Systasis

Female Systasis, from Bouček (1988).

Belongs within: Ormocerinae.

Systasis is a cosmopolitan genus of wasps of uncertain biology: they may parasitise insect larvae within galls, or they may be gall-formers themselves (Bouček 1988).

Characters (from Bouček 1988): Body with more or less distinct metallic tinge; antenna with seven segments between pedicel and clava; thoracic dorsum usually lacking coarse punctures, usually with dense reticulation (and then dull) or very dense but fine puncturation; propodeum deeply reticulate; forewing with rather sparse pilosity, with a large speculum anteriorly reaching virtually the distal end of the marginal vein, thus making conspicuous a row of longish hairs on the underside, below the marginal vein; marginal vein more than twice as long as stigmal, postmarginal vein usually only slightly longer than the stigmal; legs usually more or less infuscate.

<==Systasis Walker 1834 [incl. Guieralia Risbec 1951, Paruriella Girault 1913, Systasina Bouček 1955] B88
    |--*S. encyrtoides B88
    |--S. australiensis (Girault 1913) [=Paruriella australiensis] B88
    |--‘Miscogaster’ bato Walker 1839 B88
    |--S. cecidomyiae (Ashmead 1900) [=Asemantus cecidomyiae] B88
    |--S. cecili Girault 1929 B88
    |--S. cernus (Walker 1839) [=Miscogaster cernus, Seladerma cernum] B88
    |--S. dasyneurae A71
    |--S. dice (Walker 1839) [=Miscogaster dice, Semiotellus dice, Semiotus dice] B88
    |--S. doddi (Girault 1915) [=Schizonotus doddi] B88
    |--S. euctemon (Walker 1839) [=Pteromalus euctemon] B88
    |--S. flindersiae Girault 1934 G34
    |--S. graminis (Cameron 1912) (see below for synonymy) B88
    |--S. grotiusi Girault 1915 B88
    |--S. henrici Girault 1913 B88
    |--S. horridula (Girault 1915) [=Paruriella horridula] B88
    |--S. insularis Dodd & Girault in Girault 1915 B88
    |--S. keatsi Girault 1927 B88
    |--S. lelex (Walker 1839) [=Pteromalus lelex] B88
    |--‘Miscogaster’ letus Walker 1839 B88
    |--S. merula (Walker 1839) [=Miscogaster merula, Semiotellus merula, Semiotus merula] B88
    |--S. persimilis (Dodd & Girault in Girault 1915) [=Paruriella persimilis] B88
    |--S. punctativertex Girault 1915 B88
    |--S. seposita Girault 1915 [=S. sepositus] B88
    `--S. varipes Girault 1915 B88

Systasis graminis (Cameron 1912) [=Callimome graminis, Paruriella graminis; incl. C. reticulatus Cameron 1912, Systasis reticulata] B88

*Type species of generic name indicated

REFERENCES

Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

Diparinae

Lelaps striatus, from here.

Belongs within: Chalcidoidea.

The Diparinae are a group of chalcidoid wasps found primarily in leaf litter. Many species are strongly sexually dimorphic: males are usually macropterous, but females may be brachypterous or apterous (Desjardins 2007).

Characters (from Bouček 1988, Desjardins 2007): Antenna 13-segmented. Scutellum varied, but never smooth and convex. Metacoxa inserted unusually high, usually with transverse striations on posterior margin. Gastral tergite 1 usually expanded to cover at least half of metasoma. Cercal brush present. Females often apterous or brachypterous.

