Apotomus rufus, copyright elleelle.

Belongs within: Conjunctae.
Contains: Broscini.

The Broscinae are a group of ground beetles characterised by a pedunculate connection between the pro- and mesothorax, and a single supraorbital seta. Members of the genus Apotomus, which have elongate palps and an unmargined prothorax, are found on mudflats adjacent to rivers (Bouchard 2014).

<==Broscinae [Broschidae, Broscitae]
    |--+--Broscini R-JC01
    |  `--Melaenitae [Melaeninae] R-JC01
    |       |--Melaenus [Melaenini] R-JC01
    |       |    `--M. piger R-JC01
    |       `--Cymbionotum B14 [Cymbionotini R-JC01]
    |            `--C. (Procoscinia) B14
    |                 |--C. (P.) fernandezi Ball & Shpeley 2005 B14
    |                 `--C. (P.) negrei B14
    `--Apotomus Illiger 1807 R-JC01, LM87 [Apotominae, Apotomini, Apotomitae]
         |--*A. rufus (Rossi 1790) [=Scarites rufus] LM87
         |--A. australis Castelnau 1867 LM87
         |--A. fuscus B89
         |--A. hirsutulus R-JC01
         |--A. mastersii Macleay 1871 LM87
         |--A. novaehollandiae Castelnau 1867 LM87
         |--A. reichardti Erwin 1980 B14
         `--A. xanthotelus Bates 1874 B89

Broscinae incertae sedis:
  Brithysternum Macleay 1873 LM87
    |--*B. calcaratum Macleay 1873 LM87
    |--B. macleayi Sloane 1910 LM87
    `--B. nodosum Sloane 1910 LM87
  Adotela Castelnau 1867 LM87
    |--A. carenoides Putzeys 1873 LM87
    |--A. concolor Castelnau 1867 LM87
    |--A. esmeralda Castelnau 1867 LM87
    |--A. frenchi Sloane 1890 LM87
    |--A. nigerrima Macleay 1873 LM87
    |--A. striolata Putzeys 1873 LM87
    `--A. viridis (Macleay 1871) [=Promecoderus viridis] LM87
  Parroa Castelnau 1867 LM87
    |--P. apicalis Sloane 1893 LM87
    |--P. atronitens (Sloane 1890) [=Adotela atronitens] LM87
    |--P. australis (Sloane 1890) [=Adotela australis] LM87
    |--P. bicolor Castelnau 1867 LM87
    |--P. carbonaria Castelnau 1867 LM87
    |--P. grandis Castelnau 1867 LM87
    |--P. howittii Castelnau 1867 LM87
    |--P. laevigata Sloane 1893 LM87
    |--P. noctis Sloane 1893 LM87
    `--P. violacea Castelnau 1867 LM87
  Cerotalis Castelnau 1867 LM87
    |--C. amabilis Sloane 1890 LM87
    |--C. brachypleura Sloane 1898 LM87
    |--C. longipes Sloane 1898 LM87
    |--C. majuscula (Putzeys 1868) [=Promecoderus majusculus] LM87
    |--C. semiviolacea Castelnau 1867 LM87
    |--C. substriata Castelnau 1867 LM87
    `--C. versicolor Castelnau 1867 LM87
  Gnathoxys Westwood 1842 LM87
    |--G. barbatus Macleay 1864 LM87
    |--G. cicatricosus Reiche 1842 LM87
    |--G. crassipes Sloane 1898 LM87
    |--G. foveatus Macleay 1866 LM87
    |--G. granularis Westwood 1842 [incl. G. blissii Macleay 1866] LM87
    |--G. humeralis M80
    |--G. insignitus Macleay 1864 LM87
    |--G. irregularis Westwood 1842 LM87
    |--G. macleayi Putzeys 1868 LM87
    |--G. murrumbidgensis Macleay 1865 LM87
    |--G. obscurus Reiche 1842 LM87
    |--G. punctipennis Macleay 1873 LM87
    |--G. submetallicus Macleay 1864 LM87
    |--G. sulcicollis Sloane 1910 LM87
    |--G. tesselatus Macleay 1864 LM87
    `--G. westwoodi Putzeys 1868 LM87
  Chylnus Sloane 1920 [=Lychnus Putzeys 1868 non Matheson 1832] LM87
    `--*C. ater (Putzeys 1868) (see below for synonymy) LM87
  Percolestus Sloane 1892 LM87
    `--*P. blackburni Sloane 1892 LM87
  Percosoma Schaum 1858 LM87
    |--*P. carenoides (White 1846) (see below for synonymy) LM87
    |--P. concolor Sloane 1892 LM87
    |--P. montanum (Castelnau 1867) [=Mecodema montanum] LM87
    |--P. substriatum Moore 1960 LM87
    `--P. sulcipenne Bates 1878 LM87

*Chylnus ater (Putzeys 1868) [=*Lychnus ater; incl. L. strangulatus Bates 1878, L. striatulus Bates 1878] LM87

*Percosoma carenoides (White 1846) [=Broscus carenoides; incl. B. hopei Sturm 1843 (n. n.), Mecodema percoides Castelnau 1867] LM87

*Type species of generic name indicated


[B89] Bates, H. W. 1889. Viaggio di Leonardo Fea in Birmani e regioni vicine. XVI.—On some Carabidae from Burma collected by Mr. L. Fea. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 100–111.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[M80] Matthews, E. G. 1980. A Guide to the Genera of Beetles of South Australia. Part 1. Archostemata and Adephaga. South Australian Museum: Australia.

[R-JC01] Roig-Juñent, S., & A. C. Cicchino. 2001. Chaltenia patagonica, new genus and species belonging to Chalteniina, a new subtribe of Zolini (Coleoptera: Carabidae). Canadian Entomologist 133: 651–670.

Last updated: 3 November 2018.

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