Otitesellinae

Undescribed species of Lipothymus, photographed by Jean-Yves Rasplus.

Belongs within: Chalcidoidea.

The Otitesellinae are a group of wasps that oviposit on figs, either by entering the ostiole or through the fig wall. The larvae feed on the ovarium tissue, which swells to a gall (Bouček 1988).

Characters (from Bouček 1988): Females with whole body black, mostly with some metallic gloss, thorax often finely densely reticulate; forewing with stigmal vein diagonal; gaster tapering posteriorly, apex mostly compressed from side to side and curving downward. Males with reduced wings; antennae inserted behind anterior third of head, their sockets usually wide apart.

<==Otitesellinae B88
    |--Aepocerus B88
    |--Philosycus B88
    |--Eujacobsonia Grandi 1923 B88
    |    `--*E. mirabilis B88
    |--Guadalia Wiebes 1867 B88
    |    `--*G. vissali Wiebes 1967 B88
    |--Micrognathophora Grandi 1921 B88
    |    `--*M. leptoptera B88
    |--Lipothymus Grandi 1921 B88
    |    `--*L. sumatranus B88
    |--Grandiana Wiebes 1961 B88
    |    |--*G. wassae Wiebes 1867 B88
    |    `--G. armadillo Bouček 1988 B88
    |--Micranisa Walker 1875 [incl. Epicolystichus Girault 1915, Sycobiella Westwood 1883] B88
    |    |--*M. pteromaloides [=Idarnes pteromaloides] B88
    |    |--M. aereicorpus (Girault 1915) [=*Epicolystichus aereicorpus] B88
    |    `--‘*Sycobiella’ saundersii B88
    |--Otitesella Westwood 1883 MH11, B88 [incl. Paracolystichus Girault 1915 B88]
    |    |--*O. digitata B88
    |    |--O. amplipennis (Girault 1915) [=Paracolystichus amplipennis] B88
    |    |--O. clarae B88
    |    `--O. compressiventris (Girault 1915) [=*Paracolystichus compressiventris] B88
    `--Heterandrium MH11

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.

Ormocerinae

Female Lisseurytoma violaceitincta, from Bouček (1988).

Belongs within: Chalcidoidea.
Contains: Systasis, Trichilogaster, Terobiella, Encyrtocephalus.

The Ormocerinae are a group of wasps, most diverse in the Australian region, that are all (so far as is known) either gall-makers or inquilines of other gall-forming insects (Bouček 1988).

Characters (from Bouček 1988): Head without occipital carina. Antennae never inserted very low on face, mostly near to, sometimes above centre; either 13-segmented or 12-segmented. Toruli rather close together, except in the few males which have unusually enlarged scapes. Supraclypeal area not well delimited: clypeus rather small, its lower margin sometimes produced, then it may be bilobate. Thorax with complete and deep notauli; scutellum usually narrowed anteriorly, so that axillae are not widely separated (except in few genera). Thoracic dorsum with complete notauli, without special paired setae or bristles; scutellum without distinct frenum. Propodeum not forming a distinct nucha, always relatively short; spiracles usually large. Forewing venation variable but pilosity near basal cell not specialised. Gaster mostly sessile, first tergite not greally enlarged: hind margins of anterior tergites emarginate or excised in middle, latter state mainly in species in which the gaster is not strongly depressed dorsally.