<==Diparinae [Diparides, Diparini, Lelapinae]
    |--+--Chimaerolelaps Desjardins 2007 D07
    |  |    `--*C. villosa Desjardins 2007 D07
    |  `--Dipara D07
    `--+--Netomocera Bouček 1954 D07, B88 [Netomocerini]
       |    |--*N. setifera Bouček 1954 D07
       |    |--N. africana Hedqvist 1971 D07
       |    |--N. alboscapus Hedqvist 1971 D07
       |    |--N. nearctica Yoshimoto 1977 D07
       |    |--N. nigra Sureshan & Narendran 1990 D07
       |    |--N. rufa Hedqvist 1971 D07
       |    `--N. sedlaceki Bouček 1988 B88
       `--+--Lelaps Walker 1843 (see below for synonymy) D07
          |    |--*L. pulchricornis [=*Dilaelaps pulchricornis, *Stenopistha pulchricornis] D07
          |    |--L. abdominalis Ashmead 1904 D07
          |    |--L. aeneiceps Ashmead 1904 D07
          |    |--L. affinis Ashmead 1904 D07
          |    |--L. albipes Cameron 1884 D07
          |    |--L. albofasciatus Hedqvist 1964 D07
          |    |--L. annulicornua (Strand 1911) [=Stenopistha annulicornua] D07
          |    |--L. apicalis Ashmead 1904 D07
          |    |--L. argenticoxa Girault 1916) [=*Spalangiolaelaps argenticoxa] D07
          |    |--L. avicula Haliday 1844 D07
          |    |--L. beckeri Yoshimoto 1977 D07
          |    |--L. bimaculata Ashmead 1904 D07
          |    |--L. callisto Marshall in Buysson & Marshall 1892 [=Laelaps (l. c.) callisto] D07
          |    |--L. caudatula (Strand 1911) [=Stenopistha caudatula] D07
          |    |--L. decorata Walker 1862 [=Laelaps (l. c.) decorata] D07
          |    |--L. ferrierei Hedqvist 1964 D07
          |    |--L. ferruginea Cameron 1884 D07
          |    |--L. flagellata (Strand 1911) [=Stenopistha flagellata] D07
          |    |--L. flavescens Ashmead in Riley et al. 1894 D07
          |    |--L. floridensis Yoshimoto 1977 D07
          |    |--L. halidayi Ashmead 1904 [incl. Stenopistha halidayi sobrina Strand 1911] D07
          |    |--L. insularis Mercet 1927 D07
          |    |--L. magnifica (Strand 1911) [=Stenopistha magnifica] D07
          |    |--L. melinus Yoshimoto 1977 D07
          |    |--L. noortii Desjardins 2007 D07
          |    |--L. ornata (Strand 1911) [=Stenopistha ornata] D07
          |    |--L. paraguayensis Girault 1912 D07
          |    |--L. picta Walker 1862 [=Laelaps (l. c.) picta] D07
          |    |--L. pulchella (Strand 1911) [=Stenopistha pulchella] D07
          |    |--L. pulchra Girault 1912 D07
          |    |--L. pulchricornis Walker 1843 D07
          |    |--L. pygata (Strand 1911) [=Stenopistha pygata; incl. S. pygata conjuncta Strand 1911] D07
          |    |--L. rectivitta (Strand 1911) [=Stenopistha rectivitta] D07
          |    |--L. rhomboidea (Strand 1911) [=Stenopistha rhomboidea] D07
          |    |--L. sadales (Walker 1839) [=Merostenus sadales] D07
          |    |--L. setifrons (Strand 1911) [=Stenopistha setifrons] D07
          |    |--L. simplex (Fabricius 1804) [=Chalcis simplex] D07
          |    |--L. striaticeps (Strand 1911) [=Stenopistha striaticeps] D07
          |    |--L. striatus Yoshimoto 1977 D07
          |    |--L. stylata Ashmead 1904 D07
          |    |--L. tapajoana (Schulz 1906) [=Dilaelaps tapajoana, L. ferruginea Ashmead 1904 non Cameron 1884] D07
          |    |--L. terebrans (Strand 1911) [=Stenopistha terebrans] D07
          |    |--L. viridiceps (Strand 1911) [=Stenopistha viridiceps] D07
          |    `--L. vittipennis (Strand 1911) [=Stenopistha vittipennis] D07
          `--+--Dozodipara Desjardins 2007 D07
             |    `--*D. insularis Desjardins 2007 D07
             `--+--Boeria Hedqvist 1969 D07
                |    `--*B. saetosa Hedqvist 1969 D07
                `--+--Cerodipara Desjardins 2007 D07
                   |    `--*C. sabensis Desjardins 2007 D07
                   `--+--+--Conodipara Hedqvist 1972 [=Turneria Hedqvist 1969 non Forel 1895] D07
                      |  |    `--*C. scutellata (Hedqvist 1969) [=*Turneria scutellata] D07
                      |  `--+--Conophorisca Hedqvist 1969 D07
                      |     |    |--*C. annulata Hedqvist 1969 D07
                      |     |    |--C. grisselli Desjardins 2007 D07
                      |     |    `--C. littoriticus Desjardins 2007 D07
                      |     `--Myrmicolelaps Hedqvist 1969 [incl. Dolichodipara Hedqvist 1969] D07
                      |          |--+--*M. paradoxus Hedqvist 1969 D07
                      |          |  `--M. aurantius Desjardins 2007 D07
                      |          `--+--M. iridius Desjardins 2007 D07
                      |             `--M. scutellata (Hedqvist 1969) [=*Dolichodipara scutellata] D07
                      `--+--Pyramidophoriella Hedqvist 1969 D07
                         |    |--*P. albiclava Hedqvist 1969 D07
                         |    `--P. brunnea Hedqvist 1969 D07
                         `--+--Nosodipara Bouček 1988 D07
                            |    |--*N. monteithorum Bouček 1988 D07
                            |    `--N. ferrana Desjardins 2007 D07
                            `--Pseudoceraphron Dodd 1924 [incl. Dipareta Bouček 1988, Malinka Bouček 1988] D07
                                 |--P. regieri Desjardins 2007 D07
                                 `--+--P. albifrons (Bouček 1988) [=*Dipareta albifrons] D07
                                    `--+--+--*P. pulex Dodd 1924 D07
                                       |  `--P. burwelli Desjardins 2007 D07
                                       `--+--P. fijiensis Desjardins 2007 D07
                                          `--P. nana (Bouček 1988) [=*Malinka nana] D07

