Melampinae

Eastern melampus Melampus bidentatus, from here.

Belongs within: Pulmonata.

The Melampinae are a group of ellobiid gastropods differing from other members of the family in the presence of a planktonic larval stage (Frias Martins 1996).

Characters (from Frias Martins 1996): Protoconch heterostrophic, showing one-half of nuclear whorls above teleoconch. Reproductive system semidiaulic, with short spermoviduct, non-glandular pallial ducts, spermathecal duct entering at or near proximal end of vagina; penial complex very elongated, with marked proximal and distal portions, penis present at origin of distal portion, or absent with vas deferens opening at proximal end of penial complex through short, straight tube. Visceral nerve ring short.

<==Melampinae [Conovulidae, Melampacea, Melampidae, Melampodinae]
    |--Conovula Schweigger 1820 [Conovulinae] BR05
    `--Melampus Montfort 1810 [=Conovulus Bowdich 1822] BR05
         |--*M. luteus O27
         |--M. adamsianus Pfeiffer 1855 H09
         |--M. bidentatus Say 1822 BF89
         |--M. cristatus Pfeiffer 1855 H09
         |--M. granosa (Hombr. & Jacq. 1853) [=Auricula granosa] H09
         |--M. olivaceus Carpenter 1856 O27
         |--M. parvulus Pferiifer 1856 H09
         |--M. pulchellus (Petit 1843) [=Auricula pulchella] H09
         |--M. stutchburyi H09
         |--M. trifasciatus Kuster 1844 H09
         `--M. variabilis Gassies 1863 H09

*Type species of generic name indicated

REFERENCES

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BF89] Burnham, B. H., & P. E. Fell. 1989. Distribution of Melampus bidentatus (Say) and Succinea wilsoni (Lea) within a tidal marsh in eastern Connecticut. Nautilus 103 (3): 109–112.

Frias Martins, A. M. de. 1996. Relationships within the Ellobiidae. In: Taylor, J. (ed.) Origin and Evolutionary Radiation of the Mollusca pp. 285–294. Oxford University Press.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. II, pt I. Stanford University Press: Stanford University (California).

Dentalium

Dentalium elephantium, from here.

Belongs within: Scaphopoda.

Dentalium is a cosmopolitan genus of tusk shells known from the Middle Triassic to the present day. In members of the type subgenus, the shell is prismatic or strongly ribbed, with a polygonal cross-section toward the apex. The subgenus Antalis is less strongly ribbed than Dentalium sensu stricto, without a polygonal cross-section and often with a V-shaped notch on or near the convex side of the apex. Fissidentalium species are large and solid with many longitudinal ribs and often with a long slit at the apex (Ludbrook 1960).

Characters (from Ludbrook 1960): Shell curved and tapering, longitudinally sculptured, smooth or annulated. Embryonic whorls minute, fragile, lost at an early growth stage. Apex commonly modified by slit or notch due to absorption. Anterior aperture circular or oblique, in many forms modified by longitudinal ribbing.