<==Ormocerinae HB13
    |  i. s.: Ormocerus [Ormocerini] B88
    |--Erixestus HB13
    `--+--Systasini HB13
       |    |--Systasis B88
       |    `--Semiotellus Westwood 1839 (see below for synonymy) B88
       |         |--*S. mundus [=*Semiotus mundus] B88
       |         |--S. longifasciatipennis (Girault 1915) [=*Cheiropachysia longifasciatipennis] B88
       |         |--S. ormyroides (Girault 1915) [=Neosystasis ormyroides] B88
       |         |--S. tenebricosus (Girault 1915) [=Cheiropachysia tenebricosa] B88
       |         `--S. umbilicatus (Girault 1915) [=*Neosystasis umbilicatus, *N. umbilicata] B88
       `--Melanosomellini [Brachyscelidiphagini] HB13
            |--Trichilogaster B88
            |--Terobiella B88
            |--Encyrtocephalus B88
            |--Krivena Bouček 1988 B88
            |    `--*K. pennata Bouček 1988 B88
            |--Neoperilampus Girault & Dodd in Girault 1915 B88
            |    `--*N. niger Girault & Dodd in Girault 1915 B88
            |--Aditrochus Ruebsaamen 1902 B88
            |    |--A. coihuensis MH11
            |    `--A. fagicolus B88
            |--Plastobelyta Kieffer 1906 B88
            |    `--P. gallicola B88
            |--Espinosa Gahan 1947 B88
            |    `--E. nothofagi B88
            |--Neochalcissia Girault 1920 B88
            |    `--*N. magniscapus Girault 1920 B88
            |--Megamelanosoma Girault 1928 B88
            |    `--*M. aeschyli Girault 1928 B88
            |--Epelatus Girault 1928 B88
            |    `--*E. eurytomoideus Girault 1928 [=*E. eurytomoidea] B88
            |--Alloderma Ashmead 1904 B88
            |    `--*A. maculatipennis Ashmead 1904 (see below for synonymy) B88
            |--Alyxiaphagus Riek 1962 B88
            |    `--*A. picturatus Riek 1962 B88
            |--Wubina Bouček 1988 B88
            |    `--*W. cardaleae Bouček 1988 B88
            |--Cerna Bouček 1988 B88
            |    `--*C. kohouti Bouček 1988 B88
            |--Westra Bouček 1988 B88
            |    `--*W. nigra Bouček 1988 B88
            |--Hubena Bouček 1988 B88
            |    `--*H. ruris Bouček 1988 B88
            |--Edgaria Bouček 1988 B88
            |    `--*E. blackburni (Girault 1928) [=Tepperella blackburni, Terobiella blackburni] B88
            |--Nambouria Bouček 1988 B88
            |    `--*N. ramulorum Bouček 1988 B88
            |--Cecidoxenus Ashmead 1904 [incl. Parachrysomalla Girault 1915] B88
            |    |--*C. nigrocyaneus Ashmead 1904 [incl. Parachrysomalla secunda Girault 1917] B88
            |    `--C. aereifemur (Girault 1915) [=*Parachrysomalla aereifemur] B88
            |--Perilampomyia Girault 1916 B88
            |    |--*P. notatifrons Girault 1916 B88
            |    `--P. aeschyli Girault 1931 B88
            |--Lisseurytoma Cameron 1912 [incl. Cratodecatoma Cameron 1912, Sphegigasteroides Girault 1913] B88
            |    |--*L. violaceitincta Cameron 1912 (see below for synonymy) B88
            |    `--L. decorata Ferrière in Gahan & Ferrière 1947 B88
            |--Eurytomomma Girault 1920 B88
            |    |--*E. aurifacies Girault 1920 B88
            |    `--E. atricoxa Girault 1927 B88
            |--Xantheurytoma Cameron 1912 B88
            |    |--*X. flava Cameron 1912 B88
            |    `--X. pallidicoxa (Girault 1928) [=Coelocybelloides pallidicoxa] B88
            |--Lincolna Girault 1940 B88
            |    |--*L. aldrovandii Girault 1940 B88
            |    `--L. sismondini (Girault 1931) [=Terobiella sismondini] B88
            |--Aeschylia Girault 1929 B88
            |    |--*A. latiscapus Girault 1929 B88
            |    |--A. mira Bouček 1988 B88
            |    `--A. picta Bouček 1988 B88
            |--Perilampella Girault & Dodd in Girault 1915 (see below for synonymy) B88
            |    |--P. acaciaediscoloris (Froggatt 1892) (see below for synonymy) B88
            |    |--P. hecataeus (Walker 1839) (see below for synonymy) B88
            |    |--P. nigrisetae (Girault 1929) [=Coelocybelloides nigrisetae] B88
            |    `--P. pulchra (Girault 1929) [=Coelocybelloides pulchra] B88
            `--Brachyscelidiphaga Ashmead 1900 [incl. Coelocybella Girault 1913, Coelocybelloides Girault 1915] B88
                 |--*B. flava Ashmead 1900 [incl. B. masaccioni Girault 1931] B88
                 |--B. aeschyli Girault 1931 B88
                 |--B. aurea (Girault 1915) [=*Coelocybelloides aureus] B88
                 |--B. pulchrivariegata (Girault 1915) [=Coelocybelloides pulchrivariegatus] B88
                 |--‘Coelocybelloides’ trilineata G31
                 `--B. variegata (Girault 1913) [=*Coelocybella variegata] B88

