Achelata

Southern rock lobster or crayfish Jasus edwardsii, copyright Stemonitis.

Belongs within: Reptantia.
Contains: Stridentes.

The Achelata, spiny and slipper lobsters, are a group of marine decapods characterised by the lack of chelae on most pereiopods.

Synapomorphies (from Dixon et al. 2003): Eyestalks with row of hairs alongside eye; second antenna greatly enlarged, basal articles of second antennae fused with epistome; pereiopods (except fifth) achelate. Carapace held to body by knob emanating from last thoracic somite; lineae absent; pleon with prominent hinges; pleurites pointed; tail fan soft distally; telson with fixed lateral spines. Phyllosoma larva present.

<==Achelata [Palinuroidea]
    |--Palinuridae DAS03
    |    |  i. s.: Palinustus G05b
    |    |         Palibythus G05b
    |    |--Stridentes G05b
    |    `--Silentes G05b
    |         |--Palinurellus [Synaxidae] G05b
    |         |    `--P. gundlachi von Martens 1878 DAS03
    |         |--Sagmariasus G05b
    |         |    |--S. flemingi G05b
    |         |    `--S. verreauxii G05b
    |         |--Projasus G05b
    |         |    |--P. bahamondei G05b
    |         |    `--P. parkeri G05b
    |         `--Jasus G05b
    |              |--J. edwardsii (Hutton 1875) H86
    |              |--J. frontalis G05b
    |              |--J. lalandei (Milne Edwards 1837) H86
    |              |--J. novaehollandiae G05b
    |              |--J. paulensis L95
    |              |--J. tristani G05b
    |              `--J. verreauxi MC05
    `--Scyllaridae DAS03
         |--Thenus [Theninae] BW05
         |    `--T. orientalis (Lund 1793) DAS03
         |--Arctidinae BW05
         |    |--Arctides guineensis BW05
         |    `--Scyllarides BW05
         |         |--S. aequinoctialis BW05
         |         |--S. astori BW05
         |         |--S. herklotsii BW05
         |         |--S. latus (Latreille 1803) KK03
         |         |--S. nodifer BW05
         |         `--S. squammosus BW05
         |--Ibacinae BW05
         |    |--Evibacus Smith 1869 BW05, B55
         |    |    `--E. princeps Smith 1869 [=Evibaccus (l. c.) princeps] B55
         |    |--Parribacus BW05
         |    |    |--P. antarcticus (Lund 1793) TSH09
         |    |    |--P. caledonicus BW05
         |    |    `--P. scarlatinus BW05
         |    `--Ibacus BW05
         |         |--I. alticrenatus Bate 1888 MG-H11
         |         |--I. chacei BW05
         |         |--I. ciliatus BW05
         |         |--I. novemdentatus BW05
         |         `--I. peronii Leach 1815 MG-H11
         `--Scyllarinae BW05
              |--Acantharctus BW05
              |--Antarctus BW05
              |--Antipodarctus aoteanus [=Scyllarus aoteanus] BW05
              |--Bathyarctus BW05
              |--Biarctus sordidus BW05
              |--Chelarctus cultrifer [=Scyllarus cultrifer] BW05
              |--Gibbularctus BW05
              |--Remiarctus bertholdii (Paulson 1875) BW05, MG-H11
              |--Scammarctus batei BW05
              |--Crenarctus BW05
              |    |--C. bicuspidatus [=Scyllarus bicuspidatus] BW05
              |    `--C. crenatus (Whitelegge 1900) MG-H11
              |--Eduarctus BW05
              |    |--E. martensii BW05
              |    `--E. modestus BW05
              |--Galearctus BW05
              |    |--G. kitanoviriosus BW05
              |    `--G. timidus BW05
              |--Petrarctus BW05
              |    |--P. demani [=Scyllarus demani] BW05
              |    `--P. rugosus BW05
              `--Scyllarus BW05
                   |--S. americanus (Smith 1869) C73
                   |--S. arctus (Linnaeus 1758) DAS03
                   |--S. chacei BW05
                   |--S. depressus BW05
                   |--S. planorbis BW05
                   `--S. pygmaeus (Bate 1888) KK03

*Type species of generic name indicated

REFERENCES

[BW05] Booth, J. D., W. R. Webber, H. Sekiguchi & E. Coutures. 2005. Diverse larval recruitment strategies within the Scyllaridae. New Zealand Journal of Marine and Freshwater Research 39 (3): 581–592.

[B55] Bott, R. 1955. Dekapoden (Crustacea) aus El Salvador. 2. Litorale Dekapoden, außer Uca. Senckenbergiana Biologica 36: 45–70.

[C73] Camp, D. K. 1973. Stomatopod Crustacea. Memoirs of the Hourglass Cruises 3 (2): 1–100.

[DAS03] Dixon, C. J., S. T. Ahyong & F. R. Schram. 2003. A new hypothesis of decapod phylogeny. Crustaceana 76: 935–975.

[G05] George, R. W. 2005. Evolution of life cycles, including migration, in spiny lobsters (Palinuridae). New Zealand Journal of Marine and Freshwater Research 39: 503–514.