<==Dentalium Linnaeus 1758 DK08
    |--D. (Dentalium) S98 [incl. Paradentalium Cotton & Godfrey 1933 L60]
    |    |--*D. (D.) elephantium Linnaeus 1758 S98 [incl. D. rectum D26, D. recurvum D26]
    |    |--D. (D.) gouldii E52
    |    |    |--D. g. gouldii E52
    |    |    `--D. g. portoricense Henderson 1920 E52
    |    `--D. (D.) tiwhana Dell 1953 P61
    |--D. (Antalis Adams & Adams 1854) L60 (see below for synonymy)
    |    |--D. (*A.) entalis Linnaeus 1758 L60 (see below for synonymy)
    |    |--D. (A.) bartletti Henderson 1920 [=D. (Dentale) bartletti] E52
    |    |--D. (A.) ceratum Dall 1881 [=D. (Dentale) ceratum; incl. D. ceratum tenax] E52
    |    |--D. (A.) diarrhox Watson 1879 P61
    |    |--D. (A.) glaucarena Dell 1953 P61
    |    |--D. (A.) nanum Hutton 1873 [incl. D. huttoni Kirk 1880, D. marwicki Mestayer 1926] P61
    |    |--D. (A.) pilsbryi GD00, GO06
    |    |--D. (A.) suteri Emerson 1954 [incl. D. arenarium Suter 1907] P61
    |    `--D. (A.) vulgare Da Costa 1778 KC11, E52 [=Antalis vulgaris KC11]
    |--D. (Bathoxiphus Pilsbry & Sharp 1897) L60
    |    |--D. (*B.) ensiculus Jeffreys 1877 L60
    |    `--‘Bathoxiphus’ colmani (Palmer 1974) MG-H11
    |--D. (Coccodentalium Sacco 1896) L60
    |    |--D. (*C.) radula Schröter 1784 L60
    |    `--D. (C.) carduus Dall 1889 E52
    |--D. (Compressidens Pilsbry & Sharp 1897) L60
    |    |--D. (*C.) pressum Pilsbry & Sharp 1897 L60
    |    `--D. (C.) ophiodon Dall 1881 E52
    |--D. (Fissidentalium Fischer 1885) [incl. Schizodentalium Sowerby 1894] L60
    |    |--D. (*F.) ergasticum Fischer 1885 L60
    |    |--D. (F.) mawsoni Ludbrook 1956 DK08
    |    |--‘Fissidentalium’ otatumei AJH07
    |    |--‘Fissidentalium’ shirleyae Lamprell & Healy 1998 MG-H11
    |    `--‘Fissidentalium’ zelandicum (Sowerby 1860) P61
    |--D. (Episiphon Pilsbry & Sharp 1897) L60
    |    |--D. (*E.) sowerbyi Guilding 1834 L60
    |    |--D. (E.) johnsoni Emerson 1952 E52
    |    `--D. (E.) macilentum Pilsbry 1911 E52
    |--D. (Graptacme Pilsbry & Sharp 1897) L60
    |    |--D. (*G.) eboreum Conrad 1846 L60 [=Graptacme eborea WT06]
    |    `--D. (G.) sectum Deshayes 1826 PP78
    |--D. (Heteroschismoides Ludbrook 1960) [=Heteroschisma Simroth 1895 non Wachsmuth 1883] L60
    |    `--D. (*H.) subterfissum Jeffreys 1877) [=*Heteroschisma subterfissum] L60
    |--D. (Lobantale Cossmann 1888) L60
    |    `--D. (*L.) duplex Defrance 1819 L60
    `--D. (Tesseracme Pilsbry & Sharp 1898) L60
         |--D. (*T.) quadrapicale Sowerby 1860 L60
         `--D. (T.) tesseragonum Sowerby 1832 PP78

Dentalium incertae sedis:
  D. abbreviatum Deshayes 1826 D26
  D. acuminatum Deshayes 1826 D26
  D. acuticosta Deshayes 1826 D26
  D. aequatorium O27
  D. agassizii Pilsbry & Sharp 1897 O27
  D. annulosum Brazier 1877 H09
  D. aprinum [incl. D. striatulum] D26
  D. arcuatum D26
  D. batheri Finlay 1927 [=D. huttoni Bather 1905 non Kirk 1880] F27b
  D. bicarinatum Deshayes 1826 D26
  D. bisexangulatum Sowerby 1860 H09
  D. bouei Deshayes 1826 D26
  D. brevifissum Deshayes 1826 D26
  D. candidum [incl. D. solidum Verrill 1884 non Hutton 1873] F27b
  D. canna White 1874 W77
  D. cheverti Sharp & Pilsbry 1897 H09
  D. clathratum Martens 1881 H09
  D. clava D26
  D. coarctatum D26
  D. collinsae Lamprell & Healy 1998 MG-H11
  D. compressum d’Orbigny 1850 F27b
  D. ‘compressum’ Watson 1879 non d’Orbigny 1850 F27b
  D. conicum H86a
  D. conspicuum PR79
  D. crassum Deshayes 1826 D26
  D. dalli Pilsbry & Sharp 1897 O27
  D. dentalis Linnaeus 1758 L58 [incl. D. attenuatum D26]
  D. dispar Sowerby 1860 H09
  D. disparile G79
  D. dufresnii Deshayes 1826 D26
  D. duodecimcostatum Brazier 1877 H09
  D. eburneum D26
  D. fisheri Stearns in Pilsbry 1897 O27
  D. fissura D26
  D. fossile [incl. D. costatum] D26
  D. giganteum H86a
  D. grande Deshayes 1826 D26
  D. hexagonum Gould 1859 MG-H11
  D. inaequicostatum GO06
  D. inversum Deshayes 1826 D26
  D. javanum Sowerby 1860 H09
  D. lacteum Deshayes 1826 D26
  D. laeve Turton 1819 F27b
  D. ‘laevis’ Schlotheim 1820 non D. laeve Turton 1819 F27b
  D. lessoni Deshayes 1826 D26
  D. liratum C64
  D. longitrorsum Reeve 1842 H09
  D. majus H86a
  D. mantelli H86b [incl. D. irregulare H86a, Entalis texturatus H86b]
  D. minutum Linnaeus 1758 L58
  D. multistriatum Deshayes 1826 D26
  D. nebulosum D26
  D. neohexagonum Sharp & Pilsbry 1897 O27
  D. novemcostatum [incl. D. fasciatum] D26
  D. numerosum Dall 1897 [=D. oerstedtii numerosum] O27
  D. octangulatum Donovan 1803 H09
  D. octogonum D26
  D. oerstedtii O27
  D. otamaringaensis Marwick 1926 F27a
  D. pacificum H79
  D. panormum PP64
  D. pareoraensis Pilsbry & Sharp 1897 (see below for synonymy) F27b
  D. politum D26
  D. potteri Lamprell & Healy 1998 MG-H11
  D. pretiosum Sowerby 1860 O27 [incl. D. indianorum C64]
  D. priscum W77
  D. pseudoentalis D26
  D. pseudosexagonum Deshayes 1826 D26
  D. quadrangulare C64
  D. quadricostatum Brazier 1877 H09
  D. rectius Carpenter 1864 C64
  D. semipolitum Broderip & Sowerby 1829 O27 [incl. D. hyalinum C64]
  D. semistriatum Deshayes 1826 D26
  D. sexangulare [incl. D. sexangulum] D26
  D. solidum Hutton 1873 F27b
  D. splendidum C64
  D. strangulatum Deshayes 1826 D26
  D. striatum D26
  D. substriatum Deshayes 1826 D26
  D. sulcatum D26
  D. tenue H86a
  D. translucidum Deshayes 1826 D26
  D. vallicolens Raymond 1904 O27
  D. variabile Deshayes 1826 D26
  D. vernedei PR79
  D. watsoni Sharp & Pilsbry 1897 O27

