Calycoceras

Calycoceras spinosum, from here.

Belongs within: Acanthoceratoidea.

Calycoceras is a genus of evolute ammonites with depressed whorls known from the Upper Cretaceous (Cenomanian) (Wright et al. 1996).

Characters (from Wright et al. 1996): Rather evolute, with whorl section depressed and subcircular, oval, polygonal, or subquadrate; ribs strong, generally straight, continuous over rounded or flat but not concave venter; on early whorls at least umbilical, ventrolateral, siphonal, and, in most specimens, midlateral tubercles; in multituberculate forms umbilical tubercle more prominent than midlateral; tubercles may disappear with age and may or may not be rejuvenated on last part of shell. Marked dimorphism in size apparently general.

<==Calycoceras
    |--C. andranovaritelense K79
    |--C. asiaticum K79
    |--C. boulei K79
    |--C. canitaurinum K79
    |--C. hourcqi K79
    |--C. jeanneti K79
    |--C. latelobatum K79
    |--C. leonense K79
    |--C. (Calycoceras) naviculare (Mantell 1822) [=Acanthoceras naviculare] KK08
    |--C. newboldi K79
    |--C. orientale K79
    |--C. spinosum K79
    |--C. stoliczkai K79
    `--C. tarrantense K79

*Type species of generic name indicated

REFERENCES

[K79] Kauffman, E. G. 1979. Cretaceous. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A418–A487. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[KK08] Kennedy, W. J., & H. C. Klinger. 2008. Hypermorphosis in Salaziceras, a Cretaceous ammonite, from Madagascar. African Natural History 4: 113–116.

Wright, C. W., J. H. Callomon & M. K. Howarth. 1996. Treatise on Invertebrate Paleontology pt L. Mollusca 4, Revised vol. 4. Cretaceous Ammonoidea. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

Acanthoceratoidea

Forbesiceras largilliertianum, from here.

Belongs within: Brancoceratoidea.
Contains: Tegoceras, Calycoceras.

The Acanthoceratoidea are a Cretaceous group of usually strongly ribbed ammonites, with a tendency to develop prominent tubercles.

<==Acanthoceratoidea [Acanthocerataceae, Lyellicerataceae, Lyelliceratoidea] KK08a
    |  i. s.: Barroisiceras de Grossouvre 1894 EK02
    |           |--B. anilahyense K79
    |           `--B. haberfellneri K79
    |         Subprionocyclus TLY03
    |           |--S. neptuni TLY03
    |           `--S. normalis K79
    |--Forbesiceras Kossmat 1897 KK08c (see below for synonymy)
    |    |--*F. largilliertianum (d’Orbigny 1841) (see below for synonymy) KK08c
    |    |--F. baylissi Wright & Kennedy 1984 KK08c
    |    |--F. beaumontianum (d’Orbigny 1841) [=Ammonites beaumontianus] KK08c
    |    |--F. bicarinatum Szász 1976 KK08c
    |    |--F. brundrettei (Young 1958) KK08c
    |    |--F. chevillei (Pictet & Renevier 1866) (see below for synonymy) KK08c
    |    |--F. conlini Stephenson 1953 KK08c
    |    |--F. douvillei (Collignon 1929) [=Pulchellia (Neopulchellia) douvillei] KK08c
    |    |--F. flicki Pervinquière 1907 KK08c
    |    |--F. furnishi Collignon 1964 KK08c
    |    |--F. gignouxi (Collignon 1929) [=Pulchellia (*Neopulchellia) gignouxi] KK08c
    |    |--F. mikasaense Matsumoto 1986 KK08c
    |    |--F. obtectum (Sharpe 1853) KK08c
    |    |--F. pseudobtectum Collignon 1964 KK08c
    |    |--F. subobtectum (Stoliczka 1864) [=Ammonites subobtectus] KK08c
    |    |--F. tenuipunctatum Collignon 1964 KK08c
    |    `--F. varicostatum Zaborski 1990 KK08c
    |--Lyelliceratidae KK08a
    |    |  i. s.: Budaiceras K79
    |    |--Tegoceras KK08a
    |    `--+--Stoliczkaiinae [Neophlycticeratinae] KK08a
    |       |    |--Neophlycticeras madagascariensis K79
    |       |    `--Stoliczkaia K79
    |       |         |--S. africana K79
    |       |         |--S. clavigera K79
    |       |         |--S. crassa K79
    |       |         |--S. dispar K79
    |       |         |--S. grandidieri K79
    |       |         `--S. notha K79
    |       |--Salaziceras Breistroffer 1936 KK08b [Flickiidae KK08a, Salaziceratinae]
    |       |    |--*S. salazacense (Hébert & Munier-Chalmas 1875) [=Ammonites salazacensis] KK08b
    |       |    `--S. lemoinei Kennedy & Klinger 2008 KK08b
    |       `--Acanthoceratidae [Acanthoceratinae] KK08a
    |            |--Dunveganoceras K79
    |            |--Calycoceras K79
    |            |--Eucalycoceras K79
    |            |--Kanabiceras K79
    |            `--Acanthoceras KK08c
    |                 |--A. alvaradoense K79
    |                 |--A. amphibolum K79
    |                 |--A. amudariensis A03
    |                 |--A. beilense K79
    |                 |--A. jukesbrownei K79
    |                 |--A. muldoonense K79
    |                 |--A. rhotomagense KK08c
    |                 `--A. vicinale K79
    `--Collignoniceratidae K79
         |--Peroniceratinae K79
         |--Collignoniceratinae K79
         |    |--Reesidites minimus K79, TLY03
         |    |--Prionocyclus K79
         |    `--Collignoniceras Breistroffer 1947 EK02
         |         |--C. teshioensis K79
         |         `--C. woollgari K79
         `--Texanitinae K79
              |--Submortoniceras K79
              `--Texanites Spath 1932 EK02
                   |--T. americanus K79
                   |--T. hispanica K79
                   |--T. hourcqi K79
                   |--T. kawasakii K79
                   |--T. oliveti K79
                   |--T. pseudotexanus K79
                   |--T. shiloensis K79
                   |--T. stangeri K79
                   |    |--T. s. stangeri K79
                   |    `--T. s. densicostus K79
                   `--T. texanus K79
                        |--T. t. texanus K79
                        `--T. t. gallicus K79