[H86] Holthuis, L. B. 1986. Decapoda. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 589–615. E. J. Brill/Dr. W. Backhuys: Leiden.

[KK03] Kocataş, A., & T. Katağan. 2003. The decapod crustacean fauna of the Turkish seas. Zoology in the Middle East 29: 63–74.

[L95] Laubitz, D. R. 1995. Caprellidea (Crustacea: Amphipoda) from the southern and western Indian Ocean. Mésogée 54: 81–100.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O'Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[MC05] Montgomery, S. S., & J. R. Craig. 2005. Distribution and abundance of recruits of the eastern rock lobster (Jasus verreauxi) along the coast of New South Wales, Australia. New Zealand Journal of Marine and Freshwater Research 39 (3): 619–628.

[TSH09] Titelius, M. A., A. Sampey & C. G. Hass. 2009. Crustaceans of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, north Western Australia. Records of the Western Australian Museum Supplement 77: 145–176.

Galatheidae

Munida rugosa, copyright B. E. Picton.

Belongs within: Anomala.

The Galatheidae, squat lobsters, are a group of benthic or pelagic crustaceans somewhat similar to a lobster in superficial appearance but with the abdomen carried reflexed under the cephalothorax in the manner of a crab.

<==Galatheidae
    |  i. s.: Munidopsis Whiteaves 1874 H86
    |           |--M. andamanica MacGilchrist 1905 MG-H11
    |           |--M. comarge Taylor, Ahyong & Andreakis 2010 MG-H11
    |           |--M. crenatirostris Baba 1988 MG-H11
    |           |--M. cylindrophthalma (Alcock 1894) MG-H11
    |           |--M. dasypus Alcock 1894 MG-H11
    |           |--M. kensleyi Ahyong & Poore 2004 MG-H11
    |           |--M. levis (Alcock & Anderson 1894) MG-H11
    |           `--M. polymorpha Koelbel 1892 H86
    |         Allogalathea elegans (Adams & White 1848) MG-H11
    |         Agononida MG-H11
    |           |--A. eminens (Baba 1988) MG-H11
    |           |--A. incerta (Henderson 1888) MG-H11
    |           |--A. pilosimanus (Baba 1969) MG-H11
    |           |--A. rubrizonata Macpherson & Baba 2009 MG-H11
    |           `--A. similis (Baba 1988) MG-H11
    |         Enriquea leviantennata (Baba 1988) MG-H11
    |         Lauriea gardineri (Laurie 1926) MG-H11
    |         Paramunida stichas Macpherson 1993 MG-H11
    |         Phylladiorhynchus pusillus (Henderson 1885) MG-H11
    |         Raymunida MG-H11
    `--Galatheinae CA04
         |--Galathea MG-H11
         |    |--G. amboinensis de Man 1888 MG-H11
         |    |--G. balssi Miyake & Baba 1964 MG-H11
         |    |--G. bolivai Zariquiey-Alvarez 1950 KK03
         |    |--G. cenarroi (Zariquiey-Alvarez 1968) KK03
         |    |--G. consobrina De Man 1902 MG-H11
         |    |--G. dispersa Bate 1859 KK03
         |    |--G. intermedia Liljeborg 1851 KK03
         |    |--G. nexa Embleton 1834 KK03
         |    |--G. rostrata Milne Edwards 1880 HC79
         |    |--G. squamifera Leach 1814 DAS03
         |    `--G. strigosa (Linnaeus 1767) KK03
         `--Munida Leach 1820 [incl. Austromunida Schweitzer & Feldmann 2000] CA04
              |--*M. rugosa (Fabricius 1775) [=Pagurus rugosus] CA04
              |--M. andamanica Alcock 1894 MG-H11
              |--M. aprosoma Ahyong & Poore 2004 MG-H11
              |--M. babai Tirmizi & Javed 1976 MG-H11
              |--M. casadioi (Schweitzer & Feldmann 2000) [=*Austromunida casadioi] CA04
              |--M. disgrega Baba 2005 MG-H11
              |--M. gordoae Macpherson 2004 MG-H11
              |--M. haswelli Henderson 1885 MG-H11
              |--M. heteracantha Ortmann 1892 MG-H11
              |--M. intermedia Milne-Edwards & Bouvier 1899 KK03
              |--M. irrasa Milne Edwards 1880 HC79
              |--M. keiensis Baba 2005 MG-H11
              |--M. konara Schweitzer & Feldmann 2000 CA04
              |--M. primaeva (Segerberg 1900) CA04
              |--M. quadrispina H47
              |--M. quadroblonga Schweitzer & Feldmann 2000 CA04
              |--M. roshanei Tirmizi 1996 MG-H11
              |--M. rubridigitalis Baba 1994 MG-H11
              |--M. rugosa (Fabricius 1775) KK03
              |--M. rutlanti Zariquiey-Alvarez 1952 KK03
              `--M. tenuimana Sars 1872 KK03

*Type species of generic name indicated

REFERENCES

[CA04] Casadío, S., A. de Angeli, R. M. Feldmann, A. Garassino, J. L. Hetler, A. Parras & C. E. Schweitzer. 2004. New decapod crustaceans (Thalassinidea, Galatheoidea, Brachyura) from the Middle Oligocene of Patagonia, Argentina. Annals of Carnegie Museum 73 (2): 25–47.

