Liochthonius

Liochthonius proprinquus, from here.

Belongs within: Enarthronota.

Liochthonius is an cosmopolitan genus of unornamented brachychthoniid mites in which the adanal plates are fused to form a U-shaped plate surrounding the anal plate (Balogh & Balogh 1992).

Characters (from Balogh & Balogh 1992): Notogaster with two transverse sutures and three shields, no setae arising within sutures; without prominent ornamentation. Shield Na bearing two rows of setae (c and d); seta d2 originating more medially than longitudinal row cp-e2. Pleural region with suprapleural plates absent or adnate to their shields and forming lobes. Epimeral setal formula 3-1-3-4. Genital plates with seven pairs of setae. Peranal plate present. Adanal plates fused posteriorly, forming U-shaped plate. Seta ad2 hypertrophied, widened, blade-like.

<==Liochthonius Hammen 1959 S04
    |--L. (Liochthonius) S04
    |    |--L. (L.) brevis (Michael 1888) (see below for synonymy) S04
    |    |--L. (L.) africanus Mahunka 1983 S04
    |    |--L. (L.) alius Chinone 1978 [=L. plumosus alius; incl. L. plumosus Chinone & Aoki 1972 non Mahunka 1969] S04
    |    |--L. (L.) alpestris (Forsslund 1958) [=Brachychthonius alpestris] S04
    |    |--L. (L.) altimonticola (Hammer 1958) [=Brachychthonius altimonticola] S04
    |    |--L. (L.) altus (Hammer 1958) [=Brachychthonius altus] S04
    |    |--L. (L.) andinus (Hammer 1958) [=Brachychthonius andinus] S04
    |    |--L. (L.) andrewi Evison 1981 S04
    |    |--L. (L.) approximatus (Jacot 1939) [=Brachychthonius approximatus] S04
    |    |--L. (L.) asotthalomensis Marshall, Reeves & Norton 1987 (see below for synonymy) S04
    |    |--L. (L.) asper Chinone 1978 S04
    |    |--L. (L.) attenuatus (Jacot 1938) [=Brachychthonius attenuatus] S04
    |    |--L. (L.) australis Covarrubias 1968 S04
    |    |--L. (L.) bifurcatus (Jacot 1936) [=Brachychthonius bifurcatus] S04
    |    |--L. (L.) breviseta (Hammer 1958) [=Brachychthonius breviseta] S04
    |    |--L. (L.) clavatus (Forsslund 1942) [=Brachychthonius clavatus] S04
    |    |--L. (L.) crassus Chinone 1978 [=L. alpestris crassus] S04
    |    |--L. (L.) dilutus Moritz 1976 S04
    |    |--L. (L.) evansi (Forsslund 1958) [=Brachychthonius evansi; incl. L. forsslundi levis Chinone 1974] S04
    |    |--L. (L.) fimbriatissimus Hammer 1962 [incl. Brachychthonius fimbriatus Hammer 1958 non Jacot 1936] S04
    |    |--L. (L.) fimbriatus (Jacot 1936) [=Brachychthonius fimbriatus] S04
    |    |--L. (L.) forsslundi (Hammer 1952) [=Brachychthonius forsslundi] S04
    |    |--L. (L.) furcillatus (Willmann 1942) [=Brachychthonius furcillatus; incl. B. ensifer Strenzke 1951] S04
    |    |--L. (L.) galba Chinone 1978 S04
    |    |--L. (L.) gisini (Schweizer 1948) [=Brachychthonius gisini] S04
    |    |--L. (L.) horridus (Sellnick 1928) [=Brachychthonius horridus] S04
    |    |--L. (L.) hystricinus (Forsslund 1942) [=Brachychthonius hystricinus; incl. B. ocellatus Hammer 1952] S04
    |    |--L. (L.) idem Hammer 1966 S04
    |    |--L. (L.) intermedius Chinone & Aoki 1972 S04
    |    |--L. (L.) khencensis Hammer 1961 S04
    |    |--L. (L.) kirghisicus Krivolutsky 1971 S04
    |    |--L. (L.) lacunosus Wang & Cui 1992 S04