Lelaps Walker 1843 [=Laelaps (l. c.), Dilaelaps Schulz 1906; incl. Spalangiolaelaps Girault 1916, Stenopistha Strand 1910] D07

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[D07] Desjardins, C. A. 2007. Phylogenetics and classification of the world genera of Diparinae (Hymenoptera: Pteromalidae). Zootaxa 1647: 1-88.

Terobiella

Antenna of male Terobiella nigriceps, from Bouček (1988).

Belongs within: Ormocerinae.

Terobiella is an Australian genus of wasps causing galls on leaves of Eucalyptus (Bouček 1988).

Characters (from Bouček 1988): Body compact, non-metallic. Antennae 12-segmented, with seven segments between pedicel and clava, bearing long branches in the males, females with most funicular segments dorsally only slightly produced appearing serrate. Scutellum anteriorly pointed, axillae converging medially.

<==Terobiella Ashmead 1900 (see below for synonymy) B88
    |--*T. flavifrons Ashmead 1900 [incl. *Melanosomella flavipes Girault 1913] B88
    |--T. adolphi (Girault 1931) [=Perilampoides adolphi] B88
    |--T. bicolor (Girault 1913) [=*Perilampoides bicolor] B88
    |--T. cinctitibiae (Girault 1931) [=Perilampoides cinctitibiae] B88
    |--T. dilutiventris (Girault 1913) [=Epiperilampus dilutiventris] B88
    |--T. eucalypti (Gallard 1930) [=Tepperella eucalypti] B88
    |--T. nigriceps (Ashmead 1904) (see below for synonymy) B88
    |--T. particolor (Girault 1915) [=Perilampoides particolor] B88
    |--T. regalis (Girault 1931) [=Perilampoides regalis] B88
    |--T. scutata (Girault 1931) [=Perilampoides scutatus] B88
    |--T. similis (Girault 1917) [=Perilampoides similis] B88
    `--T. tennysoni (Girault 1920) [=Perilampoides tennysoni] B88