*Alloderma maculatipennis Ashmead 1904 [=Coelocybelloides bioculatus Girault 1916, Perilampella punctata Girault 1931] B88

*Lisseurytoma violaceitincta Cameron 1912 [incl. L. bicolor Gahan in Gahan & Ferrière 1947, *Cratodecatoma ruficeps Cameron 1912, *Sphegigasteroides rufinotum Girault 1913] B88

Perilampella Girault & Dodd in Girault 1915 [incl. Trichilogastroides Ferrière in Gahan & Ferrière 1947] B88

Perilampella acaciaediscoloris (Froggatt 1892) [=Cynips acaciaediscoloris, *Trichilogastroides acaciaediscoloris; incl. *Perilampella flammeithorax Girault 1915] B88

Perilampella hecataeus (Walker 1839) [=Miscogaster hecataeus, Lamprotatus hecataeus; incl. Coelocybelloides mediolineatus Girault 1916, Tepperella trilineata Cameron 1912, Tep. trimaculata (l. c.), Terobiella trilineata, Trichilogaster trilineata] B88

Semiotellus Westwood 1839 [=Semiotus Walker 1834 non Eschscholtz 1829; incl. Cheiropachysia Girault 1915, Neosystasis Girault 1915] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[G31] Girault, A. A. 1931. A new habit in an old insect, Homo pudicus and new Eurytomidae. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 280-283).

[HB13] Heraty, J. M., R. A. Burks, A. Cruaud, G. A. P. Gibson, J. Liljeblad, J. Munro, J.-Y. Rasplus, G. Delvare, P. Janšta, A. Gumovsky, J. Huber, J. B. Woolley, L. Krogmann, S. Heydon, A. Polaszek, S. Schmidt, D. C. Darling, M. W. Gates, J. Mottern, E. Murray, A. D. Molin, S. Triapitsyn, H. Baur, J. D. Pinto, S. van Noort, J. George & M. Yoder. 2013. A phylogenetic analysis of the megadiverse Chalcidoidea (Hymenoptera). Cladistics 29: 466-542.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.

Neanastatinae

Potentially undescribed Neanastatus, photographed by Maherjos.

Belongs within: Chalcidoidea.

The Neanastatinae are a group of eupelmid wasps characterised by a large subconical pronotum and limited ability to contract the thorax compared to other eupelmid subfamilies (Bouček 1988).

Characters (from Bouček 1988, as Metapelmatinae): Head with very convex long frons, scrobal depressions almost absent; thorax form the same in both sexes; pronotum undivided, large, conical or almost regularly convex, its hind margin capping the horizontal scutum; scutum with shallow weak notaular depressions; mesopleuron transformed into a large convex shield delimited ventrally by a subhorizontal groove in front of mid coxa; always macropterous, with very long postmarginal vein; mid tibial spur unusually strong; epipygium subconical, cerci not strongly shifted forward.

