Rhombognathus

Rhombognathus magnirostris magnirostris, from the Yale Peabody Museum of Natural History.

Belongs within: Halacaridae.

Rhombognathus is a genus of halacarid mites with minute palpi and two setae each on the ocular plates (Luxton 1990).

Characters (from Luxton 1990): Dorsal plates well developed; ventral plates often fused; ocular plates each with two setae; genital opening of female ventral; legs bidactylous; sclerite between end of leg and base of claws rod-like.

<==Rhombognathus Trouessart 1888 [incl. Aletes Lohmann 1889 nec Rafinesque 1815 nec Carpenter 1857 nec Chapuis 1874] N47
    |--*R. notops (Gosse 1855) (see below for synonymy) N47
    |--R. apsteini Lohmann 1907 N47
    |--R. biscutatus Bartsch 1993 H98
    |--R. caribaeus Bartsch 2007 B07
    |--R. foveolatus Bartsch 1993 H98
    |--R. fractus Bartsch 1979 L90
    |--R. gressitti Newell 1967 [incl. R. magnirostris var. lionyx Trouessart 1914] SL71
    |--R. intermedius Schulz 1933 N47
    |--R. lacunosus Bartsch 1979 L90
    |--R. latens Bartsch 1993 H98
    |--R. latibulus Bartsch 1993 H98
    |--R. lepidus Bartsch 1993 H98
    |--R. leurodactylus WL09
    |--R. levigatoides DPD16
    |--R. magnirostris Trouessart 1889 B07 [=R. magnipalpus (l. c.) N47]
    |    |--R. m. magnirostris N47
    |    |--R. m. plumifer Trouessart 1889 N47
    |    `--R. m. praegracilis Viets 1939 N47
    |--R. marginalis Bartsch 1993 H98
    |--R. novaezelandicus Bartsch 1986 L90
    |--R. parvulus Viets 1939 N47
    |--R. pectinatus Viets 1936 B07
    |--R. peltatus Viets 1939 B07
    |--R. placidus Bartsch 1993 H98
    |--R. psammophilus Bartsch 1993 H98
    |--R. pseudolionyx Newell & André 1959 B07
    |--R. reticulatus WL09
    |--R. scutulatus Bartsch 1983 H98
    |--R. sphaerorhynchus Police 1909 N47
    |--R. spinipes Viets 1933 N47
    |--R. thalassinus Bartsch 1993 H98
    |--R. uniscutatus Viets 1939 N47
    `--R. validipes Bartsch 2000 B07

*Rhombognathus notops (Gosse 1855) [=Pachygnathus notops, *Aletes notops; incl. A. gracilis Lohmann 1889, Rhombognathus longirostris Lohmann 1889] N47

*Type species of generic name indicated

REFERENCES

[B07] Bartsch, I. 2007. Rhombognathus from the Caribbean area, with the description of a new species (Arachnida, Acari, Halacaridae). Senckenbergiana Biologica 87 (1): 17–22.

[DPD16] Dabert, M., H. Proctor & J. Dabert. 2016. Higher-level molecular phylogeny of the water mites (Acariformes: Prostigmata: Parasitengonina: Hydrachnidiae). Molecular Phylogenetics and Evolution 101: 75–90.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[L90] Luxton, M. 1990. The marine littoral mites of the New Zealand region. Journal of the Royal Society of New Zealand 20 (4): 367–418.

[N47] Newell, I. M. 1947. A systematic and ecological study of the Halacaridae of eastern North America. Bulletin of the Bingham Oceanographic Collection 10 (3): 1–232.

[SL71] Spain, A. V., & M. Luxton. 1971. Catalog and bibliography of the Acari of the New Zealand subregion. Pacific Insects Monograph 25: 179–226.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

Halacarus

Halacarus frontiporus, from the Yale Peabody Museum of Natural History.

Belongs within: Halacaridae.

Halacarus is a genus of halacarid mites in which genu I is subequal in length to the adjoining femur and tibia (Luxton 1990).

Characters (from Luxton 1990): Ocular and posterior dorsal plates may be reduced or absent; anterior epimeral plates not divided into two parts; five large dorsal pores often present; palps four-segmented, penultimate segment with a spiniform seta; genu I subequal in length to femur I and tibia I, genu I with one pair of spiniform setae and tibia I with two pairs.

