Notaphus varius, copyright Udo Schmidt.

Belongs within: Conjunctae.
Contains: Elaphropus, Pericompsus, Tachys, Philipis, Bembidion, Peryphus.

The Bembidiini are a group of carabid beetles with reduced terminal segments on the labial and maxillary palps, and two unequally dilated segments in the tarsi of males. The majority of bembidiins are surface-dwelling hygrophiles though some are xerophilous and others are endogeous (Ball & Bousquet 2001). Members of the genus Micratopus have truncate elytra and a single pair of supraorbital setae whereas other Bembidiini possess two pairs of supraorbital setae and the elytra are not truncate (Bell 1990). Members of the subtribe Tachyina are characterised by an apicolateral notch on the protibiae, and usually an apical recurrent groove on the elytra.

<==Bembidiini [Bembidiinae]
    |--Gouleta [Xystosomina] R-JC01
    |    `--G. cayennensis R-JC01
    |--Tachyina R-JC01
    |    |--Pericompsus R-JC01
    |    |--Tachys BB01
    |    `--Philipis B02
    `--Bembidiina R-JC01
         |--Bembidion LM87
         |--Notaphus Dejean 1821 R-JC01, BB01
         |    |--N. varius (Olivier 1795) LM87 (see below for synonymy)
         |    |--N. brullei (Gemminger & Harold 1868) (see below for synonymy) LM87
         |    `--N. convergens [=Austronotaphus convergens] R-JC01
         `--Peryphus R-JC01

Bembidiini incertae sedis:
  Lymnastis Motschulsky 1862 B08
    |--*L. indicus (Motschulsky 1851) [=Lymnaeum indicum] B08
    |--L. atricapillus B08
    |--L. brooksi Baehr 2008 B08
    |--L. coomani B08
    |--L. decorsei B55
    |--L. inops B08
    |--L. macrops B55
    |--L. niloticus B55
    |--L. pilosus Bates 1892 [incl. Tachys setiger Sloane 1903] LM87
    `--L. pullulus B08
  Brachytachys Basilewsky 1953 B55
    |--B. curtulus Basilewsky 1953 B55
    `--B. villiersi Basilewsky 1953 B55
  Sphaerotachys B55
    |--S. fumigatus B55
    `--S. haemorrhoidalis B55
  Micratopus Casey 1914 [Micratopini] BB01
    `--M. aenescens (LeConte 1848) BB01
  Anillinus Casey 1918 [incl. Troglanillus Jeannel 1963] BB01
    `--A. valentinei (Jeannel 1963) BB01
  Asaphidion des Gozis 1886 BB01
    |--A. cyanicorne F05
    |    |--A. c. cyanicorne F05
    |    `--A. c. atlanticum Ortuño & Toribio 2005 F05
    |--A. flavipes (Linnaeus 1761) [=Cicindela flavipes] B55
    `--A. stierlini (Heyden 1880) VPB11
  Mioptachys Bates 1882 [incl. Tachymenis Motschulsky 1862 (preoc.)] BB01
    `--M. flavicauda (Say 1823) B14
  Tachyta Kirby 1837 B90, LM87
    |  i. s.: T. subvirens B55
    `--T. (Tachyta) LM87
         |--T. (T.) nana LM87
         |    |--T. n. nana LM87
         |    `--T. n. inornatus (Say 1823) [incl. *T. picipes Kirby 1837] LM87
         `--T. (T.) brunnipennis (Macleay 1871) [=Bembidium brunnipenne] LM87
  Porotachys Netolitzky 1914 BB01
    `--P. bisulcatus (Nicolai 1822) BB01
  Illaphanus Macleay 1865 LB91, LM87
    |--*I. stephensii Macleay 1865 LM87
    `--I. macleayi Lea 1906 LM87
  Phrypeus Casey 1924 BB01
    `--P. rickseckeri (Hayward 1897) BB01
  Elaphropus BB01
  Polyderis Motschulsky 1862 [incl. Microtachys Casey 1918] BB01
    |--*P. brevicornis (Chaudoir 1846) [=Tachys brevicornis] LM87
    |--P. antigua Erwin 1971 BB01
    |--P. laevis (Say 1823) BB01
    `--P. leleupi Basilewsky 1953 B55
  Paratachys Casey 1918 BB01
    `--*P. austinicus Casey 1918 LM87
  Micranillodes Jeannel 1963 BB01
    `--M. depressus Jeannel 1963 BB01
  Anillodes Jeannel 1963 BB01
  Serranillus Barr 1995 BB01
    `--S. jeanneli Barr 1995 BB01
  Anillaspis Casey 1918 BB01
  Horologion Valentine 1932 BB01
    `--H. speokoites Valentine 1932 BB01

Notaphus brullei (Gemminger & Harold 1868) [=Bembicidium brullei, Bembidion (Notaphus) brullei, Bembidion variegatum Brullé 1838 non Say 1823] LM87

Notaphus varius (Olivier 1795) LM87 [=Carabus varius LM87, Bembidion (*Notaphus) varium LM87, Elaphrus (Bembidion) varius G20]

*Type species of generic name indicated


[B02] Baehr, M. 2002. New species and additional records of the tachyine Philipis Erwin (Coleoptera: Carabidae: Bembidiinae). Memoirs of the Queensland Museum 48: 1–7.

[B08] Baehr, M. 2008. A new cavernicolous and eyeless species of the genus Lymnastis from Christmas Island, Australia (Insecta: Coleoptera: Carabidae: Bembidiinae). Records of the Western Australian Museum 24 (2): 199–203.

[BB01] Ball, G. E., & Y. Bousquet. 2001. Carabidae Latreille, 1810. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 32–132. CRC Press: Boca Raton.

[B55] Basilewsky, P. 1955. Carabidae (Coleoptera) de l’Angola (première partie). Diamang Publicações Culturais 27: 91–137.

[B90] Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

[VPB11] Varet, M., J. Pétillon & F. Burel. 2011. Comparative responses of spider and carabid beetle assemblages along an urban-rural boundary gradient. Journal of Arachnology 39 (2): 236–243.

Last updated: 20 December 2020.

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