Progonyleptoidellinae

Progonyleptoidellus striatus, copyright Jan van Bodegraven.

Belongs within: Gonyleptidae.

The Progonyleptoidellinae are a group of gonyleptid harvestmen with the fourth coxa hidden in dorsal view, and the pedipalp long and slender.

Characters (from Soares & Soares 1985): Coxa IV (in dorsal view) with the apex only visible outside the dorsal shield, or, if visible to a greater degree, with the visible section very narrow. Dorsal scutum with four sulci, areas I and II joined or not by median longitudinal sulcus. Eyes set in a common ocularium. Palps slightly longer than the body, femur cylindrical and slender or somewhat thick and compressed, unarmed or with an internal apical spine. Tarsi with large numbers of segments, tarsus III with at least three segments more than tarsus I (usually with about twice as many segments as tarsus I), rarely with one or two more segments, tarsus I with more than 6 segments, or, if with six, with three segments less than tarsus III. Distitarsus II with four segments. Tarsus III and IV with pseudonychium, with two smooth claws, without scopula or with rudimentary scopula.

<==Progonyleptoidellinae
    |--Iporangaia Mello-Leitão 1935 K03
    |    `--I. pustulosa Mello-Leitão 1935 K03
    |--Cadeadoius Mello-Leitão 1936 DG09
    |    `--C. niger (Mello-Leitão 1935) (see below for synonymy) K03
    |--Heliella Soares 1945 K03
    |    `--*H. singularis Soares 1945 K03
    |--Leptocnemus Koch 1839 K03, KA-Z11 [=Leptocnema Koch 1839 KA-Z11]
    |    `--*L. sulphureus (Perty 1832) K03, DG09 (see below for synonymy)
    |--Deltigalus Roewer 1931 K03
    |    `--D. curvispina (Perty 1833) (see below for synonymy) K03
    |--Mitopernoides Soares 1945 K03
    |    `--*M. variabilis Soares 1945 [=Mitoperna variabilis] K03
    |--Iguapeia Mello-Leitão 1935 [incl. Angistripygus Roewer 1943, Langesia Soares 1945] K03
    |    |--*I. melanocephala Mello-Leitão 1935 (see below for synonymy) K03
    |    `--I. gengnageli (Soares & Soares 1949) [=Metagonyleptes gengnageli] K03
    |--Progonyleptoidellus Piza 1940 K03 (see below for synonymy)
    |    |--P. striatus (Roewer 1913) DG09 (see below for synonymy)
    |    |--P. fuscopictus (Soares 1942) [=*Laneius fuscopictus, Gonazula fuscopicta] K03
    |    `--P. orguensis (Soares & Soares 1954) [=Geraecormobius orguensis] K03
    |--Gonyleptoides Roewer 1913 K03
    |    |--*G. acanthoscelis (Bertkau 1880) (see below for synonymy) K03
    |    |--G. curvifemur Soares 1944 [=Geraecormobius curvifemur] K03
    |    `--G. marumbiensis Soares 1945 [=Geraecormobius marumbiensis] K03
    `--Moreiranula Roewer 1930 K03
         |--*M. moreirae (Mello-Leitão 1922) (see below for synonymy) K03
         |--M. mamillata (Soares 1944) [=Metagonyleptes mamillatus; incl. *Currala luteolimbata Piza 1946] K03
         `--M. picta (Soares & Soares 1982) [=Metagonyleptes pictus] K03

Cadeadoius niger (Mello-Leitão 1935) [=Gonyleptoides niger, Geraecormobius niger; incl. *C. pungens Mello-Leitão 1936] K03

Deltigalus curvispina (Perty 1833) [=Gonyleptes curvispina, Caelopygus curvispina, Coelopygus curvispina; incl. Ca. alter Roewer 1923, *Deltigalus bifrons Roewer 1931, Ca. kochii Roewer 1923, Coelopygus kochii] K03

Gonyleptoides acanthoscelis (Bertkau 1880) [=Ancistrotus acanthoscelis, Geraecormobius acanthoscelis, Gonyleptoides acathoscelis (l. c.)] K03

*Iguapeia melanocephala Mello-Leitão 1935 [incl. *Angistripygus patellaris Roewer 1943, *Langesia unica Soares 1945, A. unicus] K03

Leptocnemus sulphureus (Perty 1832) K03, DG09 [=Goniosoma sulphureum K03, Asarcus sulphureus K03, Leptocnema sulfurea (l. c.) K03, Leptocnema sulphurea K03, Leptocnemus sulfureus (l. c.) K03]

*Moreiranula moreirae (Mello-Leitão 1922) [=Gonyleptoides moreirae, Geraecormobius moreirae; incl. Moreiranula melanostoma Mello-Leitão 1931, Geraecormobius melanostoma, Ge. melanostomus, Gonyleptoides melanostoma, Go. melanostomus] K03

Progonyleptoidellus Piza 1940 K03 [=Progonyleptiodellus (l. c.) KA-Z11; incl. Laneius Soares 1942 K03, Piraquara Piza 1943 K03]

Progonyleptoidellus striatus (Roewer 1913) DG09 [=Ampheres striatus K03; incl. *P. androgynus Piza 1940 K03, Geraecormobius androgynus K03, Gonyleptoides androgynus K03, Cadeadoius atroluteus Roewer 1943 K03, Geraecormobius atroluteus K03, *Piraquara schubarti Piza 1943 K03, Ge. schubarti K03, Gonyleptoides schubarti K03]

*Type species of generic name indicated

REFERENCES

[DG09] DaSilva, M. B., & P. Gnaspini. 2009. A systematic revision of Goniosomatinae (Arachnida: Opiliones: Gonyleptidae), with a cladistic analysis and biogeographical notes. Invertebrate Systematics 23: 530–624.

[K03] Kury, A. B. 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Ibérica de Aracnología, volumen especial monográfico 1: 1–337.

[KA-Z11] Kury, A. B., & M. A. Alonso-Zarazaga. 2011. Addenda and corrigenda to the "Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones)". Zootaxa 3034: 47–68.

Soares, H. E. M., & B. A. M. Soares. 1985. Opera Opiliologica Varia XXII. Opiliones Gonyleptidae. Naturalia 10: 157–200.