Dentalium pareoraensis Pilsbry & Sharp 1897 [=D. laevis Hutton 1873 nec D. laeve Turton 1819 nec D. laevis Schlotheim 1820] F27b

Dentalium (Antalis Adams & Adams 1854) L60 [incl. Dentale Da Costa 1778 (nom. rej.) E52, Entaliopsis Newton & Harris 1894 L60, Entalis Gray 1847 non Sowerby 1839 L60; Antalinae]

Dentalium (*Antalis) entalis Linnaeus 1758 L60 [incl. D. abyssorum Sars 1865 N79, D. agile Sars 1872 N79, D. entalis var. agile N79, D. striolatum Stimpson 1851 N79, D. entalis var. striolatum N79, D. tarentinum D26]

*Type species of generic name indicated

REFERENCES

[AJH07] Amano, K., R. G. Jenkins & Y. Hikida. 2007. A new gigantic Nucinella (Bivalvia: Solemyoida) from the Cretaceous cold-seep deposit in Hokkaido, northern Japan. Veliger 49 (2): 84–90.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[DK08] Darragh, T. A., & G. W. Kendrick. 2008. Silicified Eocene molluscs from the Lower Murchison district, Southern Carnarvon Basin, Western Australia. Records of the Western Australian Museum 24 (3): 217–246.

[D26] Deshayes, G.-P. 1826. Anatomie et monographe du genre Dentale. Mémoires de la Société d'Histoire Naturelle de Paris, ser. 2, 2 (2): 321–378, 4 pls.

[E52] Emerson, W. K. 1952. The scaphopod mollusks collected by the first Johnson-Smithsonian deep-sea expedition. Smithsonian Miscellaneous Collections 117 (6): 1–14.

[F27a] Finlay, H. J. 1927a. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F27b] Finlay, H. J. 1927b. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[GD00] Giribet, G., D. L. Distel, M. Polz, W. Sterrer & W. C. Wheeler. 2000. Triploblastic relationships with emphasis on the acoelomates and the position of Gnathostomulida, Cycliophora, Plathelminthes, and Chaetognatha: a combined approach of 18S rDNA sequences and morphology. Systematic Biology 49: 539–562.

[GO06] Giribet, G., A. Okusu, A. R. Lindgren, S. W. Huff, M. Schrödl & M. K. Nishiguchi. 2006. Evidence for a clade composed of molluscs with serially repeated structures: monoplacophorans are related to chitons. Proceedings of the National Academy of Sciences of the USA 103 (20): 7723–7728.

[G79] Guppy, R. J. L. 1879. First sketch of a marine invertebrate fauna of the Gulf of Paria and its neighbourhood. Part I.—Mollusca. Journal of Conchology 2: 151–172.

[H79] Haast, J. von. 1879. Geology of the Provinces of Canterbury and Westland, New Zealand. A report comprising the results of official explorations. "Times" Office: Christchurch.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H86a] Hutton, F. W. 1886a. The Mollusca of the Pareora and Oamaru systems of New Zealand. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 205–237.

[H86b] Hutton, F. W. 1886b. Notes on some Australian Tertiary fossils. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 481–482.

[KC11] Kocot, K. M., J. T. Cannon, C. Todt, M. R. Citarella, A. B. Kohn, A. Meyer, S. R. Santos, C. Schander, L. L. Moroz, B. Lieb & K. M. Halanych. 2011. Phylogenomics reveals deep molluscan relationships. Nature 477: 452–456.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[L60] Ludbrook, N. H. 1960. Scaphopoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I37–I41. Geological Society of America, and University of Kansas Press.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[N79] Norman, A. M. 1879. The Mollusca of the fiords near Bergen, Norway. Journal of Conchology 2: 8–77.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. 2 pt 1. Stanford University Press: Stanford University (California).

