Spiroloculina

Spiroloculina excavata, copyright Bernard Remaud.

Belongs within: Nubeculariidae.

Spiroloculina is a genus of porcelaneous Foraminifera known from the Upper Cretaceous to the present that is planispiral throughout development (Loeblich & Tappan 1964).

Characters (from Loeblich & Tappan 1964): Test free, commonly with flattened sides and lanceolate or fusiform outline, earliest stage may consist of single chamber completely encircling proloculus, later chambers (or all chambers in megalospheric forms) being added two to whorl on alternate sides and in single plane; wall calcareous, porcelaneous; aperture at open end of final chamber, with simple or bifid tooth.

<==Spiroloculina d’Orbigny 1826 S-VC91 (see below for synonymy)
    |--*S. depressa d’Orbigny 1826 LT64
    |--S. antillarum C40
    |--S. communis S05
    |--S. corrugata S05
    |--S. excavata M62
    |--S. nitida H04
    |--S. parvula S05
    |--S. planulata (Lamarck 1805) S-VC91
    |--S. robusta Brady 1884 [=*Flintia robusta] LT64
    |--S. rostrata M62
    |--S. tenuissima JW99
    |--S. turkomanica Brodsky 1928 G86
    `--S. venusta S05

Spiroloculina d’Orbigny 1826 S-VC91 [=Spiroculina (l. c.) LT64, Spirolocunina (l. c.) LT64; incl. Flintia Schubert 1911 LT64]

*Type species of generic name indicated

REFERENCES

[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3^rd ed. Harvard University Press: Cambridge (Massachusetts).

[G86] Golemansky, V. G. 1986. Rhizopoda: Foraminiferida. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 17–20. E. J. Brill/Dr. W. Backhuys: Leiden.

[H04] Haeckel, E. 1899–1904. Kunstformen der Natur. Bibliographisches Institut: Leipzig und Wien.

[JW99] Jian, Z.-M., L.-J. Wang, M. Kienast, M. Sarnthein, W. Kuhnt, H.-L. Lin & P.-X. Wang. 1999. Benthic foraminiferal paleoceanography of the South China Sea over the last 40,000 years. Marine Geology 156: 159–186.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[M62] Monniot, F. 1962. Recherches sur les graviers a Amphioxus de la région de Banyuls-sur-Mer. Vie et Milieu 13: 231–322.

[S-VC91] Segura-Vernis, L. R., & A. L. Carreño. 1991. Foraminíferos y ostrácodos de la Laguna de La Paz, Baja California Sur, México. Inv. Mar. CICIMAR 6 (1): 195–224.

[S05] Semeniuk, T. A. 2005. Fossil foraminiferal assemblages from Pleistocene seagrass-bank deposits of the southern Perth Basin, Western Australia, and their palaeotemperature implications. Journal of the Royal Society of Western Australia 88 (4): 177–190.

Hemigordiopsidae

Hemigordius schlumbergeri, from Loeblich & Tappan (1964).

Belongs within: Fischerinidae.

The Hemigordiopsidae are a group of porcelaneous Foraminifera with a test that is streptospirally coiled in early stages.

<==Hemigordiopsidae
    |--Hemigordiopsis Reichel 1945 LT64
    |    `--*H. renzi Reichel 1945 LT64
    |--Shanita Brönnimann, Whittaker & Zaninetti 1978 YJ04
    |    |--*S. amosi YJ04
    |    |--S. bronnimanni YJ04
    |    |--S. chagouensis YJ04
    |    `--S. intercalaria YJ04
    `--Hemigordius Schubert 1908 YJ04 (see below for synonymy)
         |--*H. schlumbergeri (Howchin 1895) [=Cornuspira schlumbergeri] LT64
         |--H. biconcavus YJ04
         |--H. calcarea Cushman & Waters 1928 [=*Hemigordiella calcarea] LT64
         |--‘*Ondogordius’ campanula Marie 1961 LT64
         |--H. harltoni C40
         |--‘*Neoangulodiscus’ leischneri Kristan-Tollmann 1962 LT64
         |--H. plautus Wang 1976 W76
         |--H. renzi [incl. Gansudiscus irregularis, G. luquensis, G. orientalis] YJ04
         `--H. zaninettiae YJ04

Hemigordius Schubert 1908 YJ04 [incl. Hemigordiella Marie in Deleau & Marie 1961 LT64, Neoangulodiscus Kristan-Tollmann 1962 LT64, Ondogordius Marie in Deleau & Marie 1961 LT64]

*Type species of generic name indicated

REFERENCES

[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3^rd ed. Harvard University Press: Cambridge (Massachusetts).