Terobiella Ashmead 1900 [incl. Melanosomella Girault 1913, Paraterobia Ashmead 1904, Perilampoides Girault 1913, Tepperella Cameron 1912] B88

Terobiella nigriceps (Ashmead 1904) [=*Paraterobia nigriceps; incl. Perilampoides aurantiscutum anna Girault 1931, Pe. aurantiscutum Girault 1916, Pe. flavithorax Girault 1915, *Tepperella maculiscutis Cameron 1912, Pe. aurantiscutum regis Girault 1931, Pe. aurantiscutum rex Girault 1931] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

Trichilogaster

Female Trichilogaster signiventris, photographed by John Hoffman.

Belongs within: Ormocerinae.

Trichilogaster is a genus of wasps forming galls on Acacia species (Bouček 1988).

Characters (from Bouček 1988): Body robust, often partly rufous or yellowish brown, with thorax short and convex, sometimes broader than head, always sculptured, sometimes rather coarsely. Head with broadly concave occiput and convex frons; scrobes not large but sometimes reaching ocellus. Clypeus produced, bilobate. Antennae at centre of face or slightly above, formula 11263; anelli large, flagellum in female subfiliform. clava symmetric. Pronotum nearly as broad as scutum, medially short; notauli narrow but deep; axillae slightly advanced, medially almost meeting. Scutellum slightly oblong, with admarginai groove, apex closely applied to raised middle of dorsellum (in side view). Propodeum often with longitudinal rugae or at least with weak median carina and curved plicae posteriorly. Hind tibia with 2 spurs. Forewing with veins rather short, the marginal and base of postmarginal slightly thickened, stigma often knobbed; often infumation present below parastigma. Gaster short and high but collapsing rather irregularly; hind margins of tergites 1 and 2 emarginate, exposing large third tergite, this often with coarse sculpture.

<==Trichilogaster Mayr 1906 [incl. Epiperilampus Girault 1913] B88
    |--*T. maideni (Froggatt 1892) [=Cynips maideni] B88
    |--T. acaciaelongifoliae (Froggatt 1892) [=Cynips acaciaelongifoliae, T. longifoliae Mayr 1905] B88
    |--T. arabica B88
    |--T. atra (Girault 1915) [=Epiperilampus ater] B88
    |--T. atricorpus (Girault 1931) [=Epiperilampus atricorpus; incl. E. atricorpus socratis Girault 1931] B88
    |--T. channingi (Girault 1913) [=Epiperilampus channingi] B88
    |--T. esculenta Ferrière in Gahan & Ferrière 1947 [=T. esculentus] B88
    |--T. flavivena (Girault 1931) [=Coelocybelloides flavivena] B88
    |--T. pendulae Mayr 1905 B88
    |--T. significata (Girault 1913) [=Epiperilampus significatus] B88
    |--T. signiventris (Girault 1931) (see below for synonymy) B88
    `--T. xanthocephala (Girault 1913) [=*Epiperilampus xanthocephalus] B88

Trichilogaster signiventris (Girault 1931) [=Perilampella signiventris; incl. Epiperilampus adolphi Girault 1931, Trichilogaster adolphi, Coelocybelloides atraticorpus Girault 1931] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

Encyrtocephalus

Female Encyrtocephalus albiclava, from Bouček (1988).

Belongs within: Ormocerinae.

Encyrtocephalus is a genus of wasps found in Africa and Australia that cause galls in trees such as Acacia, Eucalyptus and Brachychiton (Bouček 1988).