[DAS03] Dixon, C. J., S. T. Ahyong & F. R. Schram. 2003. A new hypothesis of decapod phylogeny. Crustaceana 76: 935–975.

[H47] Hatch, M. H. 1947. The Chelifera and Isopoda of Washington and adjacent regions. University of Washington Publications in Biology 10 (5): 155–274.

[H86] Holthuis, L. B. 1986. Decapoda. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 589–615. E. J. Brill/Dr. W. Backhuys: Leiden.

[HC79] Huff, J. A., & S. P. Cobb. 1979. Penaeoid and sergestoid shrimps (Crustacea: Decapoda). Memoirs of the Hourglass Cruises 5 (4): 1–101.

[KK03] Kocataş, A., & T. Katağan. 2003. The decapod crustacean fauna of the Turkish seas. Zoology in the Middle East 29: 63–74.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O'Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

Coenobitoidea

Electric blue hermit Calcinus elegans, copyright Fafner.

Belongs within: Anomala.

The Coenobitoidea are a group of crab-like decapods that mostly live as hermit crabs, though the large coconut crab Birgus latro abandons the use of a protective shell as it grows to maturity.

Characters (from McLaughlin & Lemaitre 2001): Mandible with incisor process calcareous, lacking acute teeth; third maxillipeds generally approximate basally; chelipeds equal, subequal or unequal, left frequently largest; fourth pereopods not developed as walking legs; sternite XI fused to sternite XII.

<==Coenobitoidea
    |--Pylochelidae [Pomatochelidae] ML01
    |    |--Pylocheles mortensenii Boas 1926 MG-H11
    |    |--Parapylocheles Alcock 1901 ML01
    |    |--Cancellocheles Forest 1987 ML01
    |    `--Mixtopagurus Milne-Edwards 1880 ML01
    |         `--M. paradoxus Milne-Edwards 1880 ML01
    |--Coenobitidae TSH09
    |    |--Birgus latro (Linnaeus 1758) H86
    |    `--Coenobita Latreille 1829 B55
    |         |--C. brevimanus Dana 1852 DAS03
    |         |--C. clypeatus (Fabricius 1787) ML01
    |         |--C. diogenes (Fabricius 1775) [=Pagurus diogenes, Cenobita (l. c.) diogenes] B55
    |         |--C. perlatus Milne Edwards 1837 TSH09
    |         |--C. rugosus Milne-Edwards 1837 TSH09 [incl. C. compressa Milne-Edwards 1837 B55, C. compressus B55]
    |         `--C. spinosa [incl. C. spinosa var. olivieri] H09
    `--Diogenidae [Diogeninae] ML01
         |--Diogenes pugilator (Roux 1829) KK03
         |--Bathynarius L95
         |--Cancellus parfaiti L95
         |--Aniculus ursus (Olivier 1811) TSH09
         |--Strigopagurus elongatus Forest 1995 MG-H11
         |--Ciliopagurus MG-H11
         |    |--C. shebae (Lewinsohn 1969) MG-H11
         |    `--C. strigatus (Herbst 1804) TSH09
         |--Petrochirus Stimpson 1858 L95, B55
         |    |--P. diogenes L95
         |    `--P. granulatus B55
         |         |--P. g. granulatus B55
         |         `--P. g. californiensis Bouvier 1895 B55
         |--Clibanarius TSH09
         |    |--C. corallinus (Milne Edwards 1848) TSH09
         |    |--C. englaucus W01
         |    |--C. erythropus (Latreille 1818) KK03
         |    |--C. eurysternus (Hilgendorf 1878) TSH09
         |    |--C. padavensis H09
         |    |--C. panamensis Stimpson 1859 B55
         |    |--C. striolatus Dana 1852 TSH09
         |    |--C. virescens (Krauss 1843) TSH09
         |    `--C. vittatus Bosc 1802 ML01
         |--Dardanus Paulson 1875 MG-H11, H14
         |    |--D. arrosor (Herbst 1796) KK03
         |    |--D. callidus (Risso 1827) KK03
         |    |--D. crassimanus (Milne Edwards 1836) TSH09
         |    |--D. deformis (Milne Edwards 1836) TSH09
         |    |--D. gemmatus (Milne Edwards 1848) TSH09
         |    |--D. guttatus (Olivier 1811) TSH09
         |    |--D. lagopodes (Forskal 1775) TSH09
         |    |--D. megistos (Herbst 1804) TSH09
         |    |--D. pedunculatus (Herbst 1804) TSH09
         |    `--D. scutellatus (Milne Edwards 1848) TSH09
         |--Paguristes MG-H11
         |    |--P. aciculus Grant 1905 MG-H11
         |    |--P. barbatus HS01
         |    |--P. brevirostris Baker 1905 TSH09
         |    |--P. eremita (Linnaeus 1767) KK03
         |    |--P. longisetosus Morgan 1987 MG-H11
         |    |--P. pilosus HS01
         |    |--P. puncticeps L95
         |    |--P. purpureantennatus Morgan 1987 MG-H11
         |    |--P. runyanae W01
         |    `--P. syrtensis De Saint Laurent 1971 KK03
         `--Calcinus Dana 1851 PB06
              |--C. californiensis Bouvier 1898 PB06
              |--C. chilensis (Milne Edwards 1836) [=Pagurus chilensis] PB06
              |--C. elegans (Milne Edwards 1836) TSH09
              |--C. explorator Boone 1930 PB06
              |--C. gaimardii (Milne Edwards 1848) TSH09
              |--C. gaumensis Wooster 1984 TSH09
              |--C. herbstii De Man 1887 B55
              |--C. laevimanus (Randall 1840) PB06
              |--C. latens (Randall 1840) TSH09
              |--C. lineapropodus Morgan & Forest 1991 TSH09
              |--C. mclaughlinae Poupin & Bouchard 2006 PB06
              |--C. minutus Buitendijk 1937 TSH09
              |--C. obscurus Stimpson 1859 PB06
              |--C. pulcher Forest 1958 TSH09
              |--C. seurati Forest 1951 PB06
              |--C. talismani Milne-Edwards & Bouvier 1892 PB06
              |--C. tibicen (Herbst 1791) PB06
              |--C. tubularis (Linnaeus 1767) KK03
              `--C. vachoni Forest 1958 TSH09