[PP64] Peres, J. M., & J. Picard. 1964. Nouveau manuel de bionomie benthique de la mer Mediterranee. Recueil des Travaux de la Station Marine d'Endoume, Bulletin 31 (27): 5–137.

[PR79] Pojeta, J., Jr & B. Runnegar. 1979. Rhytiodentalium kentuckyensis, a new genus and new species of Ordovician scaphopod, and the early history of scaphopod mollusks. Journal of Paleontology 53 (3): 530–541.

[PP78] Poorman, F. L., & L. H. Poorman. 1978. Additional molluscan records from Bahía de Los Angeles, Baja California Norte. Veliger 20 (4): 369–374.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4^th ed. Whitcombe and Tombs Limited: Christchurch.

[S98] Stilwell, J. D. 1998. Late Cretaceous Mollusca from the Chatham Islands, New Zealand. Alcheringa 22 (1): 29–85.

[WT06] Waeschenbach, A., M. J. Telford, J. S. Porter & D. T. J. Littlewood. 2006. The complete mitochondrial genome of Flustrellidra hispida and the phylogenetic position of Bryozoa among the Metazoa. Molecular Phylogenetics and Evolution 40 (1): 195–207.

[W77] White, C. A. 1877. Report upon the invertebrate fossils collected in portions of Nevada, Utah, Colorado, New Mexico, and Arizona, by parties of the expeditions of 1871, 1872, 1873, and 1874. U.S. Geographical Surveys West of the One Hundredth Meridian 4 (1): 1–219, pls 1–21.

Last updated: 10 August 2021.

Mesotritia

Mesotritia nuda, copyright R. Penttinen.

Belongs within: Euphthiracaroidea.

Mesotritia is a semi-cosmopolitan (unknown from Australasia) genus of euphthiracaroid box mites (Subías 2004).

Characters (from Balogh & Balogh 1992): Aspis with one or two lateral carinae. Posterior median apodeme present. Genu IV without solenidion; seta d on tibia IV short, coupled with solenidion. Genital-aggenital suture complete, genital and aggenital plates well separated; genital setae in paraxial position, in longitudinal row; interlocking triangle absent.

<==Mesotritia Forsslund 1963 S04
    |--M. (Mesotritia) [incl. Entomotritia Märkel 1964] S04
    |    |--M. (M.) flagelliformis (Ewing 1909) [=Phthiracarus flagelliformis; incl. *M. testacea Forsslund 1963] S04
    |    |--M. (M.) australis Mahunka 1984 S04
    |    |--M. (M.) brasiliensis (Sellnick 1923) [=Tritia brasiliensis] S04
    |    |--M. (M.) breviseta Niedbala & Schatz 1996 S04
    |    |--M. (M.) clara Starý 1992 S04
    |    |--M. (M.) dissimilis Hammer 1977 [=M. (Entomotritia) dissimilis] S04
    |    |--M. (M.) elegantula Starý 1992 S04
    |    |--M. (M.) exobothridialis Bayartogtokh & Aoki 1998 S04
    |    |--M. (M.) faeroensis (Sellnick 1923) [=Tritia faeroensis] S04
    |    |--M. (M.) glabra Niedbala 2002 S04
    |    |--M. (M.) glabrata (Say 1821) [=Oribata glabrata] S04
    |    |--M. (M.) grandjeani (Feider & Suciu 1957) [=Oribotritia grandjeani] S04
    |    |--M. (M.) indica Sanyal 1988 S04
    |    |--M. (M.) jacoti Niedbala 2001 S04
    |    |--M. (M.) maerkeli Sheals 1965 [incl. M. flagellata Bayoumi & Mahunka 1979, M. spinosa Aoki 1980] S04
    |    |--M. (M.) minima Niedbala 2002 S04
    |    |--M. (M.) nitida Hammer 1977 [=M. (Entomotritia) nitida] S04
    |    |--M. (M.) nova Starý 1992 S04
    |    |--M. (M.) nuda (Berlese 1887) (see below for synonymy) S04
    |    |--M. (M.) okuyamai Aoki 1980 S04
    |    |--M. (M.) ruwenzorii Niedbala 1998 S04
    |    |--M. (M.) similis Niedbala 2000 S04
    |    `--M. (M.) virginiensis (Jacot 1930) [=Oribotritia virginiensis] S04
    `--M. (Perutritia Märkel 1964) S04
         |--M. (*P.) amazonensis (Maerkel 1964) [=*Perutritia amazonensis] S04
         |--M. (P.) asensillata (Ojeda 1992) [=Perutritia asensillata] S04
         `--M. (P.) curviseta (Hammer 1961) [=Oribotritia curviseta] S04

Mesotritia (Mesotritia) nuda (Berlese 1887) [=Tritia nuda; incl. Oribotritia brachythrix Walker 1965, Mesotritia elastica Sergienko 1988, M. (Entomotritia) piffli Märkel 1964] S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Austrotritia

Austrotritia lebronneci, copyright R. Penttinen.