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[W76] Wang K. 1976. The foraminifera from the Changhsing Formation in western Guizhou. Acta Palaeontologica Sinica 15 (2): 187–195.

[YJ04] Yang X., Jin X., Ji Z., Wang Y., Yao J. & Yang H. 2004. New materials of the Shanita-Hemigordius assemblage (Permian foraminifers) from the Baoshan Block, western Yunnan. Acta Geologica Sinica (English Edition) 78: 15–21.

Fischerinidae

Cornuspira involvens, from here.

Belongs within: Miliolida.
Contains: Hemigordiopsidae.

The Fischerinidae are a group of porcelaneous Foraminifera known from the Carboniferous to the present, in which the test is undivided after the proloculus (Loeblich & Tappan 1964).

Characters (from Loeblich & Tappan 1964): Test free or attached, proloculus followed by undivided tubular or spreading second chamber; wall calcareous, porcelaneous; aperture terminal, rounded or slitlike.

<==<==Fischerinidae [Cornuspirida, Cornuspiridae, Cornuspiridea, Cornuspirideae]
    |--Fischerininae [Planispirinellinae, Trisegmentininae] LT64
    |    |--Fischerinella Loeblich & Tappan 1962 LT64
    |    |    `--*F. helix (Heron-Allen & Earland 1915) [=Fischerina helix] LT64
    |    |--Nautiloculina Mohler 1938 LT64
    |    |    `--*N. oolithica Mohler 1938 LT64
    |    |--Planispirinella Wiesner 1931 LT64
    |    |    `--*P. exigua (Brady 1879) LT64 [=Hauerina exigua LT64, Planispirina exigua C40]
    |    `--Fischerina Terquem 1878 [incl. Planispirina Seguenza 1880, Trisegmentina Wiesner 1920] LT64
    |         |--*F. rhodiensis Terquem 1878 LT64
    |         |--F. communis (Seguenza 1880) [=*Planispirina communis] LT64
    |         `--F. compressa (Wiesner 1931) (see below for synonymy) LT64
    |--Calcivertellinae LT64
    |    |--Carixia Macfadyen 1941 LT64
    |    |    `--*C. langi Macfadyen 1941 LT64
    |    |--Planiinvoluta Leischner 1961 LT64
    |    |    `--*P. carinata Leischner 1961 LT64
    |    |--Plummerinella Cushman & Waters 1928 LT64
    |    |    `--*P. complexa Cushman & Waters 1928 LT64
    |    |--Calcivertella Cushman & Waters 1928 LT64
    |    |    `--*C. adherens Cushman & Waters 1928 LT64
    |    `--Calcitornella Cushman & Waters 1928 [incl. Apterrinella Cushman & Waters 1928] LT64
    |         |--*C. elongata Cushman & Waters 1928 LT64
    |         |--C. grahamensis (Harlton 1928) [=Tolypammina grahamensis, *Apterrinella grahamensis] LT64
    |         |--C. heathi C40
    |         `--C. stephensi [=Nubecularia stephensi] F71
    `--Cyclogyrinae [Arcornuspirinia, Cornuspirinae, Cornuspirininae] LT64
         |  i. s.: Flectospira Crespin & Belford 1957 LT64
         |           `--*F. prima Crespin & Belford 1957 LT64
         |         Meandrospira Loeblich & Tappan 1946 [incl. Streblospira Crespin & Belford 1957] LT64
         |           |--*M. washitensis Loeblich & Tappan 1946 LT64
         |           `--M. meandrina (Crespin & Belford 1957) [=*Streblospira meandrina] LT64
         |--Orthovertella Cushman & Waters 1928 C40
         |    `--*O. protea Cushman & Waters 1928 LT64
         |--Rectocornuspira Warthin 1930 C40
         |    `--*R. lituiformis Warthin 1930 LT64
         |--Vidalina Schlumberger 1900 C40, LT64 [=Arvidaloum Rhumbler 1913 LT64]
         |    `--*V. hispanica Schlumberger 1900 [=*Arvidaloum hispanicum] LT64
         |--+--Hemigordiopsidae C40
         |  `--Gordiospira Heron-Allen & Earland 1932 C40
         |       `--*G. fragilis Heron-Allen & Earland 1932 C40
         |--+--Cornuspirella Cushman 1928 C40
         |  |    `--*C. diffusa (Heron-Allen & Earland 1913) [=Cornuspira diffusa] LT64
         |  `--Cornuspiroides Cushman 1928 C40
         |       `--*C. striolata (Brady in Tizard & Murray 1882) [=Cornuspira striolata] LT64
         `--Cyclogyra Wood 1842 LT64 (see below for synonymy)
              |--*C. multiplex Wood 1842 LT64
              |--‘Orbis’ foliaceus Philippi 1844 [=*Arcornuspirum foliaceum, *Cornuspira foliacea] LT64
              |--‘Cornuspira’ acicularis S05
              |--‘*Cornicocornuspira’ conica Marie 1961 LT64
              |--‘Cornuspira’ involvens C40
              |--C. planorbis (Schultze 1854) [=Cornuspira planorbis] LT64
              `--‘Cornuspira’ thompsoni C40