*Type species of generic name indicated

REFERENCES

[B55] Bott, R. 1955. Dekapoden (Crustacea) aus El Salvador. 2. Litorale Dekapoden, außer Uca. Senckenbergiana Biologica 36: 45–70.

[DAS03] Dixon, C. J., S. T. Ahyong & F. R. Schram. 2003. A new hypothesis of decapod phylogeny. Crustaceana 76: 935–975.

[HS01] Hayward, B. W., A. B. Stephenson, M. S. Morley, W. M. Blom, H. R. Grenfell, F. J. Brook, J. L. Riley, F. Thompson & J. J. Hayward. 2001. Marine biota of Parengarenga Harbour, Northland, New Zealand. Records of the Auckland Museum 37: 45–80.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H86] Holthuis, L. B. 1986. Decapoda. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 589–615. E. J. Brill/Dr. W. Backhuys: Leiden.

[H14] Hosie, A. M. 2014. Review of the stalked barnacle genus Koleolepas (Cirripedia: Thoracica: Koleolepadidae), with new records from Australian waters. Records of the Western Australian Museum 29 (1): 1–9.

[KK03] Kocataş, A., & T. Katağan. 2003. The decapod crustacean fauna of the Turkish seas. Zoology in the Middle East 29: 63–74.

[L95] Lemaitre, R. 1995. A review of the hermit crabs of the genus Xylopagurus A. Milne Edwards, 1880 (Crustacea: Decapoda: Paguridae), including descriptions of two new species. Smithsonian Contributions to Zoology 570: 1–27.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O'Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[ML01] McLaughlin, P. A., & R. Lemaitre. 2001. A new family for a new genus and new species of hermit crab of the superfamily Paguroidea (Decapoda: Anomura) and its phylogenetic implications. Journal of Crustacean Biology 21 (4): 1062–1076.

[PB06] Poupin, J., & J. M. Bouchard. 2006. The eastern Pacific species of the genus Calcinus Dana, 1851, with description of a new species from Clipperton Island (Decapoda, Anomura, Diogenidae). Zoosystema 28 (2): 465–486.

[TSH09] Titelius, M. A., A. Sampey & C. G. Hass. 2009. Crustaceans of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, north Western Australia. Records of the Western Australian Museum Supplement 77: 145–176.

[W01] Williams, J. D. 2001. Polydora and related genera associated with hermit crabs from the Indo-West Pacific (Polychaeta: Spionidae), with descriptions of two new species and a second polydorid egg predator of hermit crabs. Pacific Science 55 (4): 429–465.

Homoloidea

Fossil of Homolopsis edwardsii, from here.

Belongs within: Brachyura.
Contains: Latheticocarcinus.

The Homoloidea are a group of podotreme crabs in which the uropod has been transformed into a pit (Guinot & Tavares 2001).