Belongs within: Euphthiracaroidea.

Austrotritia is a genus of euphthiracaroid box mites found primarily in tropical Asia and Oceania (Subías 2004).

Characters (from Balogh & Balogh 1992): Aspis with one pair of exobothridial setae present. Palpi four- or five-segmented. Posterior median apodeme absent. Genu IV with solenidion and seta. Genital-aggenital suture absent, the two plates completely fused; genital setae in paraxial position, in longitudinal row; ano-genital cleft present; interlocking triangle absent.

<==Austrotritia Sellnick 1959 [incl. Macarotritia Pérez-Íñigo 1986] S04
    |--*A. quadricarinata Sellnick 1959 S04
    |--A. bifurca Niedbala 2000 S04
    |--A. carinata Niedbala 2000 S04
    |--A. dentata Aoki 1980 S04
    |--A. gibba Bayoumi & Mahunka 1979 S04
    |--A. glabrata Niedbala 2000 S04
    |--A. herenessica (Pérez-Íñigo 1986) S04 [=Indotritia (*Macarotritia) herenessica BB92]
    |--A. kinabaluensis Ramsay & Sheals 1969 S04
    |--A. lanceolata Aoki 1988 S04
    |--A. lebronneci (Jacot 1934) [=Indotritia lebronneci] S04
    |    |--A. l. lebronneci S04
    |    |--A. l. crassiori Jacot 1934 S04
    |    |--A. l. flagelloides Jacot 1934 S04
    |    `--A. l. tahitiana (Jacot 1934) [=Indotritia lebronneci tahitiana] S04
    |--A. ramsayi Mahunka 1991 S04
    |--A. robusta Niedbala & Corpuz-Raros 1998 S04
    |--A. saraburiensis Aoki 1965 (see below for synonymy) S04
    `--A. unicarinata Aoki 1980 S04

Austrotritia saraburiensis Aoki 1965 [=A. lebronneci saraburiensis; incl. A. ishigakiensis Aoki 1980, A. optabilis Niedbala 1991, A. shealsi Mahunka 1987] S04

*Type species of generic name indicated

REFERENCES

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Indotritia

Indotritia javensis, from here.

Belongs within: Ptyctima.

Indotritia is a cosmopolitan genus of euphthiracaroid box mites (Subías 2004). Most species possess an ano-adanal cleft on the venter and two lateral carinae on the aspis but the ano-adanal cleft is absent and only a single lateral carina is present on the aspis of the Australasian subgenus Zeaotritia (Balogh & Balogh 1992).

Characters (from Balogh & Balogh 1992, as Indotritia + Zeaotritia): Aspis with one or two lateral carinae. Genito-aggenital suture incomplete, genital and aggenital plates anteriorly fused, posteriorly separate; genital setae in paraxial position, in longitudinal row; interlocking triangle absent.

<==Indotritia Jacot 1929 S04
    |  i. s.: I. carolinae (Jacot 1930) A80
    |--I. (Indotritia) S04
    |    |--*I. (I.) krakatauensis (Sellnick 1923) (see below for synonymy) S04
    |    |--I. (I.) africana Mahunka 1984 S04
    |    |--I. (I.) aspera Niedbała 2000 S04
    |    |--I. (I.) bellingeri Niedbała & Schatz 1996 S04
    |    |--I. (I.) breviseta (Berlese 1923) [=Tritia berlesei breviseta] S04
    |    |--I. (I.) bryani (Jacot 1929) (n. d.) [=Euphthiracarus (I.) bryani] S04
    |    |--I. (I.) clavata Wallwork 1977 S04
    |    |--I. (I.) consimilis Märkel 1964 [=I. krakatauensis consimilis] S04
    |    |--I. (I.) jacoti Niedbała 2001 S04
    |    |--I. (I.) javensis (Sellnick 1923) (see below for synonymy) S04
    |    |--I. (I.) missouri Niedbała 2002 S04
    |    |--I. (I.) nuda Mahunka 1988 [incl. I. usambarensis Starý 1993] S04
    |    |--I. (I.) paulyi Niedbała 1998 S04
    |    |--I. (I.) propinqua Niedbała 1991 S04
    |    |--I. (I.) retusa Niedbała & Schatz 1996 S04
    |    |--I. (I.) tripartita Niedbała 1998 S04
    |    |--I. (I.) undulata Bayoumi & Mahunka 1979 S04
    |    `--I. (I.) zangherii Mahunka & Paoletti 1984 S04
    |--I. (Afrotritia Mahunka 1988) S04
    |    `--I. (*A.) compacta Mahunka 1988 S04
    `--I. (Zeaotritia Mahunka 1988) S04
          |--I. (*Z.) aotearoana Ramsay 1966 S04
          |--I. (Z.) brevipilosa Niedbała 2000 S04
          `--I. (Z.) brevisetosa Niedbała 2000 (see below for synonymy) S04