<==Halacarus Gosse 1855 B05
    |--H. (Halacarus) N47
    |    |--*H. (H.) ctenopus Gosse 1855 N47
    |    |--H. (H.) actenos Trouessart 1889 N47
    |    |--H. (H.) anomalus Trouessart 1894 N47
    |    |--H. (H.) frontiporus Newell 1947 N47
    |    |--H. (H.) oblongus Lohmann 1893 N47
    |    `--H. (H.) subtilis Viets 1940 N47
    `--H. (Thalassarachna Packard 1871) (see below for synonymy) N47
         |--H. (*T.) basteri (Johnston 1836) (see below for synonymy) N47
         |    |--H. b. basteri M62
         |    `--H. b. affinis M62
         |--H. (T.) longipes (Trouessart 1888) (see below for synonymy) N47
         |--‘Thalassarachna’ lubricus (Bartsch 1984) [=Halacarellus lubricus] H98
         |--H. (T.) novus Lohmann 1908 SL71
         |--H. (T.) procerus Viets 1927 N47
         |--H. (T.) psammophilus Bartsch 1993 H98, WL09, H98 [=Thalassarachna psammophila WL09]
         `--‘Thalassarachna’ raphidochela WL09

Halacarus incertae sedis:
  H. abyssorum Trouessart 1896 T96
  H. arenarius Bartsch 1993 H98
  H. bisulcus Viets 1927 M62
  H. borealis Trouessart 1894 T05
  H. caudani Trouessart 1896 T96
  H. celatus Bartsch 1993 H98
  H. chevreuxi T96
  H. discophorus Bartsch 1993 H98
  H. fabricii T96
  H. flavellus Bartsch 1993 H98
  H. fuscatus Bartsch 1993 H98
  H. gracileunguiculatus Lohmann 1908 SL71
  H. gracilipes [incl. H. gracilipes var. quadricostatus Trouessart 1896] T96
  H. harioti Trouessart 1889 SL71
  H. helenae Bartsch 1993 H98
  H. humerosus M62
    |--H. h. humerosus M62
    `--H. h. graveolus Monniot 1962 M62
  H. longior Bartsch 1981 B05
  H. mitrellus Bartsch 1993 H98
  H. nitidus Bartsch 1979 L90
  H. parvulus Bartsch 1993 H98
  H. striatus T96

Halacarus (Thalassarachna Packard 1871) [incl. Leptopsalis Trouessart 1888 non Thorell 1882, Leptosalis (l. c.), Leptospathis Trouessart 1894] N47

Halacarus (*Thalassarachna) basteri (Johnston 1836) [=Acarus basteri, Bdella basteri, Halacarellus basteri; incl. H. spinifer Lohmann 1889, H. spiniger, Thalassarachna verrilli Packard 1871, Halacarus verrilli] N47

Halacarus (Thalassarachna) longipes (Trouessart 1888) [=*Leptopsalis longipes, Halacarellus longipes, *Leptospathis longipes; incl. Halacarus levipes Trouessart 1889, Halacarus murrayi Lohmann 1889] N47

*Type species of generic name indicated

REFERENCES

[B05] Bartsch, I. 2005. Halacaridae from the depths of western Antarctica (Arachnida, Acari). Senckenbergiana Biologica 85 (1): 31–41.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[L90] Luxton, M. 1990. The marine littoral mites of the New Zealand region. Journal of the Royal Society of New Zealand 20 (4): 367–418.

[M62] Monniot, F. 1962. Recherches sur les graviers a Amphioxus de la région de Banyuls-sur-Mer. Vie et Milieu 13: 231–322.

[N47] Newell, I. M. 1947. A systematic and ecological study of the Halacaridae of eastern North America. Bulletin of the Bingham Oceanographic Collection 10 (3): 1–232.

[SL71] Spain, A. V., & M. Luxton. 1971. Catalog and bibliography of the Acari of the New Zealand subregion. Pacific Insects Monograph 25: 179–226.

[T05] Trägårdh, I. 1905. Monographie der arktischen Acariden. In: Römer, F., & F. Schaudinn (eds) Fauna Arctica. Eine Zusammenstellun der arktischen Tierformen, mit besonder Berücksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das Nördliche Eismeer im Jahre 1898 vol. 4 pp. 1–78. Gustav Fischer: Jena.