<==Neanastatinae [Metapelmatinae]
    |--+--Lambdobregma schwarzii HB13
    |  `--Metapelma Westwood 1835 HB13, B88 [incl. Halidayella Dalla Torre 1897 B88, Halidea Förster 1856 B88]
    |       |--*M. spectabilis B88
    |       |--M. bachi Girault 1922 B88
    |       |--M. compressipes B88
    |       |--M. giraulti Ferrière 1938 B88
    |       |--M. goethei Girault 1928 B88
    |       |--‘*Halidea’ nobilis Förster 1860 [=*Hallidayella nobilis] B88
    |       |--M. rufimanum B88
    |       |--M. salomonis Ferrière 1938 B88
    |       |--M. spectabile HB13
    |       `--M. westwoodi Girault 1915 B88
    `--Neanastatus Girault 1913 HB13, B88 [incl. Metaplopoda Masi 1926 B88, Solindenelleus Girault 1914 B88]
         |--*N. cinctiventris Girault 1913 [incl. *Metaplopoda grallaria Masi 1926, N. grallarius] B88
         |--N. aeschyli Girault 1921 B88
         |--N. ariostoni Girault 1922 B88
         |--N. aurifasciatus Girault 1915 B88
         |--N. aurivertex Girault 1915 B88
         |--N. bilingae Girault 1923 B88
         |--N. darci Girault 1933 B88
         |--N. desertensis Girault 1915 B88
         |--N. divinus Girault 1922 B88
         |--N. flavimesopleurum Girault 1915 B88
         |--N. flavipronotum Girault 1915 B88
         |--N. inconspicuus Girault 1915 B88
         |--N. maximicorpus Girault 1915 B88
         |--N. novus Girault 1920 B88
         |--N. parvus Girault 1920 B88
         |--N. pulchricorpus (Girault 1914) [=*Solindenelleus pulchricorpus] B88
         |--N. punctaticeps Girault 1915 B88
         |--N. purpureiscutellum Girault 1915 B88
         |--N. rabelaisi Girault 1922 B88
         |--N. reymondi Girault 1915 B88
         `--N. trinotatus Girault 1915 B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.

Ceratosolen

Female Ceratosolen arabicus on Ficus sycomorus, photographed by Jean-Yves Rasplus.

Belongs within: Agaonidae.

Ceratosolen is a genus of fig-wasps found in Africa, southern Asia and Australasia, pollinating figs of Ficus sections Sycidium, Adenosperma and Sycocarpus (Bouček 1988).

Characters (from Bouček 1988): Females with scape short but stout; forewing venation distinct, normally pigmented; spiracles of sixth tergite surrounded by large concave and elongate peritremal areas with granulate surface; ovipositor sometimes barely protruding. Males with antennae slender, placed in deep channels which are anteriorly separated by a triangular raised area bearing on front margin usually 3 (rarely 2) short teeth; pronotal sides long and converging forward, anteriorly without rim; mid tarsi well developed, in most species 5 -segmented, hind tarsi sometimes unusually large and densely hairy.

Ceratosolen Mayr 1885 [incl. Ceratosolensia Girault 1915, Sycocrypta Coquerel 1855 (nom. rej.)] B88
    |--C. bianchi MH11
    `--+--C. armipes Wiebes 1963 MH11, B88
       `--+--+--*C. appendiculatus (Mayr 1885) B88, MH11, B88 [=Blastophaga (*Ceratosolen) appendiculata B88]
          |  `--C. emarginatus MH11
          `--+--+--C. blommersi MH11
             |  `--+--C. arabicus MH11
             |     `--C. stupefactus MH11
             `--+--+--C. flabellatus MH11
                |  `--C. silvestrianus MH11
                `--+--C. gallii MH11
                   `--C. megacephalus MH11

Ceratosolen incertae sedis:
  C. abnormis Wiebes 1963 B88
  C. adenospermae Wiebes 1965 B88
  C. bimerus Wiebes 1965 B88
  C. bisulcatus (Mayr 1885) [=Blastophaga (Ceratosolen) bisulcata] B88
  ‘*Sycocrypta’ caeca B88
  C. calopilinae Wiebes 1963 B88
  C. dentifer Wiebes 1963 B88
  C. ficophagus (Girault 1915) [=*Ceratosolensia ficophaga] B88
  C. fusciceps (Mayr 1885) [=Blastophaga (Ceratosolen) fusciceps; incl. B. niveipes Girault 1927] B88
  C. grandii Wiebes 1963 B88
  C. gressitti Wiebes 1980 B88
  C. hooglandi Wiebes 1963 B88
  C. immanis Wiebes 1981 B88
  C. indigenus Wiebes 1981 B88
  C. iodotrichae Wiebes 1963 B88
  C. medlerianus Wiebes 1980 B88
  C. nanus Wiebes 1963 B88
  C. nexilis Wiebes 1980 B88
  C. nigriscapus (Girault 1925) [=Blastophaga nigriscapus] B88
  C. notus (Baker 1913) [=Blastophaga nota] B88
  C. solitarius Wiebes 1980 B88
  C. solmsi B88
    |--C. s. solmsi B88
    `--C. s. marchali Mayr 1906 B88
  C. sordidus Wiebes 1963 B88
  C. vissali Wiebes 1981 B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.