Indotritia (Indotritia) javensis (Sellnick 1923) [=Tritia javensis; incl. I. completa Mahunka 1987, Oribotritia mollis Aoki 1959] S04

*Indotritia (Indotritia) krakatauensis (Sellnick 1923) [=Tritia krakatauensis; incl. I. acanthophora Märkel 1964, I. heterotricha Mahunka 1984, I. sellnicki Aoki 1965, I. septentrionalis Mahunka 1967, I. tropica Starý 1993] S04

Indotritia (Zeaotritia) brevisetosa Niedbała 2000 [incl. I. breviseta Niedbała & Colloff 1997 non Tritia berlesei breviseta Berlese 1923, I. colloffi Balogh & Balogh 2002] S04

*Type species of generic name indicated

REFERENCES

[A80] Aoki, J. 1980. A revision of the oribatid mites of Japan. I. The families Phthiracaridae and Oribotritiidae. Bulletin of the Institute of Environmental Science and Technology, Yokohama National University 6 (2): 1–89.

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 7 March 2019.

Oribotritia

Oribotritia sp., copyright Tom Murray.

Belongs within: Euphthiracaroidea.

Oribotritia is a cosmopolitan genus of euphthiracaroid box mites (Subías 2004).

See also: Oribotritia: some mites just want to be left alone.

Characters (from Balogh & Balogh 1992, as Oribotritia + Berndotritia): Bothridial squama situated above bothridium. Posterior median apodeme present. Genital-aggenital suture complete, genital and aggenital plates well separated; genital setae in paraxial position, in longitudinal row; ano-genital cleft oblique, well developed; interlocking triangle absent.

<==Oribotritia Jacot 1924 (see below for synonymy) S04
    |--O. berlesei (Michael 1898) (see below for synonymy) S04
    |--O. africana Starý 1993 [incl. O. bicarinata Niedbala 1993] S04
    |--O. ampla Niedbala 1991 S04
    |--O. anceps Niedbala 2000 S04
    |--O. angusta Mahunka 1982 S04
    |--O. aokii Mahunka 1987 S04
    |--O. asiatica Hammer 1977 [incl. O. koreense Mahunka 1982] S04
    |    |--O. a. asiatica S04
    |    `--O. a. norikoae Suzuoka 1983 S04
    |--O. attenuata Niedbala & Schatz 1996 S04
    |--O. bagnalli (Hull 1916) [=Phthiracarus bagnalli] S04
    |--O. banksi (Oudemans 1916) [=Tritia banksi] S04
    |--O. bipartita Niedbala 2000 S04
    |--O. brevis Niedbala & Colloff 1997 S04
    |--O. brevisetosa Niedbała 2003 N07
    |--O. bulbifer (Mahunka 1987) S04 [=*Berndotritia bulbifer BB92]
    |--O. capitanea Niedbala & Corpuz-Raros 1998 S04
    |--O. carolinae Jacot 1930 S04
    |--O. cherokee Niedbala 2002 S04
    |--O. chichijimensis Aoki 1980 [incl. O. chichijimensis ryukyuensis Nakatamari 1985] S04
    |--O. contortula Niedbala 1993 S04
    |--O. contraria Niedbala 2000 S04
    |--O. corporaali (Oudemans 1926) [=Tritia corporaali] S04
    |--O. decumana (Koch 1836) (n. d.) [=Hoplophora decumana] S04
    |--O. didyma Niedbala & Schatz 1996 S04
    |--O. duplex Niedbala 2000 S04
    |--O. exilis (Sellnick 1923) [=Tritia exilis] S04
    |--O. fennica Forsslund & Märkel 1963 S04
    |--O. gigas Bayoumi & Mahunka 1979 S04
    |--O. hauseri Mahunka 1982 S04
    |--O. hawaiiensis (Jacot 1929) [=Euphthiracarus (Indotritia) hawaiiensis] S04
    |--O. henicos Niedbala 2002 S04
    |--O. hermanni Grandjean 1967 S04
    |--O. heterotricha Niedbala 2000 S04
    |--O. incognita Niedbala 2000 S04
    |--O. laselve Niedbała 2003 N07
    |--O. magna (Ewing 1907) [=Phthiracarus magnus] S04
    |--O. megale (Walker 1965) [=Plesiotritia megale] S04
    |--O. nepalensis Niedbala 2000 S04
    |--O. opipara Niedbala 2002 S04
    |--O. oregonensis Niedbala 2002 S04
    |--O. paraaokii Niedbala 2000 S04
    |--O. paraspinosa Mahunka 1999 S04
    |--O. pulla Niedbala 1998 S04
    |--O. rafalskii Niedbala 1997 S04
    |--O. samoaensis Niedbala 1998 S04
    |--O. serrata Feider & Suciu 1958 S04
    |--O. serrula Niedbala & Schatz 1996 S04
    |--O. solitaria Niedbala 1993 S04
    |--O. spinosa (Mahunka 1988) [=Philotritia spinosa] S04
    |--O. storkani Feider & Suciu 1957 S04
    |--O. submolesta Niedbala 2000 S04
    |--O. subsolana N01
    |--O. succincta Niedbala 1998 S04
    |--O. teretis Niedbala 1993 S04
    |--O. tiwi Mahunka 1987 S04
    |--O. tokukoae Aoki 1973 S04
    |--O. trisetosa Niedbala & Schatz 1996 S04
    `--O. virgulata Niedbala 2001 S04

