Ormocerinae

Female Lisseurytoma violaceitincta, from Bouček (1988).

Belongs within: Chalcidoidea.
Contains: Systasis, Trichilogaster, Terobiella, Encyrtocephalus.

The Ormocerinae are a group of wasps, most diverse in the Australian region, that are all (so far as is known) either gall-makers or inquilines of other gall-forming insects (Bouček 1988).

Characters (from Bouček 1988): Head without occipital carina. Antennae never inserted very low on face, mostly near to, sometimes above centre; either 13-segmented or 12-segmented. Toruli rather close together, except in the few males which have unusually enlarged scapes. Supraclypeal area not well delimited: clypeus rather small, its lower margin sometimes produced, then it may be bilobate. Thorax with complete and deep notauli; scutellum usually narrowed anteriorly, so that axillae are not widely separated (except in few genera). Thoracic dorsum with complete notauli, without special paired setae or bristles; scutellum without distinct frenum. Propodeum not forming a distinct nucha, always relatively short; spiracles usually large. Forewing venation variable but pilosity near basal cell not specialised. Gaster mostly sessile, first tergite not greally enlarged: hind margins of anterior tergites emarginate or excised in middle, latter state mainly in species in which the gaster is not strongly depressed dorsally.

<==Ormocerinae HB13
    |  i. s.: Ormocerus [Ormocerini] B88
    |--Erixestus HB13
    `--+--Systasini HB13
       |    |--Systasis B88
       |    `--Semiotellus Westwood 1839 (see below for synonymy) B88
       |         |--*S. mundus [=*Semiotus mundus] B88
       |         |--S. longifasciatipennis (Girault 1915) [=*Cheiropachysia longifasciatipennis] B88
       |         |--S. ormyroides (Girault 1915) [=Neosystasis ormyroides] B88
       |         |--S. tenebricosus (Girault 1915) [=Cheiropachysia tenebricosa] B88
       |         `--S. umbilicatus (Girault 1915) [=*Neosystasis umbilicatus, *N. umbilicata] B88
       `--Melanosomellini [Brachyscelidiphagini] HB13
            |--Trichilogaster B88
            |--Terobiella B88
            |--Encyrtocephalus B88
            |--Krivena Bouček 1988 B88
            |    `--*K. pennata Bouček 1988 B88
            |--Neoperilampus Girault & Dodd in Girault 1915 B88
            |    `--*N. niger Girault & Dodd in Girault 1915 B88
            |--Aditrochus Ruebsaamen 1902 B88
            |    |--A. coihuensis MH11
            |    `--A. fagicolus B88
            |--Plastobelyta Kieffer 1906 B88
            |    `--P. gallicola B88
            |--Espinosa Gahan 1947 B88
            |    `--E. nothofagi B88
            |--Neochalcissia Girault 1920 B88
            |    `--*N. magniscapus Girault 1920 B88
            |--Megamelanosoma Girault 1928 B88
            |    `--*M. aeschyli Girault 1928 B88
            |--Epelatus Girault 1928 B88
            |    `--*E. eurytomoideus Girault 1928 [=*E. eurytomoidea] B88
            |--Alloderma Ashmead 1904 B88
            |    `--*A. maculatipennis Ashmead 1904 (see below for synonymy) B88
            |--Alyxiaphagus Riek 1962 B88
            |    `--*A. picturatus Riek 1962 B88
            |--Wubina Bouček 1988 B88
            |    `--*W. cardaleae Bouček 1988 B88
            |--Cerna Bouček 1988 B88
            |    `--*C. kohouti Bouček 1988 B88
            |--Westra Bouček 1988 B88
            |    `--*W. nigra Bouček 1988 B88
            |--Hubena Bouček 1988 B88
            |    `--*H. ruris Bouček 1988 B88
            |--Edgaria Bouček 1988 B88
            |    `--*E. blackburni (Girault 1928) [=Tepperella blackburni, Terobiella blackburni] B88
            |--Nambouria Bouček 1988 B88
            |    `--*N. ramulorum Bouček 1988 B88
            |--Cecidoxenus Ashmead 1904 [incl. Parachrysomalla Girault 1915] B88
            |    |--*C. nigrocyaneus Ashmead 1904 [incl. Parachrysomalla secunda Girault 1917] B88
            |    `--C. aereifemur (Girault 1915) [=*Parachrysomalla aereifemur] B88
            |--Perilampomyia Girault 1916 B88
            |    |--*P. notatifrons Girault 1916 B88
            |    `--P. aeschyli Girault 1931 B88
            |--Lisseurytoma Cameron 1912 [incl. Cratodecatoma Cameron 1912, Sphegigasteroides Girault 1913] B88
            |    |--*L. violaceitincta Cameron 1912 (see below for synonymy) B88
            |    `--L. decorata Ferrière in Gahan & Ferrière 1947 B88
            |--Eurytomomma Girault 1920 B88
            |    |--*E. aurifacies Girault 1920 B88
            |    `--E. atricoxa Girault 1927 B88
            |--Xantheurytoma Cameron 1912 B88
            |    |--*X. flava Cameron 1912 B88
            |    `--X. pallidicoxa (Girault 1928) [=Coelocybelloides pallidicoxa] B88
            |--Lincolna Girault 1940 B88
            |    |--*L. aldrovandii Girault 1940 B88
            |    `--L. sismondini (Girault 1931) [=Terobiella sismondini] B88
            |--Aeschylia Girault 1929 B88
            |    |--*A. latiscapus Girault 1929 B88
            |    |--A. mira Bouček 1988 B88
            |    `--A. picta Bouček 1988 B88
            |--Perilampella Girault & Dodd in Girault 1915 (see below for synonymy) B88
            |    |--P. acaciaediscoloris (Froggatt 1892) (see below for synonymy) B88
            |    |--P. hecataeus (Walker 1839) (see below for synonymy) B88
            |    |--P. nigrisetae (Girault 1929) [=Coelocybelloides nigrisetae] B88
            |    `--P. pulchra (Girault 1929) [=Coelocybelloides pulchra] B88
            `--Brachyscelidiphaga Ashmead 1900 [incl. Coelocybella Girault 1913, Coelocybelloides Girault 1915] B88
                 |--*B. flava Ashmead 1900 [incl. B. masaccioni Girault 1931] B88
                 |--B. aeschyli Girault 1931 B88
                 |--B. aurea (Girault 1915) [=*Coelocybelloides aureus] B88
                 |--B. pulchrivariegata (Girault 1915) [=Coelocybelloides pulchrivariegatus] B88
                 |--‘Coelocybelloides’ trilineata G31
                 `--B. variegata (Girault 1913) [=*Coelocybella variegata] B88