Calosotinae

Calosota sp., from here.

Belongs within: Chalcidoidea.
Contains: Eusandalum.

The Calosotinae are a group of wasps that are mostly parasitic on wood-boring beetles (Bouček 1988).

Characters (from Bouček 1988): Thorax capable of limited longitudinal contraction; pronotum undivided; mesoscutum quadrangular or almost so, anteriorly with conspicuous rounded shoulders protruding outside the narrow and low pronotum; notauli present as shallow lines indicated by sculpture and confined to anterior half of scutum which is almost regularly convex; mesopleuron of same shape in both sexes, transformed into a large convex shield delimited ventrally by a subhorizontal groove in front of mid coxa.

<==Calosotinae B88
    |  i. s.: Exosandalum Bouček 1988 B88
    |           |--*E. compressiscapus (Girault 1915) [=Meseusandalum compressiscapus, Eusandalum compressiscapus] B88
    |           `--E. longiannulum (Girault 1929) [=Eusandalum longiannulum] B88
    |--+--Balcha Walker 1862 HB13, B88 [incl. Elemba Cameron 1908 B88, Sauteria Masi 1927 B88]
    |  |    `--B. indica HB13
    |  `--+--Archaeopelma tropeotergum HB13
    |     `--+--Eusandalum HB13
    |        `--Licrooides umbilicatus HB13
    `--Calosota Curtis 1836 HB13, B88 (see below for synonymy)
         |--*C. vernalis [=*Calosoter vernalis] B88
         |--C. charitolophoides (Girault 1915) [=Eupelmus charitolophoides] B88
         |--C. herodoti Girault 1934 B88
         |--C. hirsutioculus Girault 1925 B88
         |--C. metallica B88
         |--C. pagana (Girault 1915) [=Eupelmus paganus] B88
         |--C. parva Girault 1932 B88
         |--C. plutarchi Girault 1932 B88
         |--C. tullii Girault 1934 B88
         `--C. varipunctata Girault 1932 B88

Calosota Curtis 1836 HB13, B88 [=Calosoter Walker 1837 B88; incl. Hylephila Masi 1927 B88, Hylephilisca Ghesquière 1946 B88, Paracalosota Masi 1926 B88]

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[HB13] Heraty, J. M., R. A. Burks, A. Cruaud, G. A. P. Gibson, J. Liljeblad, J. Munro, J.-Y. Rasplus, G. Delvare, P. Janšta, A. Gumovsky, J. Huber, J. B. Woolley, L. Krogmann, S. Heydon, A. Polaszek, S. Schmidt, D. C. Darling, M. W. Gates, J. Mottern, E. Murray, A. D. Molin, S. Triapitsyn, H. Baur, J. D. Pinto, S. van Noort, J. George & M. Yoder. 2013. A phylogenetic analysis of the megadiverse Chalcidoidea (Hymenoptera). Cladistics 29: 466-542.

Tineobius

Female Tineobius, photographed by Stephen Thorpe.

Belongs within: Eupelminae.

Tineobius is a genus of wasps that are either ectoparasitic on caterpillars or hyperparasitic on other parasitoids of those caterpillars (Bouček 1988).

Characters (from Bouček 1988): Head nearly or quite as high as broad, frons less than 1/3 of head breadth; scrobes not deep, not forming carinate channels; scutellum only moderately convex, without a crest of unusual black bristles; wings always developed, postmarginal vein at least twice as long as stigmal vein; hind tibia widened and at least major part of its dorsal laminate edge white; sixth tergite visible; ovipositor distinctly protruding, thin.