[T96] Trouessart, E. 1896. Resultats scientifiques de la campagne du "Caudan" dans le Golfe de Gascogne: Halacariens. Annales de l'Universite de Lyon 26: 325–353, pls 9–11.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

Agauopsis

Agauopsis microrhyncha, from Lohmann (1893).

Belongs within: Halacaridae.

Agauopsis is a genus of halacarid mites in which the first pair of legs is armed with stout, peg-like setae (Luxton 1990).

Characters (from Luxton 1990): Female with three pairs of perigenital setae; moveable digits of chelicerae denticulate; palps four-segmented, penultimate segment with a spiniform seta; tibia I with unpaired spiniform setae, genu I shorter than femur I or tibia I.

<==Agauopsis Viets 1927 H98
    |--*A. brevipalpus (Trouessart 1889) [=Agaue brevipalpus, Agauopsis brevirostris (l. c.)] N47
    |    |--A. b. brevipalpus N47
    |    `--A. b. pontica (Cichkoff 1907) N47
    |--A. aequilivestita Bartsch 1996 H98
    |--A. australiensis Bartsch 1996 H98
    |--A. borealis Newell 1947 N47
    |--A. collaris Otto 1994 H98
    |--A. curvata WP99
    |--A. elaborata Bartsch 1996 H98
    |--A. filirostris WP99
    |--A. luxtoni Bartsch 1986 L90
    |--A. microrhyncha (Trouessart 1889) [=Agaue microrhyncha] H98
    |--A. mokari Otto 1994 H98
    |--A. novaezelandiae Bartsch 1986 L90
    |--A. ornatella Bartsch 1996 H98
    `--A. similis Bartsch 1979 H98

*Type species of generic name indicated

REFERENCES

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[L90] Luxton, M. 1990. The marine littoral mites of the New Zealand region. Journal of the Royal Society of New Zealand 20 (4): 367–418.

[N47] Newell, I. M. 1947. A systematic and ecological study of the Halacaridae of eastern North America. Bulletin of the Bingham Oceanographic Collection 10 (3): 1–232.

[WP99] Walter, D. E., & H. C. Proctor. 1999. Mites: Ecology, Evolution and Behaviour. CABI Publishing: Wallingford (UK).

Penthaleidae

Blue oat mite Penthaleus major, copyright Sandra-Photographie.

Belongs within: Eupodoidea.

The Penthaleidae are a group of often phytophagous mites, some of which are significant agricultural pests. They are soft-bodied with weakly chelate chelicerae, fifteen or more pairs of setae in the intercoxal region, and a dorsal or dorsoterminal anal opening (Walter et al. 2009).

<==Penthaleidae
    |--Chromotydaeus Berlese 1903 H98
    |    |--C. arcticus (Trägårdh 1900) [=Penthaleus arcticus] T05
    |    |--C. egregius (Berlese 1891) WL09, SB63 (see below for synonymy)
    |    |--C. ovatus T05
    |    |--C. quartus Qin & Halliday 1996 H98
    |    `--C. villosus [=Penthaleus villosus] T05
    |--Halotydeus Berlese 1891 H98
    |    |--*H. hydrodromus (Berlese & Trouessart 1889) [=Notophallus hydrodromus] QH96
    |    |    |--H. h. hydrodromus QH96
    |    |    `--H. h. albolineatus Halbert 1915 QH96, B96
    |    |--H. anthropus Qin & Halliday 1996 QH96
    |    |--H. bakerae Qin & Halliday 1996 H98
    |    |--H. castellus Qin & Halliday 1996 H98
    |    |--H. destructor (Tucker 1925) H98 [=Penthaleus destructor H98, Halotydaeus (l. c.) destructor H01]
    |    |--H. mollis Luxton 1986 L86
    |    |--H. signiensis WL09
    |    `--H. spectatus Qin & Halliday 1996 QH96
    |--Linopenthaloides novaezealandicus QH96, WL09
    `--Penthaleus Koch 1836 [incl. Notophallus Canestrini 1886] H98
         |--*P. major (Dugès 1834) SB63 (see below for synonymy)
         |--P. crinitus Narayan 1957 N57
         |--P. falcatus Qin & Halliday 1996 H98
         |--P. haematopus Koch 1840 [=Notophallus haematopus] T05
         |--P. illustris Berlese 1916 B16
         |--P. insulanus Thorell 1871 [=Notophallus insulanus; incl. Tetranychus borealis Koch 1878] T05
         |--P. longipilis (Canestrini 1886) SB63 (see below for synonymy)
         |--P. medius Schweizer & Bader 1963 SB63
         |--P. minor (Canestrini 1886) [=Notophallus minor] H98
         |--P. ovatus CF77
         `--P. tectus WL09