Oribotritia Jacot 1924 [incl. Berndotritia Mahunka 1987, Philotritia Mahunka 1988, Plesiotritia Walker 1965, Tritia Berlese 1883 (preoc.)] S04

Oribotritia berlesei (Michael 1898) [=Phthiracarus berlesei; incl. Hoplophora decumana Berlese 1883 non Koch 1836, *Oribotritia decumana] S04

*Type species of generic name indicated

REFERENCES

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[N01] Niedbała, W. 2001. Study of the diversity of Ptyctima (Acari: Oribatida) and quest for centres of its origin: the fauna of the Oriental and Australian regions. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10^th International Congress pp. 143–148. CSIRO Publishing: Melbourne.

[N07] Niedbała, W. 2007. Ptyctimous mites (Acari, Oribatida) of Costa Rica. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 39–42. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Acrotritia

Acrotritia sp., copyright Scott Justis.

Belongs within: Ptyctima.

Acrotritia is a cosmopolitan genus of box mites that has often been referred to by the junior synonymous name Rhysotritia (Subías 2004). The type species A. ardua is cosmopolitan in distribution; its distinctive features include a smooth dorsal surface and a gradually thickened, setulose head on the sensillus.

Characters (from Balogh & Balogh 1992a, as Rhysotritia): Interlamellar setae of normal length, originating far from bothridium. Trochanters III and IV with two setae; genu IV with at least one seta. Notogaster with V-shaped split ventrally. Plates of anogenital region fused into one pair of anogenital plates; one interlocking triangle present in middle of anogenital region; genital setae in paraxial position, in longitudinal row.