*Alloderma maculatipennis Ashmead 1904 [=Coelocybelloides bioculatus Girault 1916, Perilampella punctata Girault 1931] B88

*Lisseurytoma violaceitincta Cameron 1912 [incl. L. bicolor Gahan in Gahan & Ferrière 1947, *Cratodecatoma ruficeps Cameron 1912, *Sphegigasteroides rufinotum Girault 1913] B88

Perilampella Girault & Dodd in Girault 1915 [incl. Trichilogastroides Ferrière in Gahan & Ferrière 1947] B88

Perilampella acaciaediscoloris (Froggatt 1892) [=Cynips acaciaediscoloris, *Trichilogastroides acaciaediscoloris; incl. *Perilampella flammeithorax Girault 1915] B88

Perilampella hecataeus (Walker 1839) [=Miscogaster hecataeus, Lamprotatus hecataeus; incl. Coelocybelloides mediolineatus Girault 1916, Tepperella trilineata Cameron 1912, Tep. trimaculata (l. c.), Terobiella trilineata, Trichilogaster trilineata] B88

Semiotellus Westwood 1839 [=Semiotus Walker 1834 non Eschscholtz 1829; incl. Cheiropachysia Girault 1915, Neosystasis Girault 1915] B88

*Type species of generic name indicated

REFERENCES

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[G31] Girault, A. A. 1931. A new habit in an old insect, Homo pudicus and new Eurytomidae. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 280-283).

[HB13] Heraty, J. M., R. A. Burks, A. Cruaud, G. A. P. Gibson, J. Liljeblad, J. Munro, J.-Y. Rasplus, G. Delvare, P. Janšta, A. Gumovsky, J. Huber, J. B. Woolley, L. Krogmann, S. Heydon, A. Polaszek, S. Schmidt, D. C. Darling, M. W. Gates, J. Mottern, E. Murray, A. D. Molin, S. Triapitsyn, H. Baur, J. D. Pinto, S. van Noort, J. George & M. Yoder. 2013. A phylogenetic analysis of the megadiverse Chalcidoidea (Hymenoptera). Cladistics 29: 466-542.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.