Cyclogyra Wood 1842 LT64 [incl. Arcornuspirum Rhumbler 1913 LT64, Conicocornuspira Marie in Deleau & Marie 1961 LT64, Cornuspira Schultze 1854 C40]

Fischerina compressa (Wiesner 1931) [=*Trisegmentina compressa; incl. Hauerina compressa Sidebottom 1904 non d’Orbigny 1846, T. sidebottomi Cushman 1933] LT64

*Type species of generic name indicated

REFERENCES

[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3^rd ed) Harvard University Press: Cambridge (Massachusetts).

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[S05] Semeniuk, T. A. 2005. Fossil foraminiferal assemblages from Pleistocene seagrass-bank deposits of the southern Perth Basin, Western Australia, and their palaeotemperature implications. Journal of the Royal Society of Western Australia 88 (4): 177–190.

Rhapydionininae

Rhapydionina deserta, from Loeblich & Tappan (1964).

Belongs within: Soritidae.

The Rhapydionininae are a group of more or less conical Foraminifera known from the Jurassic to the present, though so far as is known they are extremely rare in the modern fauna (Loeblich & Tappan 1964).

See also: Porcelain fans.

Characters (from Loeblich & Tappan 1964): Test elongate, conical; chambers in rectilinear series, subdivided into chamberlets; aperture terminal, cribrate.

Rhapydionininae LT64
    |--Craterites Heron-Allen & Earland 1924 LT64
    |    `--*C. rectus Heron-Allen & Earland 1924
    |--Praerhapydionina van Wessem 1943 LT64
    |    `--*P. cubana van Wessen 1943 LT64
    |--Rhipidionina Stache 1913 LT64
    |    `--*R. liburnica (Stache 1889) [=Pavonina liburnica] LT64
    |--Ripacubana Loeblich & Tappan 1964 [=Conulina d’Orbigny in de la Sagra 1839] LT64
    |    `--*R. conica (d’Orbigny in de la Sagra 1839) [=*Conulina conica] LT64
    `--Rhapydionina Stache 1913 [=Rhapidionina van den Bold 1946; incl. Haurania Henson 1948] LT64
         |--*R. liburnica (Stache 1889) LT64 (see below for synonymy)
         |--R. deserta (Henson 1948) [=*Haurania deserta] LT64
         |--R. protocaenica C40
         `--R. urensis LT64

*Rhapydionina liburnica (Stache 1889) LT64 [=Peneroplis liburnica LT64, *Rhapidionina liburnica LT64; incl. R. liburnica var. laevigata C40, R. liburnica var. strangulata C40]

*Type species of generic name indicated

REFERENCES

[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3^rd ed. Harvard University Press: Cambridge (Massachusetts).

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

Meandropsininae

Side (left) and apertural views of Fallotia colomi, from Loeblich & Tappan (1964).

Belongs within: Soritidae.

The Meandropsininae are a group of porcelaneous Foraminifera known from the Upper Cretaceous and Palaeocene (Loeblich & Tappan 1964).

Characters (from Loeblich & Tappan 1964): Early stage planispirally coiled, later may be discoidal, operculiform, flabelliform, cylindrical or conical in shape; subepidermal chamberlets in marginal zone, with interseptal pillars; aperture commonly cribrate.