Characters (from Bouček 1988): Antenna with eight segments between pedicel and clava, first two segments of flagellum subequal in size, shorter than following ones; upper margin of occiput sometimes carinate or edged, rarely rounded; body dull, without metallic gloss; thorax with conspicuous pilosity and coarse puncturation (latter sometimes less conspicuous in small specimens); prepectus large and broadly triangular; forewing often without infumation, sometimes infumate just at stigma, sometimes with double or almost treble infumation; fringe well developed; stigmal vein strongly arched, at its perpendicular base more or less triangularly thickened, its base and apex usually darkened, distinctly shorter than marginal vein; gaster usually sessile, very rarely on conspicuous petiole.

<==Encyrtocephalus Ashmead 1900 (see below for synonymy) B88
    |--*E. simplicipes Ashmead 1900 B88
    |--E. albiclava (Girault 1915) [=Neorileyella (*Pseudorileya) albiclava] B88
    |--E. albipilum Girault 1931 B88
    |--E. atrativentris Girault 1931 B88
    |--E. fasciatus (Girault 1915) [=*Neorileyella fasciata] B88
    |--E. gallicola (Ashmead 1904) [=*Decatomothorax gallicola] B88
    |--E. hyalinus (Girault 1915) [=Neorileyella hyalina] B88
    |--E. io Girault 1931 B88
    |--E. katrina (Girault 1931) [=Neorileyella katrina] B88
    |--E. mozarti Girault 1931 B88
    |--E. notabilis [=*Eucyrtonotus notabilis] B88
    `--E. ophelia (Girault 1931) [=Neorileyella ophelia] B88

Encyrtocephalus Ashmead 1900 [incl. Decatomothorax Ashmead 1904, Encyrtonotus (l. c.), Eucyrtonotus Silvestri 1915, Neorileyella Girault 1915, Pseudorileya Girault 1915] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

Colotrechninae

Ovipositing Cameronella, from here.

Belongs within: Chalcidoidea.

The Colotrechninae are a group of chalcidoid wasps found in Australia and South America. Host data are known for only a small number of species, but colotrechnines include species associated with plant galls and seed pods (Bouček 1988).

Characters (from Bouček 1988): Occipital carina absent; large axillulae present, separated from scutellum by (mostly) outcurving grooves or lines visible dorsally.

<==Colotrechninae [Colotrechnides] B88
    |--Uzka Bouček 1988 [Uzkini] B88
    |    `--*U. bethyloides Bouček 1988 B88
    |--Divna Bouček 1988 [Divnini] B88
    |    `--*D. hirtura Bouček 1988 B88
    |--Amerostenini B88
    |    |--Errolia Bouček 1988 B88
    |    |    `--*E. cyanea Bouček 1988 B88
    |    |--Glorimontana Bouček 1988 B88
    |    |    `--*G. flaveola Bouček 1988 B88
    |    |--Omphalodipara Girault 1923 [incl. Froudeana Girault 1928] B88
    |    |    |--*O. splendida Girault 1923 B88
    |    |    `--O. clavicornis (Dodd 1924) [=Hetreulophus clavicornis, *Froudeana clavicornis] B88
    |    `--Amerostenus Girault 1913 B88
    |         |--*A. australiensis Girault 1913 B88
    |         |--A. aereipes Girault 1915 B88
    |         |--‘Miscogaster’ athanis Walker 1839 B88
    |         |--A. theope (Walker 1839) [=Miscogaster theope, Semiotellus theope, Semiotus theope] B88
    |         |--A. tridentatus Girault 1928 B88
    |         `--A. varidentatus Girault 1929 B88
    `--Colotrechnini [Cremneulophini, Dvaliniinae] B88
         |--Bofuria Hedqvist 1978 B88
         |--Bomburia Hedqvist 1978 B88
         |--Dvalinia Hedqvist 1977 B88
         |--Elachertodomyia Girault 1917 B88
         |--Pachyneuronella Girault 1913 B88
         |    `--*P. viridis Girault 1913 B88
         |--Uriellopteromalus Girault 1915 B88
         |    |--*U. subplanithorax Girault 1915 B88
         |    `--U. aeneus Bouček 1988 B88
         |--Colotrechnus Thomson 1878 [incl. Zanonia Masi 1921] B88
         |    |--*C. subcoeruleus B88
         |    |--C. agromyzae B88
         |    |--C. ignotus B88
         |    |--C. notalauris Bouček 1988 B88
         |    `--C. viridis B88
         `--Cameronella Dalla Torre 1898 (see below for synonymy) B88
              |--*C. blackburni (Cameron 1888) [=*Panthalis blackburni] B88
              |--C. apiomorphae (Girault 1939) [=Dinoura apiomorphae] B88
              |--C. auriventris (Ashmead 1900) [=*Dinoura auriventris] B88
              |--C. cyanea (Ashmead 1900) [=Dinoura cyanea] B88
              |--C. eucalypti (Girault 1917) [=Dinoura eucalypti] B88
              |--C. gigantea (Girault 1916) [=*Cremneulophus giganteus] B88
              `--C. pulchra (Girault 1927) [=Dinoura pulchra] B88

