Hernandariinae

Hernandaria scabricula, copyright Abel Pérez González.

Belongs within: Gonyleptidae.

The Hernandariinae are a group of densely granulate gonyleptid harvestmen, often carrying a coating of dirt as camouflage.

Characters (from DaSilva & Kury 2007): Dense granulation on whole body, camouflage of dirt, strong apophyses on anterior margin of dorsal scute, single opening of scent glands. Sexual dimorphism of leg IV present or absent. Sexual dimorphism of basitarsus I present or absent. Mesotergum with 3 or 4 areas separated by transverse grooves.

<==Hernandariinae [Apembolephaeninae, Apembolepheninae, Hernandariidae, Hernandaroidae]
    |  i. s.: Hernandarioides Pickard-Cambridge 1905 R23, K03 [incl. Kaluga Goodnight & Goodnight 1942 K03]
    |           `--*H. plana Pickard-Cambridge 1905 K03, KA-Z11, K03 (see below for synonymy)
    |         Ariaeus Sørensen 1932 DK07
    |           `--*A. tuberculatus Sørensen 1932 V05
    |--+--Multumbo Roewer 1927 DK07
    |  |    |--*M. terrenus Roewer 1927 DK07
    |  |    `--M. dimorphicus DaSilva & Kury 2007 DK07
    |  `--+--Piassagera Roewer 1928 DK07
    |     |    `--*P. brieni Roewer 1928 K03
    |     `--Pseudotrogulus Roewer 1932 DK07
    |          |--*P. telluris Roewer 1932 FP-R02
    |          |--P. funebris Firmo & Pinto-da-Rocha 2002 FP-R02
    |          `--P. mirim Kury 1992 FP-R02
    `--+--Acrogonyleptes Roewer 1917 DK07 (see below for synonymy)
       |    |--A. armatifrons (Roewer 1917) [=Metagonyleptes armatifrons; incl. *A. spinifrons Roewer 1917] K03
       |    |--A. curitibae (Soares 1943) (see below for synonymy) K03
       |    |--A. exochus (Mello-Leitão 1931) (see below for synonymy) K03
       |    `--A. unus (Mello-Leitão 1927) [=*Proweyhia una; incl. *Metaxundarava heterotypica Mello-Leitão 1927] K03
       `--Hernandaria Sørensen 1884 DK07 (see below for synonymy)
            |--*H. scabricula Sørensen 1884 (see below for synonymy) K03
            |--H. calcarata (Soares & Soares 1945) [=Apembolephaenus calcaratus] K03
            |--H. heloisiae (Soares 1945) (see below for synonymy) K03
            `--H. setulosa (Mello-Leitão 1933) (see below for synonymy) K03

Acrogonyleptes Roewer 1917 DK07 [incl. Acrogonyleptoides Mello-Leitão 1931 K03, Melloleitaniana Soares 1943 K03, Melloteitaniana (l. c.) K03, Metaxundarava Mello-Leitão 1927 K03, Proweyhia Mello-Leitão 1927 K03, Pseudoacrogonyleptoides Soares 1966 K03]

Acrogonyleptes curitibae (Soares 1943) [=*Melloleitaniana curitibae; incl. M. pectinifemur Soares & Soares 1947, M. riodareiensis Soares & Soares 1945, M. riodariensis (l. c.)] K03

Acrogonyleptes exochus (Mello-Leitão 1931) [=*Acrogonyleptoides exochus; incl. *Pseudoacrogonyleptoides granulatus Soares 1966, Acrogonyleptoides piraquarensis Soares & Soares 1946] K03

Hernandaria Sørensen 1884 DK07 [=Hernandria (l. c.) KA-Z11; incl. Apembolephaenus Holmberg 1909 K03, Apembolephenus (l. c.) K03, Paraproweyhia Soares & Soares 1947 K03, Proampycus Roewer 1917 K03]

Hernandaria heloisiae (Soares 1945) [=Proweyhia heloisiae; incl. *Paraproweyhia curitibae Soares & Soares 1947] K03

*Hernandaria scabricula Sørensen 1884 [incl. *Apembolephaenus jorgei Holmberg 1909, Apembolephenus (l. c.) jorgei, *Proampycus spinifrons Roewer 1917] K03

Hernandaria setulosa (Mello-Leitão 1933) [=Apembolephenus (l. c.) setulosus, Apembolephaenus setulosus, Apembolephenus setulosos (l. c.)] K03

*Hernandarioides plana Pickard-Cambridge 1905 K03, KA-Z11, K03 [=H. planus K03; incl. *Kaluga elongata Goodnight & Goodnight 1942 K03]

*Type species of generic name indicated

REFERENCES

[DK07] DaSilva, M. B., & A. B. Kury. 2007. A remarkable new species of Multumbo showing sexual dimorphism, with the transfer of Multumbo and Piassagera to the Hernandariinae (Opiliones, Gonyleptidae). Zootaxa 1558: 29–37.

[FP-R02] Firmo, C. L., & R. Pinto-da-Rocha. 2002. A new species of Pseudotrogulus Roewer and assignment of the genus to the Hernandariinae (Opiliones, Gonyleptidae). Journal of Arachnology 30: 173–176.

[K03] Kury, A. B. 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Ibérica de Aracnología, volumen especial monográfico 1: 1–337.

[KA-Z11] Kury, A. B., & M. A. Alonso-Zarazaga. 2011. Addenda and corrigenda to the "Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones)". Zootaxa 3034: 47–68.

[R23] Roewer, C.-F. 1923. Die Weberknechte der Erde: Systematische Bearbeitung der bisher bekannnten Opiliones. Gustav Fischer: Jena.

[V05] Vasconcelos, E. G. 2005. Notes on Geraecormobius clavifemur (Mello-Leitao, 1927) and description of Geraecormobius reitzi n. sp. (Arachnida: Opiliones: Gonyleptidae). Zootaxa 1088: 1–10.