Forbesiceras Kossmat 1897 KK08c [=Cenomanites Haug 1898 KK08c, Discoceras Kossmat 1895 non Barrande 1867 KK08c; incl. Neopulchellia Collignon 1929 KK08c; Forbesiceratidae, Forbesiceratinae KK08a]

Forbesiceras chevillei (Pictet & Renevier 1866) [=Ammonites chevillei; incl. F. almerae (Mallada 1891), F. clarki Collignon 1964, F. nodosum Crick 1907, F. sculptum Crick 1907] KK08c

*Forbesiceras largilliertianum (d’Orbigny 1841) [=Ammonites largilliertianus, *Cenomanites largilliertianus, *Discoceras largilliertianum] KK08c

*Type species of generic name indicated

REFERENCES

[A03] Alifanov, V. R. 2003. Two new dinosaurs of the infraorder Neoceratopsia (Ornithischia) from the Upper Cretaceous of the Nemegt Depression, Mongolian People’s Republic. Paleontologicheskii Zhurnal 2003 (5): 77–88 (transl. Paleontological Journal 37 (5): 524–534.)

[EK02] Engeser, T., & H. Keupp. 2002. Phylogeny of the aptychi-possessing Neoammonoidea (Aptychophora nov., Cephalopoda). Lethaia 35: 79–96.

[K79] Kauffman, E. G. 1979. Cretaceous. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A418–A487. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[KK08a] Kennedy, W. J., & H. C. Klinger. 2008a. Cretaceous faunas from Zululand and Natal, South Africa. The ammonite subfamily Lyelliceratinae Spath, 1921. African Natural History 4: 57–111.

[KK08b] Kennedy, W. J., & H. C. Klinger. 2008b. Hypermorphosis in Salaziceras, a Cretaceous ammonite, from Madagascar. African Natural History 4: 113–116.

[KK08c] Kennedy, W. J., & H. C. Klinger. 2008c. Cretaceous faunas from Zululand and Natal, South Africa. The ammonite family Forbesiceratidae Wright, 1952. African Natural History 4: 117–130.

[TLY03] Tanabe, K., N. H. Landman & Y. Yoshioka. 2003. Intra- and interspecific variation in the early internal shell features of some Cretaceous ammonoids. Journal of Paleontology 77 (5): 876–887.

Brancoceratoidea

Prolyelliceras ulrichi, from here.