Chromotydaeus egregius (Berlese 1891) WL09, SB63 [=Penthaleus egregius SB63, Halotydeus egregius B96, Penthalodes egregius SB63]

Penthaleus longipilis (Canestrini 1886) SB63 [=Notophallus longipilis SB63; incl. P. longilis (l. c.) var. longipes Schweizer 1951 S51]

*Penthaleus major (Dugès 1834) SB63 [=Tetranychus majeur H98, P. maior (l. c.) H98; incl. Notophallus bicolor Froggatt 1921 H98, Notophallus bicolour (l. c.) H98, Notophalus dicolor (l. c.) H98, Penthaleus bicolor H98]

*Type species of generic name indicated

REFERENCES

[B96] Baker, A. S. 1996. On the status of some species of Eupodoidea (Acariformes: Prostigmata) represented in the Berlese Acaroteca. In: Mitchell, R., D. J. Horn, G. R. Needham & W. C. Welbourn (eds) Acarology IX vol. 1. Proceedings pp. 373–375. Ohio Biological Survey: Columbus (Ohio).

[B16] Berlese, A. 1916. Centuria terza di Acari nuovi. Redia 12: 289–338.

[CF77] Canestrini, G., & F. Fanzago. 1877. Intorno agli Acari Italiani. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, Serie 5, 4: 69–208, pls 2–7.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[H01] Halliday, R. B. 2001. 250 years of Australian acarology. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10^th International Congress pp. 3–16. CSIRO Publishing: Melbourne.

[L86] Luxton, M. 1986. Mites (Arachnida: Acari) from the coast of Hong Kong. In: B. Morton (ed.) Proceedings of the Second International Marine Biological Workshop: The Marine Flora and Fauna of Hong Kong and Southern China, Hong Kong, 1986 pp. 667–671. Hong Kong University Press.

[N57] Narayan, D. S. 1957. A new species of Penthaleus from Mexico (Acarina: Penthaleidae). Journal of the Kansas Entomological Society 30 (3): 111–113.

[QH96] Qin, T. K., & R. B. Halliday. 1996. Revision of the Australian and South African species of Halotydeus (Acarina: Penthaleidae). Bulletin of Entomological Research 86: 441–450.

[S51] Schweizer, J. 1951. Die Landmilben des schweizerischen Nationalparkes. 2. Teil: Trombidiformes. Ergebnisse der wissenschaftlichen Untersuchung des schweizerischen Nationalparks [Résultats des recherches scientifiques entreprises au Parc National Suisse] 3 (Neue Folge): 51–172.

[SB63] Schweizer, J., & C. Bader. 1963. Die Landmilben der Schweiz (Mittelland, Jura und Alpen): Trombidiformes Reuter, mit 217 Arten und Unterarten und 193 Originalzeichnungen. Denkschriften der Schweizerischen Naturforschenden Gesellschaft [Mémoires de la Société Helvétique des Sciences Naturelles] 84 (2): i–vi, 209–378.

[T05] Trägårdh, I. 1905. Monographie der arktischen Acariden. In: Römer, F., & F. Schaudinn (eds) Fauna Arctica. Eine Zusammenstellun der arktischen Tierformen, mit besonder Berücksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das Nördliche Eismeer im Jahre 1898 vol. 4 pp. 1–78. Gustav Fischer: Jena.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

Speleognathinae

Dorsal (left) and ventral views of Boydaia sturni, from Skoracki et al. (2012).

Belongs within: Procurvata.

The Speleognathinae are a group of mites endoparasitic in the nasal mucosa of birds and mammals. They have no active instars between larva and adult with all three nymphal instars being calyptostatic within the larval skin (Walter et al. 2009).