Cameronella Dalla Torre 1898 [=Panthalis Cameron 1888 non Kinberg 1854;incl. Cremneulophus Girault 1916, Dinoura Ashmead 1900] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

Cleonymus

Unidentified Cleonymus, photographed by Kim Fleming.

Belongs within: Cleonyminae.

Cleonymus is a near-cosmopolitan genus of wasps (unrecorded from South America), parasitic on wood-boring beetles (Bouček 1988).

Characters (from Bouček 1988): Antenna with eight distinct segments between pedicel and clava; male antenna unbranched, with almost symmetrical segments covered by regular dense pilosity; preclava in female with distinct lateral process closely applied to side of clava; notaular grooves becoming indistinct in very dense reticulation-puncturation on posterior half of scutum; stigmal vein in both sexes fairly long, at very low angle, fully twice as long as distance of stigmal knob from postmarginal vein; hind femur often serrate or with a subapical tooth on ventral margin; gaster in female sessile.

<==Cleonymus Latreille 1809 (see below for synonymy) B88
    |--*C. depressus [=Diplolepis depressa] B88
    |--C. californicus Girault 1917 GM79
    |--C. dentatifemur (Girault 1926) [=Aplatygerrhus dentatifemur] B88
    |--C. imperialis (Dodd 1924) [=Aplatygerrhus imperialis] B88
    |--C. laticinctus (Girault 1926) [=Systolomorphella laticincta] B88
    |--C. magnificus (Ashmead 1888) [=Charitopus magnificus, *Ptinobius magnificus] B88
    |--C. magnus Bouček 1988 [=*Aplatygerrhus magnificus Girault 1913 non Charitopus magnificus Ashmead 1888] B88
    |--C. nigriclavus Girault 1917 G17
    |--C. pentlandi (Girault 1922) [=Systolomorphella pentlandi] B88
    |--C. regalis (Dodd 1924) [=Aplatygerrhus regalis] B88
    |--C. rufiscapus (Girault 1925) [=Aplatygerrhus rufiscapus] B88
    |--C. silvifilia (Girault 1927) [=Systolomorphella silvifilia] B88
    |--C. trifasciatipennis (Girault 1915) [=*Systolomorphella trifasciatipennis] B88
    |--C. unfasciatipennis (Dodd & Girault in Girault 1915) [=Systolomorphella unfasciatipennis] B88
    |--C. unnotipennis (Girault 1915) [=Systolomorphella unnotipennis] B88
    `--‘*Beharella’ viridicyanea B88

Cleonymus Latreille 1809 [incl. Aplatygerrhus Girault 1913, Beharella Risbec 1952, Ptinobius Ashmead 1896, Systolomorphella Girault 1915] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[G17] Girault, A. A. 1917. Descriptiones hymenopterorum chalcidoidicarum variorum cum observationibus. III. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 102-111).

[GM79] Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 1-400.