Belongs within: Aptychophora.
Contains: Acanthoceratoidea.

The Brancoceratoidea is a Cretaceous group of ammonites with a more or less evolute conch bearing strong ribs.

<==Brancoceratoidea [Brancocerataceae] KK08
    |--Leymeriella [Leymeriellidae] KK08
    |    |--L. regularis K79
    |    |--L. schrammeni K79
    |    `--L. tardefurcata KK08
    `--+--Acanthoceratoidea KK08
       `--Brancoceratinae [Brancoceratidae] KK08
            |--Brancoceras Hyatt 1884 KK08, H84
            |    |--*B. ixion [=Goniatites ixion] H84
            |    |--B. alternatum Kennedy 2004 KK08
            |    |--B. lineare [=Goniatites linearis] H84
            |    |--B. ovatum [=Goniatites ovatus] H84
            |    |--B. rotatorium [=Goniatites rotatorius] H84
            |    |--B. subsulcatum [=Goniatites subsulcatus] H84
            |    |--B. sulcatum [=Goniatites sulcatus] H84
            |    |--B. umbilicatum H84
            |    `--B. ungeri [=Goniatites ungeri] H84
            `--+--Parabrancoceras Breistroffer 1951 KK08
               |    `--P. dromense KK08
               `--Prolyelliceras Spath 1930 [incl. Ralphimlayites Etayo-Serna 1979] KK08
                    |  i. s.: P. alternatum (Robert 2002) (see below for synonymy) KK08
                    |         ‘Ralphimlayites’ apuloense Etayo-Serna 1979 KK08
                    |         P. flandrini (Dubourdieu 1953) [=Lyelliceras flandrini] KK08
                    |         P. radenaci (Pervinquière 1907) [=Prionotropis radenaci] KK08
                    |--P. cotteri (Spath 1930) [=Lyelliceras cotteri] KK08
                    |--P. lobatum (Riedel 1938) KK08
                    `--+--P. prorsocurvatum (Gerhardt 1897) (see below for synonymy) KK08
                       `--+--+--Eubrancoceras aegoceratoides (Steinmann 1881) KK08
                          |  `--P. mathewsi (Knechtel 1947) [=Lyelliceras mathewsi, Ralphimlayites mathewsi] KK08
                          `--+--‘Lyelliceras’ isaclei KK08
                             `--P. ulrichi (Knechtel 1947) [=Lyelliceras ulrichi, Ralphimlayites ulrichi] KK08

Prolyelliceras alternatum (Robert 2002) [=Ralphimlayites alternatum, R. alternum Robert 2001 (n. n.)] KK08

Prolyelliceras prorsocurvatum (Gerhardt 1897) [=Acanthoceras prorsocurvatum, *Ralphimlayites prorsocurvatum; incl. *Prolyelliceras peruvianum Spath 1930] KK08

*Type species of generic name indicated

REFERENCES

[H84] Hyatt, A. 1883–1884. Genera of fossil cephalopods. Boston Soc. Nat. History, Proc. 22: 253–338.

[K79] Kauffman, E. G. 1979. Cretaceous. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A418–A487. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[KK08] Kennedy, W. J., & H. C. Klinger. 2008. Cretaceous faunas from Zululand and Natal, South Africa. The ammonite subfamily Lyelliceratinae Spath, 1921. African Natural History 4: 57–111.

Ancyloceratidae

Australiceras sp., in the Houston Museum of Natural Science.

Belongs within: Aptychophora.

The Ancyloceratidae are a group of ammonites known from the Lower Cretaceous. They are commonly heteromorphic, though some derived forms reverted to a more standard tight coil.

Characters (from Wright et al. 1996): Either coiled in a loose or tight, more or less equiangular spiral (criocone) or with an initial spire followed by curved or straight shaft and terminal hook (ancyloceratoid); sometimes with phylogenetic tendency to tighter, normal ammonitic coiling. Whorl section ranging from circular to square or polygonal; tending to develop strong, periodic ribs with umbilical, lateral, and ventrolateral spines, separated by few to many, fine, untuberculate ribs. Ornament rarely constant through growth. Probably strongly dimorphic throughout, but presumed microconchs rarely matched with macroconchs. Sutures florid, with trifid lobes or simplifying.