<==Speleognathinae [Speleognathidae] AF00
    |  i. s.: Meropiboydaia Fain 1985 AF00, H98
    |           `--M. merops (Fain 1955) [=Boydaia merops, Aureliania merops, Neoboydaia merops] H98
    |         Speleochir aitkeni AF00
    |         Speleognathopsis Cooreman 1954 AF00, H98
    |           |--S. accipitris (Domrow 1969) [=Ophthalmognathus accipitris] H98
    |           |--S. benoiti Fain 1955 H98
    |           `--S. galli AF00
    |         Trispeleognathini WL09
    |--Astrida parrae AF00
    `--+--Psyttaboidaia AF00
       |--+--Speleognathus Womersley 1936 AF00
       |  |    |--*S. australis Womersley 1936 AF00
       |  |    `--S. schoutedeni Fain 1955 AF00, H98 (see below for synonymy)
       |  `--Neoboydaia Fain 1958 AF00, H98
       |       |--N. colymbiformi Clark 1964 H98
       |       `--N. philomachi (Fain 1956) [=Boydaia philomachi] H98
       `--Boydaia Womersley 1953 AF00, H98 [Boydaiini]
            |--B. aratingae AF00
            |--B. clavata AF00
            |--B. crassipes (Berlese & Trouessart 1889) [=Ereynetes crassipes, Riccardoella crassipes] H98
            |--B. falconis Fain 1956 H98
            |--B. hirundoae Fain 1956 H98
            |--B. maluri Domrow 1969 H98
            |--B. myzomelae Domrow 1969 H98
            |--B. nigra WL09
            |--B. podargi Fain & Lukoschus 1979 H98
            |--B. spatulata Fain 1955 [=B. sturni var. spatulata] H98
            |--B. sturnellae WL09
            |--B. sturni (Boyd 1948) [=Speleognathus sturni] H98
            |--B. trochila A91
            |--B. zosteropis Fain 1963 H98
            `--B. zumpti AF00

Speleognathus schoutedeni Fain 1955 AF00, H98 [=Neoboydaia schoutedeni H98, Ophthalmognathus schoutedeni H98, Trispeleognathus schoutedeni H98]

*Type species of generic name indicated

REFERENCES

[A91] André, H. M. 1991. The Tydeoidea: a striking exception to the Oudemans-Grandjean rule. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 293–296. SPB Academic Publishing: The Hague.

[AF00] André, H. M., & A. Fain. 2000. Phylogeny, ontogeny and adaptive radiation in the superfamily Tydeoidea (Acari: Actinedida), with a reappraisal of morphological characters. Zoological Journal of the Linnean Society 130: 405–448.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

Protohydrachnidia

Rhyncholimnochares sp., copyright Ray Fisher.

Belongs within: Hydrachnidia.
Contains: Eylais.

The Protohydrachnidia are a clade of water mites in which the lateral eyes lack muscles, the last pair of idiosomal dorsoventral muscles are arranged in a median position, and the ejaculatory complex has the distal-sclerite arms and apical setae reduced.