<==Homoloidea
    |  i. s.: Mithracites Gould 1859 GT01
    |           `--M. vectensis Gould 1859 GT01
    |         Lauridromia dehaani (Rathbun 1923) DAS03
    |--Rhinopoupinia Feldmann, Tshudy & Thomson 1993 [Poupiniidae] GT01
    |    `--R. bicornis Feldmann, Tshudy & Thomson 1993 GT01
    |--Latreilliidae GT01
    |    |--Eplumula australiensis (Henderson 1888) MG-H11
    |    `--Latreillia MG-H11
    |         |--L. elegans Roux 1830 DAS03
    |         `--L. pennifera Alcock 1900 MG-H11
    `--Homolidae SN04
         |--Homolax Alcock 1899 GT01
         |--Latheticocarcinus SN04
         |--Hypsophrys Wood-Mason in Wood-Mason & Alcock 1891 SN04
         |--Gastrodorus von Meyer 1864 SN04
         |--Laeviprosopon Glaessner 1933 SN04
         |--Tithonohomola Glaessner 1933 SN04
         |--Prohomola Karasawa 1992 SN04
         |--Mohola Guinot & Richer de Forges 1995 SN04
         |--Homologenus MG-H11
         |    |--H. braueri Doflein 1904 MG-H11
         |    `--H. malayensis Ihle 1912 MG-H11
         |--Yaldwynopsis MG-H11
         |--Moloha Guinot & Richer de Forges 1995 GT01
         |    `--M. majora (Kubo 1936) GT01
         |--Lignihomola Collins 1997 SN04
         |    `--‘Homolopsis’ etheridgei Woodward 1892 SN04
         |--Paromolopsis Wood-Mason in Wood-Mason & Alcock 1891 SN04
         |    |--*P. boasi Wood-Mason in Wood-Mason & Alcock 1891 SN04
         |    `--P. piersoni Schweitzer, Nyborg et al. 2004 SN04
         |--Hoplitocarcinus Beurlen 1928 SN04
         |    |--*H. johannesboehmi Beurlen 1928 SN04
         |    `--H. gibbosus (Schlüter 1879) [=Dromiopsis gibbosa] SN04
         |--Dagnaudus Guinot & Richer de Forges 1995 SN04
         |    |--D. petterdi (Grant 1905) MG-H11
         |    `--D. pritchardi (Jenkins 1977) [=Paromola pritchardi] SN04
         |--Zygastrocarcinus Bishop 1983 SN04
         |    |--Z. cardsmithi Bishop 1986 GT01
         |    `--Z. mendryki (Bishop 1982) [=Homolopsis mendryki] SN04
         |--Homola Leach 1815 SN04
         |    |--H. barbata (Fabricius 1793) [=Cancer barbatus; incl. *H. spinifrons Leach 1815] SN04
         |    |--H. orientalis Henderson 1888 MG-H11
         |    `--H. vancouverensis Schweitzer 2001 SN04
         |--Latreillopsis Henderson 1888 B64
         |    |--L. alcocki Stebbing 1920 B64
         |    |--L. bispinosa Henderson 1888 B64
         |    |--L. hawaiiensis Edmondson 1932 B64
         |    |--L. multispinosa Ihle 1912 B64
         |    |--L. petterdi Grant 1905 B64
         |    `--L. tetraspinosa Dai & Chen 1980 MG-H11
         `--Homolopsis Bell 1863 SN04
              |--*H. edwardsii Bell 1863 SN04
              |--H. chilensis Förster & Stinnesbeck 1987 SN04
              |--H. glabra Wright & Collins 1972 SN04
              |--H. hachiyai Takeda & Fujiyama 1983 SN04
              |--H. spinosa van Straelen 1936 SN04
              |--H. spinulosa Glaessner 1980 SN04
              |--H. tuberculata van Straelen 1936 SN04
              `--H. williamsi Bishop 1992 SN04

*Type species of generic name indicated

REFERENCES

[B64] Bennett, E. W. 1964. The marine fauna of New Zealand: Crustacea Brachyura. New Zealand Department of Scientific and Industrial Research Bulletin 153: 1–120.

[DAS03] Dixon, C. J., S. T. Ahyong & F. R. Schram. 2003. A new hypothesis of decapod phylogeny. Crustaceana 76: 935–975.

[GT01] Guinot, D., & M. Tavares. 2001. Une nouvelle famille de crabes du Crétacé, et la notion de Podotremata Guinot, 1977 (Crustacea, Decapoda, Brachyura). Zoosystema 23 (3): 507–546.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O'Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[SN04] Schweitzer, C. E., T. G. Nyborg, R. M. Feldmann & R. L. M. Ross. 2004. Homolidae de Haan, 1839 and Homolodromiidae Alcock, 1900 (Crustacea: Decapoda: Brachyura) from the Pacific Northwest of North America and a reassessment of their fossil records. Journal of Paleontology 78 (1): 133–149.

Leucosiidae

Ebalia tumefacta, copyright B. E. Picton.

Belongs within: Heterotremata.

The Leucosiidae, purse or pebble crabs, are a group of crabs with a rounded or pentagonal, usually very convex, carapace.

