Sciophilini

Sciophila sp., photographed by pwalter.

Belongs within: Mycetophilidae.

The Sciophilini is a group of sciophiline fungus gnats distinguished by the presence of macrotrichia on the wings (Tonnoir & Edwards 1927). However, monophyly of the group has been questioned on the basis of similarities of some genera to species lacking macrotrichia (Vockeroth 1981).

Characters (from Tonnoir & Edwards 1927): Wing with macrotrichia on the membrane; microtrichia sometimes absent. Postnotum usually with hairs or bristles; Sc nearly always long; last section of R₁ several times as long as r-m, which is oblique; 7th abdominal segment usually large and visible.

<==Sciophilini
    |--Trizygia TE27
    |--Loicia basifurca V81
    |--Azana V81
    |--Monoclona rufilatera V81
    |--Cluzobra V81
    |--Acnemia V81
    |--Polylepta guttiventris V81
    |--Anaclileia V81
    |--Baeopterogyna nudipes V81
    |--Neuratelia sayi V81
    |--Leptomorphus nebulosus V81
    |--Eudicrana obumbrata V81
    |--Neotrizygia Tonnoir & Edwards 1927 TE27
    |    `--*N. obscura Tonnoir in Tonnoir & Edwards 1927 TE27
    |--Morganiella Tonnoir & Edwards 1927 TE27
    |    `--*M. fusca Tonnoir in Tonnoir & Edwards 1927 TE27
    |--Phthinia V81
    |    |--P. longiventris Tonnoir in Tonnoir & Edwards 1927 TE27
    |    `--P. tanypus V81
    |--Taxicnemis Tonnoir & Edwards 1927 TE27
    |    |--*T. ‘hirta’ (Marshall 1896) (see below for synonymy) TE27
    |    `--T. flava Edwards in Tonnoir & Edwards 1927 TE27
    |--Paratinia V81
    |    |--P. montana Plassmann 1976 P76a
    |    `--P. recurva V81
    |--Syntemna V81
    |    |--S. hungarica P76b
    |    |--S. nitidula P76b
    |    `--S. vernalis V81
    |--Allocotocera V81 [incl. Euryceras Marshall 1896 TE27]
    |    |--A. anaclinoides (Marshall 1896) [=*Euryceras anaclinoides] TE27
    |    |--A. cephasi Edwards in Tonnoir & Edwards 1927 TE27
    |    |--A. crassipalpis Tonnoir in Tonnoir & Edwards 1927 TE27
    |    |--A. dilatata Tonnoir in Tonnoir & Edwards 1927 TE27
    |    `--A. pulchella V81
    |--Sciophila [incl. Megalopelma] V09
    |    |--S. adamsi P76b
    |    |--‘Megalopelma’ glabanum V81
    |    |--S. hirta Meigen 1818 TC04
    |    |--S. novata V81
    |    |--S. ocreata Philippi 1865 TC04
    |    `--S. parviareolata Santos Abreu 1920 TC04
    |--Aphelomera TE27
    |    |--A. elongata Tonnoir in Tonnoir & Edwards 1927 TE27
    |    |--A. forcipata Edwards in Tonnoir & Edwards 1927 TE27
    |    |--A. longicauda Edwards in Tonnoir & Edwards 1927 TE27
    |    |--A. majuscula Edwards in Tonnoir & Edwards 1927 TE27
    |    |--A. marshalli Edwards in Tonnoir & Edwards 1927 TE27
    |    |--A. opaca Tonnoir in Tonnoir & Edwards 1927 TE27
    |    `--A. skusei Marshall 1896 TE27
    |--Parvicellula Marshall 1896 TE27
    |    |--*P. triangula Marshall 1896 TE27
    |    |--P. apicalis Tonnoir in Tonnoir & Edwards 1927 TE27
    |    |--P. fascipennis Edwards in Tonnoir & Edwards 1927 TE27
    |    |--P. gracilis Tonnoir in Tonnoir & Edwards 1927 TE27
    |    |--P. hamata Edwards in Tonnoir & Edwards 1927 TE27
    |    |--P. nigricoxa Tonnoir in Tonnoir & Edwards 1927 TE27
    |    |--P. obscura Tonnoir in Tonnoir & Edwards 1927 TE27
    |    |--P. ruficoxa Tonnoir in Tonnoir & Edwards 1927 TE27
    |    `--P. subhamata Tonnoir in Tonnoir & Edwards 1927 TE27
    `--Aneura Marshall 1896 TE27
         |--*A. boletinoides Marshall 1896 TE27
         |--A. appendiculata Tonnoir in Tonnoir & Edwards 1927 TE27
         |--A. bispinosa Edwards in Tonnoir & Edwards 1927 TE27
         |--A. defecta Edwards in Tonnoir & Edwards 1927 TE27
         |--A. fagi (Marshall 1896) [=Sciophila fagi] TE27
         |--A. filiformis Tonnoir in Tonnoir & Edwards 1927 TE27
         |--A. fusca Tonnoir in Tonnoir & Edwards 1927 TE27
         |--A. longicauda Tonnoir in Tonnoir & Edwards 1927 TE27
         |--A. longipalpis Tonnoir in Tonnoir & Edwards 1927 TE27
         |--A. nitida Tonnoir in Tonnoir & Edwards 1927 TE27
         `--A. pallida Edwards in Tonnoir & Edwards 1927 TE27

*Taxicnemis ‘hirta’ (Marshall 1896) [=Sciophila hirta non Meigen 1818; incl. T. hirta var. bivittata Tonnoir & Edwards 1927] TE27

*Type species of generic name indicated

REFERENCES

[P76a] Plassmann, E. 1976a. Zwei neue Pilzmücken aus dem Kaukasus (Diptera: Mycetophilidae). Senckenbergiana Biologica 57 (1–3): 69–70.

[P76b] Plassmann, E. 1976b. Vierter Nachtrag zu der Mycetophiliden-Sammlung des Senckenberg-Museums Frankfurt am Main (Diptera; Mycetophilidae). Senckenbergiana Biologica 57 (1–3): 77–79.

[TC04] Toft, R. J., & P. J. Chandler. 2004. Three introduced species of Mycetophilidae (Diptera: Sciaroidea) established in New Zealand. New Zealand Entomologist 27: 43–49.

[TE27] Tonnoir, A. L., & F. W. Edwards. 1927. New Zealand fungus gnats (Diptera, Mycetophilidae). Transactions and Proceedings of the New Zealand Institute 57: 747–878.

[V81] Vockeroth, J. R. 1981. Mycetophilidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 223–246. Research Branch, Agriculture Canada.

[V09] Vockeroth, J. R. 2009. Mycetophilidae (fungus gnats). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 267–278. NRC Research Press: Ottawa.

Last updated: 18 June 2018.