Meandropsininae [Broekininae]
    |--Meandropsina Munier-Chalmas in Schlumberger 1898 [incl. Cyclomeandropsina Henson 1950 (n. n.)] LT64
    |    `--*M. vidali Schlumberger 1898 LT64
    |--Edomia Henson 1948 LT64
    |    `--*E. reicheli Henson 1948 LT64
    |--Pseudedomia Henson 1948 LT64
    |    `--*P. multistriata Henson 1948 LT64
    |--Taberina Keijzer 1945 LT64
    |    `--*T. cubana Keijzer 1945 LT64
    |--Pseudorbitolina Douvillé 1910 LT64
    |    `--*P. marthae Douvillé 1910 LT64
    |--Broeckina Munier-Chalmas 1882 [=Broekina Marie 1958; incl. Praesorites Douvillé 1902] LT64
    |    |--*B. dufresnoyi (d’Archiac in d’Archiac & Haime 1854) [=Cyclolina dufresnoyi, *Broekina dufresnoyi] LT64
    |    `--B. moureti (Douvillé 1902) [=*Praesorites moureti] LT64
    |--Nummofallotia Barrier & Neumann 1959 LT64
    |    |--*N. cretacea (Schlumberger 1899) [=Nonionina cretacea; incl. Goupillaudina sanctipetri Marie 1957] LT64
    |    `--N. apula LSM-CG00
    `--Fallotia Douvillé 1902 [incl. Ayalaina Seiglie 1961, Fascispira Silvestri 1940] LT64
         |--*F. jacquoti Douvillé 1902 LT64
         |--F. colomi (Silvestri 1940) [=*Fascispira colomi] LT64
         `--‘Meandropsina’ rutteni Palmer 1934 [=*Ayalaina rutteni] LT64

*Type species of generic name indicated

REFERENCES

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[LSM-CG00] Luperto Sinni, E., J. Martín-Chivelet & R. Giménez. 2000. Praemurgella valenciana nov. gen. et sp. (Foraminifera) in the Sierra de Utiel Formation (Coniacian–Santonian) of the Prebetic domain (SE Spain). Geobios 32 (2): 145–151.

Soritinae

Sorites orbiculus, copyright Jan Steger.

Belongs within: Soritidae.

The Soritinae are a group of porcellaneous Foraminifera known from the Eocene to the present, characterised by a distinct pattern of chamber addition (Loeblich & Tappan 1964).

Characters (from Loeblich & Tappan 1964): Growth occurs by addition of numerous small chambers in arcuate or annular series, without intercommunications between those of single series, apertures connect adjacent chambers of successive series.

Soritinae [Orbitolitidinae, Orbitolitinae]
    |  i. s.: Orbitolites Lamarck 1801 (see below for synonymy) LT64
    |           `--*O. complanata Lamarck 1801 (see below for synonymy) LT64
    |         Opertorbitolites Nuttall 1925 LT64
    |           `--*O. douvillei Nuttall 1925 LT64
    |         Somalina Silvestri 1939 LT64
    |           `--*S. stefaninii Silvestri 1939 LT64
    |         Yaberinella Vaughan 1928 LT64
    |           `--*Y. jamaicensis Vaughan 1928 LT64
    |--+--Amphisorus Ehrenberg 1839 PHT13, LT64 [incl. Bradyella Munier-Chalmas 1902 LT64]
    |  |    `--*A. hemprichii Ehrenberg 1839 [incl. Orbitolites duplex Carpenter 1883, *Bradyella duplex] LT64
    |  `--Marginopora Quoy & Gaimard in de Blainville 1830 PHT13, LT64
    |       `--*M. vertebralis Quoy & Gaimard in de Blainville 1830 [incl. Orbitolites complanata laciniata] LT64
    `--Sorites Ehrenberg 1839 PHT13, LT64 [=Taramellina Munier-Chalmas 1902 LT64]
         |--S. marginalis (Lamarck 1816) (see below for synonymy) LT64
         |--S. duplex C40
         |--S. orbiculus C-SC03
         `--S. variabilis S05

Orbitolites Lamarck 1801 [=Discolithus Fortis 1802, Orbitolithes Oken 1815, Orbitulites Bronn in Bronn & Roemer 1854 nec Berthold 1827 nec Eichwald 1829 nec Griffith & Pidgeon 1834; incl. Discolites de Montfort 1808] LT64

*Orbitolites complanata Lamarck 1801 [=*Discolithus complanatus, *Orbitulites complanatus; incl. *Discolites concentricus de Montfort 1808] LT64

Sorites marginalis (Lamarck 1816) [=Orbulites marginalis, Orbitolites marginatis; incl. *S. dominicensis Ehrenberg 1839, *Taramellina dominicensis] LT64

*Type species of generic name indicated

REFERENCES

[C-SC03] Cavalier-Smith, T., & E. E.-Y. Chao. 2003. Phylogeny and classification of phylum Cercozoa. Protist 154: 341–358.

[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3^rd ed. Harvard University Press: Cambridge (Massachusetts).

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[PHT13] Pawlowski, J., M. Holzmann & J. Tyszka. 2013. New supraordinal classification of Foraminifera: molecules meet morphology. Marine Micropalaeontology 100: 1–10.