Tineobius Ashmead 1896 [incl. Anastatoidea Gahan 1927, Australeupelmus Girault 1921] B88
    |--*T. citri Ashmead 1896 B88
    |--T. beenleighi (Girault 1930) B88 (see below for synonymy)
    |--T. brachartonae [=*Anastatoidea brachartonae] B88
    |--T. columbi (Girault 1923) [=Metapelma columbi, Anastatoidea columbi] B88
    |--T. decoratus (Ferrière 1938) [=Anastatoidea decorata] B88
    |--T. indicus [=Anastatoidea indica] B88
    |--T. longfellowi (Girault 1923) [=Metapelma longfellowi, Anastatoidea longfellowi] B88
    |--T. longicauda [=Anastatoidea longicauda] B88
    |--T. multicolor (Girault 1915) [=Eupelmus multicolor, *Australeupelmus multicolor] B88
    |--T. nassaui (Girault 1933) [=Metapelma (Tineobius) nassaui] B88
    |--T. salomonis (Ferrière 1938) [=Anastatoidea salomonis] B88
    |--T. sidneyi (Girault 1930) [=Metapelma sidneyi] B88
    `--T. superbus (Dodd 1917) [=Metapelma superba, Anastatoidea superba] B88

Tineobius beenleighi (Girault 1930) B88 [=Metapelma beenleighi B88, Anastatoidea beenleighi B88, Matapelma (l. c.) beenleighi G30]

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[G30] Girault, A. A. 1930. New pests from Australia. VII. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 270-272).

Anastatus

Female Anastatus bifasciatus ovipositing on eggs of Nezara viridula, photographed by H. Dumas.

Belongs within: Eupelminae.

Anastatus is a cosmopolitan genus of wasps that parasitise eggs of other insects, mostly Lepidoptera and Heteroptera but also some other taxa such as cockroaches or Orthoptera (Bouček 1988).

Characters (from Bouček 1988): Length of body usually less than 5 mm; first fumcle segment shorter than scape; vertex convex, eye longest in subvertical direction; forewing not unusually long and narrow, stigmal vein normally developed, not very short (or species brachypterous); mid tibia widened at the apex and there, on the anterior side, usually as if partly cleft by a distinct groove outside of a group of several black peg-like spines; gaster posteriorly more or less flattened; apex of last tergite more or less rounded, convex, with a stiffened flat rim which may be slightly upturned.