<==Protohydrachnidia
    |--Hydrovolzioidea TBG00
    |    |--Hydrovolziidae [Hydrovolziinae] WWO01
    |    |    |--Stygovolzia Imamura 1957 TBG00
    |    |    |    `--S. uenoi (Imamura 1957) TBG00
    |    |    |--Hydrovolziella Viets 1935 TBG00
    |    |    |    `--*H. lata (Walter 1935) [=Hydrovolzia lata] TBG00
    |    |    `--Hydrovolzia Thor 1905 TBG00
    |    |         |--H. baderi G96
    |    |         |--H. cancellata Walter 1906 TBG00
    |    |         |--H. gerhardi TBG00
    |    |         |--H. japonica Imamura 1961 TBG00
    |    |         |--H. marshallae WL09
    |    |         |--H. persica Bader & Sepasgozarian 1980 TBG00
    |    |         `--H. placophora (Monti 1905) TBG00
    |    `--Acherontacaridae [Acherontacarinae] TBG00
    |         |--Bharatavolzia Cook 1967 TBG00
    |         |    |--B. (Bharatavolzia) muscicola Cook 1967 S86a
    |         |    `--B. (Bharatavolziella Cook 1967) TBG00
    |         |         |--B. (B.) cooki Schwoerbel & Sepasgozarian 1980 S86a
    |         |         `--B. (B.) pallida Cook 1967 S86a
    |         `--Acherontacarus Viets 1932 (see below for synonymy) TBG00
    |              |--*A. halacaroides Viets 1932 TBG00
    |              |--A. bicornis Cook 1974 TBG00
    |              |--A. cedro Lundblad 1962 TBG00
    |              |--A. cicolanii GS96
    |              |--A. dividuus TBG00
    |              |--A. fonticolus Viets 1934 TBG00
    |              |--A. rutilans Angelier 1951 TBG00
    |              |--A. tuberculatus TBG00
    |              `--A. vietsi Angelier 1951 TBG00
    `--Eylaoidea WL09
         |  i. s.: Apheviderulix [Apheviderulicidae] WL09
         |           |--A. santana WL09
         |           `--A. welwitchioides WL09
         |--Piersigiidae WWO01
         |    |--Stygolimnocharinae WL09
         |    |    |--Parawandesia Angelier 1951 WL09, S86b
         |    |    |    `--P. chappuisi Angelier 1951 S86b
         |    |    `--Stygolimnochares Cook 1967 H98
         |    |         |--S. australica Cook 1986 H98
         |    |         `--S. elongata Cook 1967 S86b
         |    `--Piersigiinae WL09
         |         |--Austrapiersigia montana WL09
         |         `--Piersigia WL09
         |              |--P. crusta WL09
         |              |--P. intermedia Wi91
         |              `--P. limnophila WL09
         `--+--Eylais Wo91
            `--Limnocharidae WWO01
                 |--Limnocharinae WL09
                 |    |--Limnochares Latreille 1796 H98
                 |    |    |--L. (Cyclothrix) americana WP99
                 |    |    |--L. aquatica (Linnaeus 1758) Wo91 [=Acarus aquaticus L02]
                 |    |    |--L. australica Lundblad 1941 H98
                 |    |    |--L. crinita WL09
                 |    |    `--L. holoserica [=Hydrachna holoserica] G20
                 |    `--Neolimnochares Lundblad 1937 WL09, RFF05
                 |         |--N. breviscuta Lundblad 1941 RFF05
                 |         |--N. kakadu Harvey 1990 H98
                 |         |--N. petrophila Lundblad 1937 RFF05
                 |         |--N. placophora Lundblad 1937 RFF05
                 |         `--N. pugiunculata Viets 1954 RFF05
                 `--Rhyncholimnocharinae WL09
                      |--Austrolimnochares womersleyi WL09
                      `--Rhyncholimnochares Lundblad 1936 RFF05
                           |  i. s.: R. kittitanniana WL09
                           |         R. womersleyi (Lundblad 1952) [=Paralimnochares womersleyi] H98
                           |--R. (Rhyncholimnochares) RFF05
                           |    |--R. (R.) andina Lundblad 1953 RFF05
                           |    |--R. (R.) expansiseta Cook 1980 RFF05
                           |    |--R. (R.) gracilirostris Lundblad 1953 RFF05
                           |    |--R. (R.) lamellipalpis Lundblad 1936 RFF05
                           |    |--R. (R.) longipalpis Lundblad 1953 RFF05
                           |    `--R. (R.) longiscuta Lundblad 1941 RFF05
                           `--R. (Paralimnochares Lundblad 1937) RFF05
                                |--R. (P.) dipersiai (Rosso de Ferradás 1975) RFF05
                                `--R. (P.) sursumhians (Lundblad 1937) RFF05

Acherontacarus Viets 1932 [incl. Acherontacarellus Lundblad 1962, Acherontacaropsis Cook 1967, Neoacherontacarus Bader 1989] TBG00

*Type species of generic name indicated

REFERENCES

[GS96] Gerecke, R., & A. Di Sabatino. 1996. Historical zoogeography and evolution of habitat preference in water mites (Hydracarina) of the central Mediterranean region. In: Mitchell, R., D. J. Horn, G. R. Needham & W. C. Welbourn (eds) Acarology IX vol. 1. Proceedings pp. 523–527. Ohio Biological Survey: Columbus (Ohio).

[G96] Gil, M. J. 1996. Water-mite (Hydracarina) larvae of some selected genera from Spain: a phylogenetic inference. In: Mitchell, R., D. J. Horn, G. R. Needham & W. C. Welbourn (eds) Acarology IX vol. 1. Proceedings pp. 543–546. Ohio Biological Survey: Columbus (Ohio).

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

[S86a] Schwoerbel, J. 1986a. Acari: Limnohalacaridae and Hydrovolziidae. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 643–647. E. J. Brill/Dr W. Backhuys: Leiden.