<==Leucosiidae
    |--Heteronucia venusta Nobili 1906 TSH09
    |--Myrine kessleri (Paul’son 1875) MG-H11
    |--Oreophorus reticulatus Adams & White 1849 MG-H11
    |--Parilia major Sakai 1962 MG-H11
    |--Philyra MG-H11
    |--Merocryptus Milne Edwards 1873 B64
    |    `--M. lambriformis Milne Edwards 1873 [incl. Ebalia rugulosa Yokoya 1933] B64
    |--Myra MG-H11
    |    |--M. curtimana Galil 2001 MG-H11
    |    |--M. fugax P71
    |    `--M. subgranulata (Fabricius 1798) KK03
    |--Randallia MG-H11
    |    |--R. eburnea Alcock 1896 MG-H11
    |    `--R. pustuloides Sakai 1961 MG-H11
    |--Persephona Leach 1817 BF07
    |    |--P. lichtensteinii Leach 1817 BF07
    |    |--P. mediterranea (Herbst 1794) BF07
    |    `--P. punctata (Linnaeus 1758) BF07
    |--Leucosia MG-H11
    |    |--L. haematosticta Adams & White 1848 MG-H11
    |    |--L. ocellata Bell 1855 MG-H11
    |    |--L. signata Paulson 1975 KK03
    |    `--L. whitei Bell 1855 MG-H11
    |--Arcania MG-H11
    |    |--A. cornuta (MacGilchrist 1905) MG-H11
    |    |--A. elongata Yokoya 1933 MG-H11
    |    |--A. gracilis (Henderson 1898) MG-H11
    |    |--A. muricata Galil 2002 MG-H11
    |    |--A. novemspinosa (Adams & White 1849) MG-H11
    |    `--A. septemspinosa (Fabricius 1787) MG-H11
    `--Ebalia Leach 1817 B64
         |--E. algirica Lucas 1846 E12
         |--E. cranchii Leach 1817 KK03
         |--E. deshayesii Lucas 1846 E12
         |--E. edwardsii Costa 1838 KK03
         |--E. granulosa Milne-Edwards 1837 KK03
         |--E. insignis Lucas 1846 E12
         |--E. laevis (Bell 1855) [=Phlyxia laevis; incl. P. cheesmani Filhol 1886, E. cheesmani] B64
         |--E. nux Milne-Edwards 1883 KK03
         |--E. tuberculosa (Milne Edwards 1873) [=Persephona tuberculosa; incl. Phlyxia granulosa Haswell 1880] B64
         |--E. tuberosa (Pennant 1777) KK03
         `--E. tumefacta M62

*Type species of generic name indicated

REFERENCES

[B64] Bennett, E. W. 1964. The marine fauna of New Zealand: Crustacea Brachyura. New Zealand Department of Scientific and Industrial Research Bulletin 153: 1–120.

[BF07] Braga, A. A., A. Fransozo, G. Bertini & P. B. Fumis. 2007. Bathymetric distribution and recruitment of the spider crab Libinia spinosa H. Milne Edwards 1834 in the Ubatuba and Caraguatatuba regions, northern coast of São Paulo, Brazil (Crustacea, Brachyura, Majoidea, Pisidae). Senckenbergiana Biologica 87 (1): 7–16.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[KK03] Kocataş, A., & T. Katağan. 2003. The decapod crustacean fauna of the Turkish seas. Zoology in the Middle East 29: 63–74.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O'Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[M62] Monniot, F. 1962. Recherches sur les graviers a Amphioxus de la région de Banyuls-sur-Mer. Vie et Milieu 13: 231–322.

[P71] Por, F. D. 1971. One hundred years of Suez Canal—a century of Lessepsian migration: retrospect and viewpoints. Systematic Zoology 20 (2): 138–159.

[TSH09] Titelius, M. A., A. Sampey & C. G. Hass. 2009. Crustaceans of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, north Western Australia. Records of the Western Australian Museum Supplement 77: 145–176.

Inachidae

Cyrtomaia, from the Smithsonian Institution.

Belongs within: Heterotremata.

The Inachidae are a group of spider crabs in which the often long eyestalks lack orbits.

Characters (from Bennett 1964, as Inachinae): Eyes without orbits; eyestalks generally long, either non-retractile or retractile against sides of carapace or against acute postocular spine; basal article of antenna extremely slender throughout its length, usually long.

<==Inachidae [Inachinae]
    |--Camposcioidea B64
    |    |--Camposcia retusa Latreille 1829 MG-H11
    |    |--Stenorhynchus fissifrons Haswell 1879 B64
    |    `--Trichoplatus Milne Edwards 1876 B64
    |         `--*T. huttoni Milne Edwards 1876 (see below for synonymy) B64
    `--Macrocheiroidea [Macrocheiridae] B64
         |--Macrocheira kaempferi MD01
         `--Cyrtomaia Miers 1886 [incl. Echinomaia Borradaile 1916] B64
              |--C. balssi iIhle & Ihle-Landenberg 1931 B64
              |--C. bicornis Ihle & Ihle-Landenberg 1931 B64
              |--C. echinata Rathbun 1916 B64
              |--C. goodridgei McArdle 1900 B64
              |--C. hispida (Borradaile 1916) [=Echinomaia hispida; incl. C. platypes Yokoya 1933] B64
              |--C. horrida Rathbun 1916 (see below for synonymy) B64
              |--C. lamellata Rathbun 1906 B64
              |--C. maccullochi Rathbun 1918 MG-H11
              |--C. murrayi Miers 1886 MG-H11
              |--C. septemspinosa Rathbun 1932 B64
              |--C. smithi Rathbun 1893 [incl. C. smithi var. tenuipedunculata Ihle & Ihle-Landenberg 1931] B64
              `--C. suhmi Miers 1886 B64

