Olpioidea

Synsphyronus sp., from Western Australian Museum.

Belongs within: Panctenata.
Contains: Olpiinae.

The Olpioidea was recognised by Harvey (1992) for the families Menthidae and Olpiidae on the basis of the presence of trichobothria on the pedipalpal femur. However, molecular phylogenetic analysis suggests that the Garypidae also fall within the clade formed by these families (Murienne et al. 2008).

<==Olpioidea
    |--Menthidae H06
    |    |--Oligomenthus H06
    |    |--Paramenthus H06
    |    |--Thenmus Harvey in Harvey & Muchmore 1990 H06
    |    |    |--*T. aigialites Harvey in Harvey & Muchmore 1990 H06
    |    |    `--T. augustus Harvey 2006 H06
    |    `--Menthus Chamberlin 1930 H06
    |         |--*M. rossi (Chamberlin 1923) [=Minniza rossi] H06
    |         |--M. californicus Chamberlin 1930 H06
    |         |--‘Atemnus’ gracilis [=Lustrochernes gracilis; incl. Menthus lindahli] H06
    |         `--M. mexicanus Hoff 1945 H06
    `--Olpiidae HL08
         |--+--Olpiinae HL08
         |  `--Garypidae HL08
         |       |  i. s.: Archeolarca rotunda M90
         |       |--Garypus Koch 1873 H10
         |       `--+--Ammogarypus Beier 1962 H10
         |          |--Anagarypus Chamberlin 1930 H10
         |          |    |--A. australianus OW01
         |          |    `--A. heatwolei Muchmore 1982 HL08
         |          |--Elattogarypus Beier 1964 H10
         |          |    |--E. cicatrosus Mahnert 2007 H10
         |          |    |--E. cruciatus Beier 1964 H10
         |          |    `--E. somalicus Mahnert 1984 H10
         |          |--Eremogarypus Beier 1955 H10
         |          |    `--E. eximius Beier 1973 H10
         |          |--Meiogarypus Beier 1955 H10
         |          |    `--M. mirus H10
         |          |--Neogarypus Vachon 1937 H10
         |          |--Paragarypus Vachon 1937 H10
         |          |--Thaumastogarypus Beier 1947 H10
         |          |    `--T. mancus Mahnert 1982 H10
         |          `--Synsphyronus Chamberlin 1930 H10 (see below for synonymy)
         |               |--*S. paradoxus Chamberlin 1930 H12
         |               |--S. apimelus Harvey 1987 HL08
         |               |--S. attiguus Harvey 1987 H12
         |               |--S. christopherdarwini Harvey 2012 H12
         |               |--S. elegans Beier 1954 H10
         |               |--S. ellenae Harvey 2010 H10
         |               |--S. francesae Harvey 2010 H10
         |               |--S. hansenii (With 1908) [=Garypus hansenii, *Idiogarypus hansenii] H10
         |               |--S. leo Harvey 1987 H10
         |               |--‘*Maorigarypus’ melanochelatus Chamberlin 1930 H12
         |               |--S. mimulus Chamberlin 1943 H12
         |               `--S. nullarborensis Beier 1969 H10
         `--Hesperolpiinae [Hesperolpiini] HL08
              |  i. s.: Aphelolpium Hoff 1964 HL08
              |           `--A. cayanum M90
              |         Calocheirus Chamberlin 1930 HL08
              |           `--C. tenerifae Mahnert 2002 HL08
              |         Cardiolpium Mahnert 1986 HL08
              |         Ectatolpium Beier 1947 HL08
              |         Hesperolpium Chamberlin 1930 HL08
              |           |--H. andrewsi M90
              |           `--H. slevini M90
              |         Planctolpium Hoff 1964 HL08
              |           |--P. peninsulae CS90
              |           `--P. suteri M90
              |         Stenolpiodes Beier 1959 HL08
              |         Stenolpium Beier 1955 HL08
              |--Nanolpium Beier 1947 MHG08, HL08
              `--+--Progarypus Beier 1931 MHG08, HL08
                 |    |--P. gracilis Mahnert 2001 HL08
                 |    |--P. liliae Mahnert 2001 HL08
                 |    |--P. nigrimanus Mahnert 2001 HL08
                 |    `--P. setifer Mahnert 2001 HL08
                 `--+--Apolpium Chamberlin 1930 MHG08, HL08
                    |    `--A. parvum Hoff 1945 HL08
                    `--Pachyolpium Beier 1931 MHG08, HL08
                         |--P. atlanticum Mahnert & Schuster 1981 H09
                         `--P. monae M90

Synsphyronus Chamberlin 1930 H10 [incl. Idiogarypus Chamberlin 1930 H12, Maorigarypus Chamberlin 1930 H12; Synsphyronidae]

*Type species of generic name indicated

REFERENCES

[CS90] Corey, D. T., & I. J. Stout. 1990. Ground surface arachnids in sandhill communities of Florida. Journal of Arachnology 18: 167–172.

Harvey, M. S. 1992. The phylogeny and classification of the Pseudoscorpionida (Chelicerata: Arachnida). Invertebrate Taxonomy 6: 1373–1435.

[H06] Harvey, M. S. 2006. New species and records of the pseudoscorpion family Menthidae (Pseudoscorpiones). Records of the Western Australian Museum 23 (2): 167–174.

[H09] Harvey, M. S. 2009. The first Australasian species of the halophilic pseudoscorpion genus Paraliochthonius (Pseudoscorpiones: Chthoniidae). Records of the Western Australian Museum 25 (3): 329–344.

[H10] Harvey, M. S. 2010. Two new species of Synsphyronus (Pseudoscorpiones: Garypidae) from southern Western Australian granite landforms. Records of the Western Australian Museum 26 (1): 11–22.

[H12] Harvey, M. S. 2012. A new species of Synsphyronus (Pseudoscorpiones: Garypidae) from Western Australia. Records of the Western Australian Museum 27 (1): 55–61.

[HL08] Harvey, M. S., & M. C. Leng. 2008. The first troglomorphic pseudoscorpion of the family Olpiidae (Pseudoscorpiones), with remarks on the composition of the family. Records of the Western Australian Museum 24 (4): 387–394.

[M90] Muchmore, W. B. 1990. Pseudoscorpionida. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 503–527. John Wiley & Sones: New York.

[MHG08] Murienne, J., M. S. Harvey & G. Giribet. 2008. First molecular phylogeny of the major clades of Pseudoscorpiones (Arthropoda: Chelicerata). Molecular Phylogenetics and Evolution 49: 170–184.

[OW01] Otto, J. C., & K. J. Wilson. 2001. Assessment of the usefulness of ribosomal 18S and mitochondrial COI sequences in Prostigmata phylogeny. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10^th International Congress pp. 100–109. CSIRO Publishing: Melbourne.