Anastatus Motschulsky 1859 (see below for synonymy) B88
    |--*A. mantoidae B88
    |--A. aereicorpus (Girault 1925) [=Eupelmus aereicorpus] B88
    |--A. apterus (Girault 1913) [=*Parooderella aptera] B88
    |--A. aristoteleus Girault 1915 [=A. aristotelea] B88
    |--A. atriflagellum (Girault 1924) [=Eupelmus atriflagellum] B88
    |--A. axiagasti Ferrière 1933 [incl. A. axiagasti var. rufithorax Ferrière 1933] B88
    |--A. bernardi Ferrière 1954 AN-A04
    |--A. bifasciatus (Geoffroy in Fourcroy 1785) [=Cynips bifasciatus] AN-A04
    |--A. biproruli (Girault 1925) [=Eupelmus biproruli] B88
    |--A. blattidifurax Girault 1915 B88
    |--A. bombax Girault 1915 B88
    |--A. catalonicus Bolivar 1934 AN-A04
    |--A. darwini Girault 1915 B88
    |--A. dodone (Walker 1839) [=Calosoter dodone, Eupelmus dodone] B88
    |--‘*Cacotropia’ echidna B88
    |--A. extraordinarius (Girault 1929) [=Eupelmus extraordinarius] B88
    |--A. flavithorax (Girault & Dodd in Girault 1915) [=Parooderella flavithorax] B88
    |--A. floridanus HB13
    |--A. fuligispina (Girault 1939) [=Eupelmus fuligispina] B88
    |--A. gibboni Girault 1920 B88
    |--A. giottini (Girault 1922) [=Eupelmus giottini] B88
    |--A. giraudi (Ruschka 1921) [=Eupelmus giraudi; incl. A. dolichopterus Bolívar 1934] AN-A04
    |--A. goethei (Girault 1929) [=Parooderella goethei] B88
    |--A. hemipterus (Girault 1913) [=*Parasolindenia hemiptera, Parooderella hemiptera] B88
    |--A. hookeri (Girault 1915) [=Eupelmus hookeri] B88
    |--‘Halidea’ insignis Förster 1860 B88
    |--A. intonsioculus (Girault 1934) [=Eupelmus intonsioculus] B88
    |--A. japonicus Ashmead 1904 [incl. A. bifasciatus var. disparis Ruschka 1921] AN-A04
    |--A. lichtensteini (Ruschka 1921) [=Eupelmus lichtensteini; incl. A. ameleophagus Bernard 1935] AN-A04
    |--A. lutheri (Girault 1934) [=Parooderella lutheri] B88
    |--A. maculosus Askew in Askew & Nieves-Aldrey 2004 AN-A04
    |--A. madagascariensis (Risbec 1952) [=Oodera madagascariensis; incl. A. (Cladanastatus) umae] B88
    |--A. magnoculus Askew in Askew & Nieves-Aldrey 2004 AN-A04
    |--A. mancus (Girault 1915) [=Parooderella manca] B88
    |--A. maximus Girault 1915 B88
    |--A. motschulskini (Girault 1915) [=Eupelmus motschulskini] B88
    |--A. nasonini (Girault 1925) [=Eupelmus nasonini] B88
    |--‘*Eupelmoides’ obscuratus B88
    |--A. octavia (Girault 1939) [=Eupelmus octavia] B88
    |--A. oscari (Ruthe 1859) [=Eupelmus oscari] AN-A04
    |--A. osmyli (Girault 1927) [=Eupelmus osmyli] B88
    |--A. pasteuri Girault 1915 B88
    |--A. pentatomidivorus Girault 1915 (see below for synonymy) B88
    |--A. pipunculi Perkins 1906 B88
    |--A. poggioni (Girault 1922) [=Eupelmus poggioni] B88
    |--A. racinei Girault 1915 [incl. A. racinei var. praecipuus Girault 1915] B88
    |--A. redini (Girault 1925) [=Eupelmus redini] B88
    |--A. ruficaudus AN-A04
    |--A. saintpierrei Girault 1917 B88
    |--A. semitectus (Girault 1915) [=Parooderella semitecta] B88
    |--A. simplicifrons (Girault 1924) [=Parooderella simplicifrons] B88
    |--A. tennysoni (Girault 1921) [=Eupelmus tennysoni] B88
    |--A. tenuipes Bolivar 1925 [incl. A. blattidarum Ferrière 1930, Solindenia blattiphaga Mani 1936] B88
    |--A. thoreauini Girault 1917 B88
    |--A. tricolor Girault 1915 B88
    `--A. worcesteri (Girault 1915) [=Eupelmus worcesteri, Calosota worcesteri] B88

Anastatus Motschulsky 1859 [incl. Anastatimorpha Erdös 1957, Antigaster Walsh & Riley 1869, Cacotropia Motschulsky 1863, Cerycium Erdös 1946 non Philippi 1841, Cladanastatus Bouček 1979, Eupelmoides Masi 1917, Lutnes Cameron 1884, Misocoris Rondani 1877, Paraguaya Girault 1911, Parasolindenia Girault 1913 non Brues 1907, Paravignalia Risbec 1951, Parooderella Girault 1913, Pseudanastatus Masi 1917, Pseudooderella Brethes 1922, Solindenia Cameron 1883, Vignalia Risbec 1951] B88

Anastatus pentatomidivorus Girault 1915 [=A. pentatomidivora; incl. A. pentatomidivora var. nonexcisus Girault 1915] B88

*Type species of generic name indicated

REFERENCES

[AN-A04] Askew, R. R. & J. L. Nieves-Aldrey. 2004. Further observations on Eupelminae (Hymenoptera, Chalcidoidea, Eupelmidae) in the Iberian Peninsula and Canary Islands, including descriptions of new species. Graellsia 60 (1): 27-39.

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[HB13] Heraty, J. M., R. A. Burks, A. Cruaud, G. A. P. Gibson, J. Liljeblad, J. Munro, J.-Y. Rasplus, G. Delvare, P. Janšta, A. Gumovsky, J. Huber, J. B. Woolley, L. Krogmann, S. Heydon, A. Polaszek, S. Schmidt, D. C. Darling, M. W. Gates, J. Mottern, E. Murray, A. D. Molin, S. Triapitsyn, H. Baur, J. D. Pinto, S. van Noort, J. George & M. Yoder. 2013. A phylogenetic analysis of the megadiverse Chalcidoidea (Hymenoptera). Cladistics 29: 466-542.