Inachidae incertae sedis:
  Inachus TF08
    |--I. aguiarii De Brito Capello 1876 KK03
    |--I. communissimus Rizza 1839 KK03
    |--I. dorsettensis (Pennant 1777) TF08
    |--I. leptochirus Leach 1817 KK03
    |--I. mauritanicus Lucas 1846 E12
    `--I. thoracicus Roux 1830 KK03
  Dorhynchus MG-H11
    |--D. ramusculus (Baker 1906) MG-H11
    `--D. thomsoni (Thomson 1873) KK03
  Dumea latipes (Haswell 1880) MG-H11
  Ephippias endeavouri Rathbun 1918 MG-H11
  Grypachaeus hyalinus (Alcock & Anderson 1894) MG-H11
  Physachaeus ctenurus Alcock 1895 MG-H11
  Pleistacantha maxima Ahyong & Lee 2006 MG-H11
  Sunipea indicus (Alcock 1895) MG-H11
  Platymaia MG-H11
    |--P. fimbriata Rathbun 1916 MG-H11
    `--P. wyvillethomsoni Miers 1886 MG-H11
  Achaeus Leach 1817 B64
    |--A. brevirostris (Haswell 1879) MG-H11
    |--A. cranchii Leach 1817 KK03
    |--A. curvirostris (Milne-Edwards 1873) MG-H11
    |--A. gracilis Costa 1839 KK03
    `--A. lacertosus Stimpson 1858 MG-H11
  Oncinopus MG-H11
    |--O. angustifrons Takeda & Miyake 1969 MG-H11
    |--O. aranea De Haan 1839 MG-H11
    `--O. neptunus Adams & White 1848 MG-H11
  Achaeopsis Stimpson 1857 B64
  Macropodia Leach 1814 B64
    |--M. czernjawskii (Brandt 1880) KK03
    |--M. linaresi Forest & Zariquiey-Alvarez 1964 KK03
    |--M. longipes (Milne-Edwards & Bouvier 1899) KK03
    |--M. rostrata (Linnaeus 1761) KK03
    `--M. tenuirostris (Leach 1814) KK03

Cyrtomaia horrida Rathbun 1916 [incl. C. horrida var. japonica Balss 1924, C. horrida var. pilosa Ihle & Ihle-Landenberg 1931] B64

*Trichoplatus huttoni Milne Edwards 1876 [=Naxia huttoni; incl. Halimus hectori Miers 1876, H. rubiginosus Kirk 1881, Naxia (Halimus) rubiginosus] B64

*Type species of generic name indicated

REFERENCES

[B64] Bennett, E. W. 1964. The marine fauna of New Zealand: Crustacea Brachyura. New Zealand Department of Scientific and Industrial Research Bulletin 153: 1–120.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[KK03] Kocataş, A., & T. Katağan. 2003. The decapod crustacean fauna of the Turkish seas. Zoology in the Middle East 29: 63–74.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1–124.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O'Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[TF08] Teixeira, G. M., V. Fransozo, A. L. Castilho, R. C. da Costa & F. A. d. M. Freire. 2008. Size distribution and sex ration in the spider crab Epialtus brasiliensis (Dana 1852) associated with seaweed on a rocky shore in southeastern Brazil (Crustacea, Decapoda, Brachyura, Majoidea, Epialtidae). Senckenbergiana Biologica 88 (2): 169–175.

Majidae

Seaweed crab Paramithrax peroni, from McLintock 1966.

Belongs within: Heterotremata.
Contains: Leptomithrax.

The Majidae, spider crabs, are a group of crabs with highly mobile chelipeds that may commonly be used to attach algae and debris to themselves for camouflage.

Characters (from Bennett 1964): Chelipeds especially mobile, rarely much greater than other legs or with fingers bent at angle on hand. Second joint of antenna well developed, generally fused with epistome and often with front. Orbits generally more or less incomplete. Hooked hairs almost always present. Male apertures coxal. Palp of external maxillipeds articulated wither at summit or at antero-internal angle of merus.