[S86b] Schwoerbel, J. 1986b. Acari: “Hydrachnellae”. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 652–696. E. J. Brill/Dr W. Backhuys: Leiden.

[TBG00] Tuzovskij, P., D. Benfatti & R. Gerecke. 2000. The water mite family Acherontacaridae Cook, 1967 nov. stat. and diagnosis of the superfamily Hydrovolzioidea Thor, 1905 (Acariformes, Hydrachnidia). Acarologia 41: 451–473.

[WL09] Walter, D. E., E. E. Lindquist, I. M. Smith, D. R. Cook & G. W. Krantz. 2009. Order Trombidiformes. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 233–420. Texas Tech University Press.

[WP99] Walter, D. E., & H. C. Proctor. 1999. Mites: Ecology, Evolution and Behaviour. CABI Publishing: Wallingford (UK).

[Wi91] Witte, H. 1991. The phylogenetic relationships within the Parasitengonae. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 171–182. SPB Academic Publishing: The Hague.

[Wo91] Wohltmann, A. 1991. Nutrition and respiratory rate of water mites (Actinedida: Hydrachnidia) from a phylogenetic point of view. In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 543–548. SPB Academic Publishing: The Hague.

[WWO01] Wohltmann, A., H. Witte & R. Olomski. 2001. Organismal patterns causing high potential for adaptive radiation in Parasitengonae (Acari: Prostigmata). In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10^th International Congress pp. 83–99. CSIRO Publishing: Melbourne.

Wandesia

Wandesia thori, copyright Faculty of Science and Mathematics, Podgorica.

Belongs within: Hydryphantidae.

Wandesia is a genus of elongate-bodied water mites with widely separated coxae and lacking dorsal plates or genital flaps.

<==Wandesia Schechtel 1912 RFF05
    |  i. s.: W. cavecerberum GS96
    |         W. saginata GS96
    |         W. troma Cook 1986 H98
    |--W. (Wandesia) S86
    |    |--W. (W.) centensis Habeeb 1973 S86
    |    |--W. (W.) extendens Cook 1980 S86
    |    |--W. (W.) racovitzai Gledhill 1970 S86
    |    `--W. (W.) thori Schechtel 1912 S86
    |--W. (Allowandesia Schwoerbel 1961) S86
    |    `--W. (A.) japonica Imamura 1956 S86
    |--W. (Mesowandesia André & Naudo 1962) S86
    |    `--W. (M.) riparia Habeeb 1954 S86
    |--W. (Partnuniella Viets 1938) RFF05
    |    |--W. (P.) chechoi Cook 1988 [=W. (Pseudowandesia) chechoi] RFF05
    |    |--W. (P.) cubana Orghidan & Georgesco 1977 S86
    |    |--W. (P.) lethaea (Besch 1964) RFF05
    |    |--W. (P.) stalagmophila Lundblad 1941 RFF05
    |    |--W. (P.) thermalis (Viets 1938) RFF05
    |    `--W. (P.) walteri Lundblad 1937 RFF05
    `--W. (Pseudowandesia Habeeb 1958) RFF05
         |--W. (P.) andiana Cook 1980 RFF05
         |--W. (P.) approximata Cook 1967 S86
         |--W. (P.) gaspensis Habeeb 1955 S86
         |--W. (P.) glareosa Weigmann & Schminke 1970 S86
         |--W. (P.) stygophila Szalay 1944 S86
         |--W. (P.) tanasachii Orgidan & Georgescu 1977 S86
         `--W. (P.) vermiformis Cook 1967 S86

*Type species of generic name indicated

REFERENCES

[GS96] Gerecke, R., & A. Di Sabatino. 1996. Historical zoogeography and evolution of habitat preference in water mites (Hydracarina) of the central Mediterranean region. In: Mitchell, R., D. J. Horn, G. R. Needham & W. C. Welbourn (eds) Acarology IX vol. 1. Proceedings pp. 523–527. Ohio Biological Survey: Columbus (Ohio).

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[RFF05] Rosso de Ferradás, B., & H. R. Fernández. 2005. Elenco y biogeografía de los ácaros acuáticos (Acari, Parasitengona, Hydrachnidia) de Sudamérica. Graellsia 61 (2): 181–224.

[S86] Schwoerbel, J. 1986. Acari: “Hydrachnellae”. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 652–696. E. J. Brill/Dr W. Backhuys: Leiden.