<==Ancyloceratidae [Ancylocerataceae] EK02
    |  i. s.: Parancyloceras bidentatum W69, K79
    |         Menuthiocrioceras W69
    |         Pedioceras W69
    |--Lithancylus W69
    `--+--+--Colchidites shaoriensis W69
       |  `--Turkmeniceras turkmenicum W69
       |--Ancyloceras d’Orbigny 1842 W69, EK02 [Ancyloceratinae]
       |    |--A. durrelli K79
       |    |--A. flindersi F71
       |    |--A. innexum K79
       |    `--A. vandenheckii K79
       |--+--Epancyloceras W69
       |  `--Ammonitoceras W69
       |       |--A. tovilense W69
       |       `--A. ucetiae W69
       `--+--Tropaeum W69
          |    |  i. s.: T. arcticum K79
          |    |         T. australe K79
          |    |         T. jacki K79
          |    |         T. leptum (Etheridge 1909) [=Crioceras leptum] F71
          |    |--T. hillsi W69
          |    `--+--T. bowerbanki W69
          |       `--T. subarcticum W69
          `--Australiceras W69
               |  i. s.: A. gracile F71
               |         A. irregulare [=Crioceras irregulare] F71
               |         A. lampros (Etheridge 1909) [=Crioceras lampros] F71
               |         A. robustum Whitehouse 1926 F71
               |--A. gigas W69
               `--+--A. jacki W69 [=Crioceras jackii F71; incl. Tropaeum undatum F71]
                  `--A. tuberculatum W69

*Type species of generic name indicated

REFERENCES

[EK02] Engeser, T., & H. Keupp. 2002. Phylogeny of the aptychi-possessing Neoammonoidea (Aptychophora nov., Cephalopoda). Lethaia 35: 79–96.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[K79] Kauffman, E. G. 1979. Cretaceous. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A418–A487. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[W69] Wiedmann, J. 1969. The heteromorphs and ammonoid extinction. Biological Reviews 44: 563–602.

Wright, C. W., J. H. Callomon & M. K. Howarth. 1996. Treatise on Invertebrate Paleontology pt L. Mollusca 4, Revised vol. 4. Cretaceous Ammonoidea. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

Marathonitidae

Peritrochia, from here.

Belongs within: Ammonoidea.

The Marathonitidae are a group of more or less globular ammonoids known from the Upper Carboniferous to the Upper Permian (Miller et al. 1957).

Characters (from Miller et al. 1957, as Marathonitinae): Conch subglobular to globular. Lateral lobes of sutures tending to be trifid.

<==Marathonitidae RR79
    |--Marathonites smithi P68
    |--Kargalites RR79
    |--Tabantalites RR79
    |--Demarezites RR79
    |--Hyattoceras subgeinitzi RR79
    |--Pseudovidrioceras RR79
    `--Peritrochia RR79
         |--P. erebus Girty 1908 P68
         |--P. novomexicana Miller & Parizek 1948 P68
         `--P. sellardsi P68

*Type species of generic name indicated

REFERENCES

Miller, A. K., W. M. Furnish & O. H. Schindewolf. 1957. Paleozoic Ammonoidea. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt L. Mollusca 4. Cephalopoda: Ammonoidea pp. L11–L79. Geological Society of America, and University of Kansas Press.

[P68] Purnell, L. R. 1968. Catalog of the type specimens of invertebrate fossils. Part I: Paleozoic Cephalopoda. United States National Museum Bulletin 262: 1–198.

[RR79] Ross, C. A., & J. R. P. Ross. 1979. Permian. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A291–A350. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

Popanoceratidae

Popanoceras bowmani, from here.

Belongs within: Ammonoidea.

The Popanoceratidae are a group of discoidal to globular ammonoids known from the Upper Carboniferous to the Upper Permian (Miller et al. 1957).

Characters (from Miller et al. 1957): Conch discoidal to globular, umbilicus small. Sutures ceratitic to ammonitic, forming series of subequal lateral lobes.