[S05] Semeniuk, T. A. 2005. Fossil foraminiferal assemblages from Pleistocene seagrass-bank deposits of the southern Perth Basin, Western Australia, and their palaeotemperature implications. Journal of the Royal Society of Western Australia 88 (4): 177–190.

Peneroplinae

Live Peneroplis planatus, copyright Martina Prazeres.

Belongs within: Soritidae.

The Peneroplinae are a group of porcelaneous Foraminifera known from the Upper Triassic to the present (Loeblich & Tappan 1964).

Characters (from Loeblich & Tappan 1964): Test close-coiled in early stage, later may be uncoiled or annular; chambers simple, not divided into chamberlets; aperture rounded, slitlike, or series of pores on final septal face.

Peneroplinae [Cristellarinae, Peneroplidinae, Spirolininae]
    |--Praepeneroplis Hofker 1952 [=Protopeneroplis Hofker 1950 non Weynschenk 1950] LT64
    |    `--*P. senoniensis (Hofker 1949) [=Peneroplis senoniensis, *Protopeneroplis senoniensis] LT64
    |--Renulina Lamarck 1804 [=Renulites Lamarck 1804, Renulinites (l. c.)] LT64
    |    `--*R. opercularia Lamarck 1804 [=*Renulites opercularia] LT64
    |--Triasina Majzon 1954 LT64
    |    `--*T. hantkeni Majzon 1954 LT64
    |--Vandenbroeckia Marie 1958 [=Vandenbroekia] LT64
    |    `--*V. munieri Marie 1958 LT64
    |--Monalysidium Chapman 1900 LT64
    |    |--*M. sollasi (Chapman 1900) [=Peneroplis (*Monalysidium) sollasi] LT64
    |    `--M. politum C40
    |--Dendritina d’Orbigny 1826 [incl. Meneghinia Silvestri 1889 non Fucini 1931, Neopeneroplis Didkovskiy 1958] LT64
    |    |--*D. arbuscula d’Orbigny 1826 [=Peneroplis (*Dendritina) arbusculus] LT64
    |    |--‘*Meneghinia’ nautiliformis Silvestri 1889 LT64
    |    `--‘*Neopeneroplis’ sarmaticus Didkovskiy 1958 LT64
    |--Spirolina Lamarck 1804 (see below for synonymy) LT64
    |    |--*S. cylindracea Lamarck 1804 (see below for synonymy) LT64
    |    |--S. arietinus C40
    |    `--‘*Coscinospira’ hemprichii Ehrenberg 1839 LT64
    `--Peneroplis de Montfort 1808 S-VC91 (see below for synonymy)
         |--*P. planatus (Fichtel & Moll 1798) (see below for synonymy) LT64
         |--P. armorica (d’Archiac in Tournour 1868) [=Cyclolina armorica, *Archiacina armorica] LT64
         |--‘Cristellaria’ calcar H04
         |--‘Cristellaria’ compressa H04
         |--‘Cristellaria’ echinata H04
         |--P. karreri Wiesner 1923 [=*Laevipeneroplis karreri] LT64
         |--P. pertesus (Forskal 1775) S-VC91
         |--P. proteus d’Orbigny 1839 [=*Puteolina protea, *Puteolus proteus] LT64
         `--‘Cristellaria’ siddalliana H04

Peneroplis de Montfort 1808 S-VC91 [=Cristellaria Lamarck 1816 LT64; incl. Archiacina Munier-Chalmas in Vasseur 1878 LT64, Laevipeneroplis Šulc 1936 LT64, Puteolina Hofker 1952 LT64, Puteolus Hofker 1950 non Monterosato 1888 LT64]

*Peneroplis planatus (Fichtel & Moll 1798) [=Nautilus planatus, *Cristellaria planata, C. squammula Lamarck 1822] LT64

Spirolina Lamarck 1804 [=Spiralina Brown 1844 non Hartmann 1840, Spirolinites Lamarck 1804, Spirulina Ehrenberg 1843 non Bory 1826; incl. Coscinospira Ehrenberg 1839] LT64

*Spirolina cylindracea Lamarck 1804 [=*Spiralina cylindracea, *Spirolinites cylindracea, *Spirulina cylindracea] LT64

*Type species of generic name indicated

REFERENCES

[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3^rd ed. Harvard University Press: Cambridge (Massachusetts).

[H04] Haeckel, E. 1899–1904. Kunstformen der Natur. Bibliographisches Institut: Leipzig und Wien.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[S-VC91] Segura-Vernis, L. R., & A. L. Carreño. 1991. Foraminíferos y ostrácodos de la Laguna de La Paz, Baja California Sur, México. Inv. Mar. CICIMAR 6 (1): 195–224.