    |    |--L. (L.) laetepictus (Berlese 1910) (n. d.) [=Brachychthonius laetepictus] S04
    |    |--L. (L.) lapponicus (Trägardh 1910) [=Hypochthonius brevis lapponicus] S04
    |    |--L. (L.) latus (Jacot 1936) [=Brachychthonius latus] S04
    |    |--L. (L.) lentus Chinone 1974 S04
    |    |--L. (L.) leptaleus Moritz 1976 S04
    |    |--L. (L.) longipilus (Womersley 1945) [=Brachychthonius longipilus] S04
    |    |--L. (L.) mollis (Hammer 1958) [=Brachychthonius mollis] S04
    |    |--L. (L.) moritzi Balogh & Mahunka 1983 (see below for synonymy) S04
    |    |--L. (L.) muscorum Forsslund 1964 S04
    |    |--L. (L.) mussardi Mahunka 1974 S04
    |    |--L. (L.) neglectus Moritz 1976 S04
    |    |--L. (L.) neonomminatus Subías 2004 [=L. latus Mahunka 1982 non Brachychthonius latus Jacot 1936] S04
    |    |--L. (L.) nodifer Hammer 1962 S04
    |    |--L. (L.) nortoni Martínez & Casanueva 1995 S04
    |    |--L. (L.) oceanicus Hammer 1973 S04
    |    |--L. (L.) ohnishi Chinone 1978 S04
    |    |--L. (L.) patagoniensis Hammer 1962 S04
    |    |--L. (L.) peduncularius (Strenzke 1951) [=Brachychthonius peduncularius] S04
    |    |--L. (L.) penicillus Chinone 1978 S04
    |    |--L. (L.) pepitensis Hammer 1962 S04
    |    |--L. (L.) perelegans Moritz 1976 S04
    |    |--L. (L.) perfusorius Moritz 1976 S04
    |    |--L. (L.) phitosi Mahunka 1982 S04
    |    |--L. (L.) plumosus Mahunka 1969 S04
    |    |--L. (L.) prior (Jacot 1938) [=Brachychthonius prior] S04
    |    |--L. (L.) propinquus Niedbala 1972 S04
    |    |--L. (L.) pseudohystricinus Balogh & Mahunka 1983 S04
    |    |--L. (L.) pusillus Chinone 1978 S04
    |    |--L. (L.) rigidisetosus Hammer 1962 S04
    |    |    |--L. r. rigidisetosus S04
    |    |    `--L. r. curtus Hammer 1962 S04
    |    |--L. (L.) saltaensis (Hammer 1958) [=Brachychthonius saltaensis] S04
    |    |--L. (L.) sellnicki (Thor 1930) (see below for synonymy) S04
    |    |--L. (L.) similis Mahunka 1974 S04
    |    |--L. (L.) simplex (Forsslund 1942) [=Brachychthonius simplex] S04
    |    |--L. (L.) strenzkei Forsslund 1963 S04
    |    |--L. (L.) szemmelveiszi Balogh & Balogh 1983 S04
    |    |--L. (L.) tanzanicus Mahunka 1984 S04
    |    |--L. (L.) tuberculatus Hammer 1961 S04
    |    |--L. (L.) tuxeni (Forsslund 1957) [=Brachychthonius hystricinus tuxeni] S04
    |    |--L. (L.) tyrrhenicus Bernini 1985 S04
    |    `--L. (L.) unilateralis Hammer 1962 S04
    `--L. (Afroliochthonius Mahunka 1995) S04
         `--L. (*A.) reductus Mahunka 1995 S04

Liochthonius (Liochthonius) asotthalomensis Marshall, Reeves & Norton 1987 [incl. L. forsslundi Mahunka 1969 non Brachychthonius forsslundi Hammer 1952] S04

Liochthonius (Liochthonius) brevis (Michael 1888) [=Hypochthonius brevis; incl. Brachychthonius perpusillus Berlese 1910, *Liochthonius perpusillus] S04

Liochthonius (Liochthonius) moritzi Balogh & Mahunka 1983 [incl. L. hystricinus tuxeni Chinone 1974 non Brachychthonius hystricinus tuxeni Forsslund 1957] S04

Liochthonius (Liochthonius) sellnicki (Thor 1930) [=Brachychthonius sellnicki; incl. B. brevis expolitus Berlese 1910 (n. d.), B. nodosus Willmann 1952, B. scalaris Forsslund 1942] S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.