<==Popanoceratidae [Popanoceratoidea] SLE01
    |--Protopopanoceras RR79
    |--Propopanoceras RR79
    |--Mongoloceras RR79
    |--Tauroceras Toumanskaya 1938 SLE01
    |    `--*T. scrobiculatum (Gemmellaro 1887) [=Popanoceras scrobiculatum] SLE01
    `--Popanoceras Hyatt 1884 [incl. Pamiropopanoceras Leonova 1989] ZL04
         |--*P. sobolewskyanus (de Verneuil 1845) (see below for synonymy) ZL04
         |--P. antiquum [=Arcestes antiquus] H84
         |--P. bowmani RR79
         |--P. ganti Smith 1903 P68
         |--‘*Pamiropopanoceras’ meridionale Leonova 1989 ZL04
         `--P. walcotti White 1889 P68

*Popanoceras sobolewskyanus (de Verneuil 1845) [=Goniatites sobolewskyanus, P. soboleskianum (l. c.); incl. Goniatites kiangianus de Verneuil 1845, P. kiangianum, G. koninckianus de Verneuil 1845, P. koninckianum] ZL04

*Type species of generic name indicated

REFERENCES

[H84] Hyatt, A. 1883–1884. Genera of fossil cephalopods. Boston Soc. Nat. History, Proc. 22: 253–338.

Miller, A. K., W. M. Furnish & O. H. Schindewolf. 1957. Paleozoic Ammonoidea. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt L. Mollusca 4. Cephalopoda: Ammonoidea pp. L11–L79. Geological Society of America, and University of Kansas Press.

[P68] Purnell, L. R. 1968. Catalog of the type specimens of invertebrate fossils. Part I: Paleozoic Cephalopoda. United States National Museum Bulletin 262: 1–198.

[RR79] Ross, C. A., & J. R. P. Ross. 1979. Permian. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A291–A350. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[SLE01] Sone, M., M. S. Leman & M. Ehino. 2001. Middle Permian cephalopods from central Peninsular Malaysia: implications for faunal migration through the southern Tethys. Journal of Asian Earth Sciences 19: 805–814.

[ZL04] Zhou, Z., & M. Liengjarern. 2004. Lower Permian perrinitid ammonoid faunas from Thailand. Journal of Paleontology 78: 317–339.

Adrianitidae

Adrianites, from here.

Belongs within: Ammonoidea.

The Adrianitidae are a group of discoidal to globular ammonoids known from the Upper Carboniferous to the Middle Permian (Miller et al. 1957).

Characters (from Miller et al. 1957): Discoidal to globular conchs with variable umbilici. Goniatitic suture with ten to thirty lobes that tend to be subequal.

<==Adrianitidae [Adrianitoidea] ZL04
    |  i. s.: Crimites RR79
    |         Texoceras texanum RR79, P68
    |         Doryceras spinosum RR79
    |         Aricoceras RR79
    |         Palermites RR79
    |         Epadrianites timorense RR79
    |         Basleoceras RR79
    |         Pseudoagathiceras RR79
    |         Sizilites RR79
    `--Adrianitinae ZL04
         |--Adrianites SLE01
         |    |--A. adamsi P68
         |    |--A. elegans SLE01
         |    `--A. newelli Miller & Furnish 1940 P68
         `--Neocrimites Ruzhencev 1940 [incl. Metacrimites Ruzhencev 1950] ZL04
              |--*N. fredericksi (Emel’yancev 1929) [=Adrianites fredericksi] ZL04
              |--N. meridionalis [=Adrianites (Neocrimites) meridionalis] F71
              |--N. nalivkini (Toumanskaya 1949) [=Adrianites nalivkini, Waagenina nalivkini] ZL04
              |--N. newelli RR79
              `--N. plummeri RR79

*Type species of generic name indicated

REFERENCES

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

Miller, A. K., W. M. Furnish & O. H. Schindewolf. 1957. Paleozoic Ammonoidea. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt L. Mollusca 4. Cephalopoda: Ammonoidea pp. L11–L79. Geological Society of America, and University of Kansas Press.

[P68] Purnell, L. R. 1968. Catalog of the type specimens of invertebrate fossils. Part I: Paleozoic Cephalopoda. United States National Museum Bulletin 262: 1–198.

[RR79] Ross, C. A., & J. R. P. Ross. 1979. Permian. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A291–A350. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[SLE01] Sone, M., M. S. Leman & M. Ehino. 2001. Middle Permian cephalopods from central Peninsular Malaysia: implications for faunal migration through the southern Tethys. Journal of Asian Earth Sciences 19: 805–814.

[ZL04] Zhou, Z., & M. Liengjarern. 2004. Lower Permian perrinitid ammonoid faunas from Thailand. Journal of Paleontology 78: 317–339.