<==Majidae [Periceridae, Periceroida]
    |--Ophthalmiinae [Stenocionopoida] B64
    |--Macrocoelominae B64
    |--Micippoides [Hyasteniinae] B64
    |    `--M. angustifrons B64
    |--Acanthonychinae B64
    |    |--Simocarcinus B64
    |    `--Acanthonyx TF08
    |         |--A. lunulatus (Risso 1816) KK03
    |         `--A. scutiformis (Dana 1851) TF08
    |--Mithracinae [Mithracidae] TSH09
    |    |--Micippa TSH09
    |    |    |--M. cristata (Linnaeus 1758) TSH09
    |    |    `--M. thalia (Herbst 1803) KK03
    |    |--Paramicippa spinosa B64
    |    |--Jacquinotia Rathbun 1915 [incl. Prionorhynchus Jacq. in Jacq. & Lucas 1853 (preoc.)] B64
    |    |    `--*J. edwardsi (Jacquinot 1853) [=Prionorhynchus edwardsi; incl. Paramicippa grandis Hector 1899] B64
    |    |--Tiarinia TSH09
    |    |    |--T. angusta Dana 1852 TSH09
    |    |    `--T. cornigera (Latreille 1825) TSH09
    |    `--Mithrax Latreille 1817 GT01
    |         `--M. (Mithraculus) B55
    |              |--M. (M.) forceps (Milne-Edwards 1875) [=Mithraculus forceps] B55
    |              `--M. (M.) sculptus (Lamarck 1818) [=Maia sculpta] B55
    `--Majinae [Mioida, Mamaiidae] TSH09
         |--Cyclax suborbicularis (Stimpson 1858) TSH09
         |--Schizophrys aspera (Milne Edwards 1834) TSH09
         |--Pseudomicippe TSH09
         |--Maiopsis B64
         |--Campbellia Balss 1930 B64
         |    `--*C. kohli Balss 1930 B64
         |--Gonatorhynchus Haswell 1880 [incl. Lobophrys Filhol 1886] B64
         |    |--*G. tumidus B64
         |    `--G. barbicornis (Latreille 1828) (see below for synonymy) B64
         |--Entomonyx Miers 1884 MG-H11, B64
         |    |--E. depressus Sakai 1974 MG-H11
         |    `--E. spinosus Miers 1884 MG-H11 [=Acanthophrys spinosus B64; incl. Macrocoeloma nummifer Alcock 1895 B64]
         |--Maja MG-H11
         |    |--M. confragosa Griffin & Tranter 1986 MG-H11
         |    |--M. crispata Risso 1827 KK03
         |    |--M. gibba Alcock 1895 MG-H11
         |    |--M. goltziana D’Oliveira 1888 KK03
         |    |--M. squinado (Herbst 1788) DAS03
         |    `--M. suluensis Rathbun 1916 MG-H11
         |--Acanthophrys Milne Edwards 1865 [incl. Chlorinoides Haswell 1879] B64
         |    |--*A. cristimanus B64
         |    |--A. filholi Milne Edwards 1865 B64
         |    |--A. goldsboroughi Rathbun 1906 B64
         |    |--A. spatulifer (Haswell 1882) [=Paramithrax spatulifer, Chlorinoides spatulifer] B64
         |    `--A. tenuirostris [=*Chlorinoides tenuirostris] B64
         |--Paramithrax Milne Edwards 1834 B64
         |    |--*P. peroni Milne Edwards 1834 [=P. peronii] B64
         |    |--P. backstromi Balss 1923 B64
         |    |--P. minor Filhol 1886 [incl. P. parvus Borradaile 1916] B64
         |    |--P. parvispinosus Ward 1933 B64
         |    |--P. pipa (Herbst 1788) [=Cancer pipa, Maja pipa] B64
         |    |--P. spinosus Miers 1879 B64
         |    `--P. ursus (Herbst 1788) (see below for synonymy) B64
         `--Leptomithrax B64

Majidae incertae sedis:
  Hyas Leach 1814 GT01
    |--H. araneus Linnaeus 1758 DK89
    `--H. coarctatus Leach 1815 TF08
  Microphrys Milne-Edwards 1851 B55
    `--M. bicornutus (Latreille 1825) [=Pisa bicornuta] B55
  Planotergum mirabile Balss 1935 MG-H11
  Prismatopus MG-H11
    |--P. brevispinosus (Yokoya 1933) MG-H11
    `--P. occidentalis (Griffin 1970) MG-H11
  Naxia tumida B64
  Telophrys cristulipes B64

Gonatorhynchus barbicornis (Latreille 1828) [=Pisa barbicornis, *Lobophrys barbicornis, Paramithrax barbicornis] B64

Paramithrax ursus (Herbst 1788) [=Cancer (Mithrax) ursus, Inachus ursus, Maja ursus; incl. P. cristatus Filhol 1886, P. latreillei Miers 1876] B64

*Type species of generic name indicated

REFERENCES

[B64] Bennett, E. W. 1964. The marine fauna of New Zealand: Crustacea Brachyura. New Zealand Department of Scientific and Industrial Research Bulletin 153: 1–120.

[B55] Bott, R. 1955. Dekapoden (Crustacea) aus El Salvador. 2. Litorale Dekapoden, außer Uca. Senckenbergiana Biologica 36: 45–70.

[DK89] Deaton, L. E., & K. D. Kempler. 1989. Occurrence of the ribbed mussel, Geukensia demissa, on the book gills of a horseshoe crab, Limulus polyphemus. Nautilus 103 (1): 42.

[GT01] Guinot, D., & M. Tavares. 2001. Une nouvelle famille de crabes du Crétacé, et la notion de Podotremata Guinot, 1977 (Crustacea, Decapoda, Brachyura). Zoosystema 23 (3): 507–546.

[KK03] Kocataş, A., & T. Katağan. 2003. The decapod crustacean fauna of the Turkish seas. Zoology in the Middle East 29: 63–74.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O'Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[TF08] Teixeira, G. M., V. Fransozo, A. L. Castilho, R. C. da Costa & F. A. d. M. Freire. 2008. Size distribution and sex ration in the spider crab Epialtus brasiliensis (Dana 1852) associated with seaweed on a rocky shore in southeastern Brazil (Crustacea, Decapoda, Brachyura, Majoidea, Epialtidae). Senckenbergiana Biologica 88 (2): 169–175.

[TSH09] Titelius, M. A., A. Sampey & C. G. Hass. 2009. Crustaceans of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, north Western Australia. Records of the Western Australian Museum Supplement 77: 145–176.