Staffellidae

Diagram of axial section of Nankinella, from Loeblich & Tappan (1964).

Belongs within: Loeblichiidae.

The Staffellidae are a group of smaller fusulinids known from the Upper Carboniferous and Permian (Loeblich & Tappan 1964).

Characters (from Loeblich & Tappan 1964): Shell small, spherical to discoidal, with umbilicate or flush axial ends; septa strongly arched forward, closely spaced, and totally unfluted from end to end; tunnel singular, bordered by distinctly asymmetrical chomata except in terminal part of last volution; walls composed of upper and lower layers that are in turn underlain and overlain by secondary deposits in most forms, except for outer part of last volution.

<==Staffellidae [Staffellacea, Staffellinae, Staffellinidae, Staffellininae]
    |--Nummulostegina Schubert 1907 LT64
    |    `--*N. velebitana Schubert 1908 LT64
    |--Pisolina Lee 1933 LT64
    |    `--*P. excessa Lee 1933 LT64
    |--Sphaerulina Lee 1933 LT64
    |    |--*S. crassispira Lee 1933 LT64
    |    `--S. ogbinensis K04
    |--Staffella Ozawa 1925 LT64
    |    |--*S. sphaerica (Abich 1859) [=Fusulina sphaerica, Fusulinella sphaerica] LT64
    |    |--S. expansa LT64
    |    `--S. moellerana Thompson 1935 LT64
    `--Nankinella Lee 1933 [incl. Hayasakaina Fujimoto & Kawada 1953] LT64
         |--*N. discoides (Lee 1931) [=Staffella discoides] LT64
         |--N. gigas C40
         |--N. kotakiensis (Fujimoto & Kawada 1953) [=*Hayasakaina kotakiensis] LT64
         |--N. minor PE02
         `--N. plummeri LT64

*Type species of generic name indicated

REFERENCES

[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3^rd ed. Harvard University Press: Cambridge (Massachusetts).

[K04] Kobayashi, F. 2004. Late Permian foraminifers from the Limestone Block in the Southern Chichibu Terrane of west Shikoku, SW Japan. Journal of Paleontology 78 (1): 62–70.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[PE02] Pan, H.-Z., & D. H. Erwin. 2002. Gastropods from the Permian of Guangxi and Yunnan Provinces, south China. Journal of Paleontology 76 (Memoir 56): 1–49.