<==Acrotritia Jacot 1923 [incl. Rhysotritia Märkel & Meyer 1959] S04
    |--A. ardua (Koch 1841) S04 [=Hoplophora ardua S04, Hummelia ardua G32, *Rhysotritia ardua BB92a]
    |    |--A. a. ardua (see below for synonymy) S04
    |    |--A. a. affinis (Sergienko 1989) [=Rhysotritia ardua affinis] S04
    |    |--A. a. jinyunia (Li & Chen 1990) [=Rhysotritia ardua jinyunia] S04
    |    `--A. a. neotropicalis (Pérez-Íñigo & Baggio 1993) [=Rhysotritia ardua neotropicalis] S04
    |--A. aokii (Niedbała 2000) [=Rhysotritia aokii] S04
    |--A. bipartita (Neidbala 2000) [=Rhysotritia bipartita] S04
    |--A. brasiliana (Mahunka 1983) [=Rhysotritia brasiliana] S04
    |--A. clavata (Märkel 1964) S04 [=Rhysotritia clavata N07]
    |    |--A. c. clavata S04
    |    `--A. c. sextiana (Lions 1966) [=Rhysotritia clavata sextiana; incl. R. riezuae Moreno & Moraza 1994] S04
    |--A. corletti (Mahunka 2000) [=Rhysotritia corletti] S04
    |--A. curticephala (Jacot 1938) [=Pseudotritia ardua curticephala; incl. Rhysotritia lucida Niedbała 1998] S04
    |--A. diaphoros (Niedbała 2002) [=Rhysotritia diaphoros] S04
    |--A. dikra (Niedbała & Schatz 1996) S04 [=Rhysotritia dikra N07]
    |--A. dinota (Niedbała & Schatz 1996) [=Rhysotritia dinota] S04
    |--A. divida (Mahunka 1991) [=Rhysotritia divida] S04
    |--A. dixa (Niedbała & Schatz 1996) S04 [=Rhysotritia dixa N07]
    |--A. duplicata (Grandjean 1953) S04 [=Pseudotritia duplicata S04, Rhysotritia duplicata BB92b]
    |    |--A. d. duplicata S04
    |    `--A. d. limbata (Märkel & Meyer 1959) [=Rhysotritia duplicata limbata] S04
    |--A. ejuncida (Niedbała & Schatz 1996) S04 [=Rhysotritia ejuncida N07]
    |--A. furcata (Bayoumi & Mahunka 1979) [=Rhysotritia furcata] S04
    |--A. gracile (Niedbała 2000) [=Rhysotritia gracile] S04
    |--A. granulata (Mahunka 1999) [=Rhysotritia granulata] S04
    |--A. hallasanensis (Niedbała 2002) [=Rhysotritia hallasanensis] S04
    |--A. hyeroglyphica (Berlese 1916) (see below for synonymy) S04
    |--A. koreensis Mahunka 1997 (see below for synonymy) S04
    |--A. lentula (Koch 1841) (n. d.) [=Tritia lentula] S04
    |--A. loricata (Rathke 1799) (n. d.) S04 [=Trombidium loricatum S04, Pseudotritia loricata S59]
    |--‘Rhysotritia’ meristos Niedbała 2003 N07
    |--A. monodactyla (Niedbała 2002) [=Rhysotritia monodactyla] S04
    |--A. ornata (Niedbała 2002) [=Rhysotritia ornata] S04
    |--A. otaheitensis (Hammer 1972) [=Rhysotritia ardua otaheitensis] S04
    |--‘Rhysotritia’ paenimina W85
    |--‘Rhysotritia’ parallelos Niedbała 2003 N07
    |--A. parareticulata (Niedbała 2002) [=Rhysotritia parareticulata] S04
    |--A. peruensis (Hammer 1961) S04 [=Oribotritia peruensis S04, Rhysotritia peruensis N07]
    |--A. refracta (Niedbała 1998) S04 [=Rhysotritia refracta N07]
    |--A. reticulata (Mahunka 1988) [=Rhysotritia reticulata] S04
    |--A. rustica (Niedbała 1991) [=Rhysotritia rustica] S04
    |--A. scotti (Walker 1965) [=Rhysotritia scotti] S04
    |--A. simile (Mahunka 1982) [=Rhysotritia simile] S04
    |--A. sinensis Jacot 1923 [incl. Rhysotritia clavata spiculifera Mahunka 1991, R. rasile Mahunka 1982] S04
    |--A. sterigma (Niedbała 1998) [=Rhysotritia sterigma] S04
    |--A. vestita (Berlese 1913) (see below for synonymy) S04
    |--‘Rhysotritia’ ventosus Hammer 1961 N01
    `--A. wallworki (Lee 1981) [=Rhysotritia wallworki] S04

Acrotritia ardua ardua (Koch 1841) [incl. Phthiracarus americanus Ewing 1909, *A. americana, Hoplophora arctata Riley 1874, Pseudotritia ardua antetriheterodactylum Jacot 1933, Phthiracarus canestrinii Michael 1898, Ph. pectinatus Ewing 1917] S04

Acrotritia hyeroglyphica (Berlese 1916) [=Tritia lentula var. hyeroglyphica; incl. Rhysotritia hauseri Mahunka 1991, R. ardua penicillata Pérez-Íñigo 1969, Acrotritia sinensis triheterodactylis Jacot 1923] S04

Acrotritia koreensis Mahunka 1997 [incl. Rhysotritia penicillata Mahunka 1982 non R. ardua penicillata Pérez-Íñigo 1969] S04

Acrotritia vestita (Berlese 1913) [=Hoploderma vestitum; incl. Rhysotritia anchiseta Niedbała 1998, R. bifurcata Niedbała 1993, R. comteae Mahunka 1983, R. niedbalai Balogh & Balogh 2002] S04

*Type species of generic name indicated

REFERENCES

[BB92a] Balogh, J., & P. Balogh. 1992a. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[BB92b] Balogh, J., & P. Balogh. 1992b. The Oribatid Mites Genera of the World vol. 2. Plates. Hungarian Natural History Museum: Budapest.

[G32] Grandjean, F. 1932. La famille des Protoplophoridae (acariens). Bulletin de la Société Zoologique de France 57: 10–36.

[N01] Niedbała, W. 2001. Study of the diversity of Ptyctima (Acari: Oribatida) and quest for centres of its origin: the fauna of the Oriental and Australian regions. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10^th International Congress pp. 143–148. CSIRO Publishing: Melbourne.

[N07] Niedbała, W. 2007. Ptyctimous mites (Acari, Oribatida) of Costa Rica. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 39–42. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[S59] Sellnick, M. 1959. Acarina from southeastern Polynesia—II (Oribatidae). Occasional Papers of the Bernice P. Bishop Museum 22 (9): 109–152.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

[W85] Walter, D. E. 1985. The effects of litter type and elevation on colonization of mixed coniferous litterbags by oribatid mites. Pedobiologia 28: 383–387.

Last updated: 7 March 2019.