--A. lutosa (Newman 1840) [=Lebia lutosa; incl. Thyreopterus subangulatus Germar 1848] LM87
    |--A. macleayi Sloane 1911 [=Agonochila macleayi] LM87
    |--A. minima (Macleay 1864) (see below for synonymy) LM87
    |--A. mollis (Newman 1842) [=Lebia mollis] LM87
    |--A. ovalis Sloane 1923 [=Agonochila ovalis] LM87
    |--A. perplexa Blackburn 1894 LM87
    |--A. plagiata Sloane 1916 [=Agonochila plagiata] LM87
    |--A. punctulata Sloane 1911 [=Agonochila punctulata] LM87
    |--A. quadricollis Sloane 1911 [=Agonochila quadricollis] LM87
    |--A. ruficollis Sloane 1898 [=Agonochila ruficollis] LM87
    |--A. signata Moore 1963 [=Agonochila signata] LM87
    |--A. sinuosa Chaudoir 1869 [=Agonochila sinuosa] LM87
    |--A. stictica Blackburn 1895 LM87
    |--A. subfasciata Chaudoir 1869 [=Agonochila subfasciata] LM87
    |--A. sublaevis Chaudoir 1869 [=Agonochila sublaevis] LM87
    |--A. suturalis Macleay 1871 LM87
    `--A. vittula Chaudoir 1869 [=Agonochila vittula] LM87

Agonocheila minima (Macleay 1864) [=Sarathrocrepis (l. c. for Sarothrocrepis) minima; incl. Agonochila litterav Chaudoir 1869] LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

Philophloeus

Philophloeus luculentus, photographed by Stephen Thorpe.

Belongs within: Pericalina.

Philophloeus is an Australian genus of ground beetles with a long penultimate joint to the palpi, setulose-punctate interstices on the elytra, and simple, dorsally setulose tarsi with two or three joints in the males ventrally squamose.

Philophloeus Chaudoir 1844 [=Gomelina Blanchard 1853; incl. Idius Chaudoir 1869] LM87
    |--*P. australis (Dejean 1826) [=Cymindis australis, *Gomelina australis] LM87
    |--P. angulatus Chaudoir 1869 LM87
    |--P. australasiae Chaudoir 1869 LM87
    |--P. bivittatus Sloane 1920 LM87
    |--P. brunnipennis Macleay 1871 LM87
    |--P. confertus Blackburn 1892 LM87
    |--P. discorufus Sloane 1898 LM87
    |--P. distinguendus Chaudoir 1869 LM87
    |--P. eucalypti Germar 1848 [incl. P. grandiceps Chaudoir 1869, P. eucalypti tasmaniae Blackburn 1892] LM87
    |--P. fuscipennis Germar 1848 LM87
    |--P. immaculatus Chaudoir 1869 LM87
    |--P. intermedius Chaudoir 1869 LM87
    |--P. laticollis Blackburn 1892 LM87
    |--P. luculentus (Newman 1842) [=Lebia luculenta; incl. P. luculentus guttifera Sloane 1911] LM87
    |--P. maculatus Macleay 1871 LM87
    |--P. moestus (Chaudoir 1869) [=*Idius moestus] LM87
    |--P. monticola Blackburn 1892 LM87
    |--P. myrmecophilus Lea 1912 LM87
    |--P. obtusus Chaudoir 1869 LM87
    |--P. occidentalis Blackburn 1890 LM87
    |--P. opaciceps Blackburn 1890 LM87
    |--P. ornatus Blackburn 1892 [incl. P. truncatus Sloane 1898] LM87
    |--P. planus (Newman 1840) [=Lebia plana] LM87
    |--P. puberulus Chaudoir 1869 LM87
    |--P. pygmaeus Blackburn 1892 LM87
    |--P. quadripennis Chaudoir 1869 LM87
    |--P. rectangulus Chaudoir 1869 LM87
    |--P. simsoni Sloane 1920 LM87
    |--P. sydneyensis Blackburn 1892 LM87
    |--P. unicolor Chaudoir 1869 LM87
    `--P. vittatus Macleay 1871 LM87

*Type species of generic name indicated

REFERENCES

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.