Belongs within: Boraginaceae.
Contains: Pectocarya, Plagiobothrys, Amsinckia, Hackelia, Trichodesmeae, Myosotis, Cryptantha, Cynoglossum.
The Cynoglosseae were recognised by Långström & Chase (2002) as a clade within the Boraginaceae containing taxa with heterocolpate pollen, an undivided style with one stigma, and usually well-developed appendages or wings on the nutlets. A previously recognised division between tribes Cynoglosseae, Eritrichieae and Myosotideae was not supported by Långström & Chase's (2002) molecular analysis. Members of the clade include the stickseeds of the genus Lappula, a Northern Hemisphere genus of annual herbs producing nutlet fruits covered more or less long, barbed prickles (Hickman 1993).
See also: Sending forget-me-nots.
<==Cynoglosseae [Eritrichieae]
|--Omphalodes verna LC02
`--+--Trichodesmeae LC02
`--+--Lappula LC02
| |--L. barbata GR98
| |--L. microcarpa O88
| |--L. redowskii H93
| | |--L. r. var. redowskii H93
| | `--L. r. var. cupulata [incl. L. redowskii var. desertorum] H93
| `--L. squarrosa [incl. L. echinata] H93
`--+--Myosotis LC02
|--+--Pardoglossum cheirifolium LC02
| |--Cryptantha LC02
| `--+--Lindelofia longiflora LC02
| `--Cynoglossum LC02
`--Trigonotis LC02
|--T. brevipes LC02
|--T. multicaulis O88
|--T. ovalifolia O88
|--T. peduncularis LC02
|--T. rotundifolia O88
`--T. tibetica O88
Cynoglosseae incertae sedis:
Actinocarya LC02
Afrotysonia LC02
Antiotrema LC02
Diploloma LC02
Gyrocaryum LC02
Harpagonella palmeri LC02, H93
Heliocarya LC02
Heterocaryum LC02
Krynitzkia LC02
Kuschakewiczia LC02
Mattia LC02
Mattiastrum LC02
Paracaryum LC02
Pectocarya LC02
Rindera LC02
Solenanthus LC02
Thyrocarpus LC02
Trachelanthus LC02
Tysonia LC02
Anoplocaryum LC02
Asperugo procumbens LC02, C55
Bothriospermum LC02
Craniospermum LC02
Echiochospermum LC02
Eremocarya LC02
Lepechiniella LC02
Myosotidium Hooker 1859 LC02, A61
`--M. hortensia (Decne) Baill. 1891 (see below for synonymy) A61
Nesocaryum LC02
Omphalolappula Brand 1931 LC02, BR65
`--O. concava (Mueller) Brand 1931 (see below for synonymy) BR65
Oreocarya LC02
Plagiobothrys LC02
Pseudomertensia LC02
Rochelia stellulata LC02, C55
Schistocaryum LC02
Tianschaniella LC02
Tretocarya LC02
Amsinckia LC02
Hackelia LC02
Echinospermum LC02
|--E. deflexum C55
`--E. lappula C55
Microula LC02
|--M. pustulosa O88
|--M. sikkimensis O88
`--M. tibetica O88
Eritrichium LC02
|--E. canum O88
|--E. fulvum C55
|--E. minimum O88
|--E. nanum H09
| |--E. n. ssp. nanum H09
| `--E. n. ssp. jankae H09
`--E. thomsonii O88
Myosotidium hortensia (Decne) Baill. 1891 [=Myosotis hortensia Decne in Deless. 1846; incl. Cynoglossum nobile Hooker 1858, Myosotidium nobile Hook. 1859] A61
Omphalolappula concava (Mueller) Brand 1931 [=Echinospermum concavum Mueller 1860, Cynoglossum concavum Kuntze 1891, Lappula concava Mueller 1882] BR65
*Type species of generic name indicated
REFERENCES
[A61] Allan, H. H. 1961. Flora of New Zealand vol. 1. Indigenous Tracheophyta: Psilopsida, Lycopsida, Filicopsida, Gymnospermae, Dicotyledones. R. E. Owen, Government Printer: Wellington (New Zealand).
[BR65] Black, J. M., & E. L. Robertson. 1965. Flora of South Australia. Part IV. Oleaceae–Compositae. W. L. Hawes, Government Printer: Adelaide.
[C55] Candolle, A. de. 1855. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 2. Librairie de Victor Masson: Paris.
[GR98] Greuter, W., & T. Raus (eds.) 1998. Med-Checklist Notulae, 17. Willdenowia 28: 163–174.
[H09] Heltmann, H. 2009. Der Königstein (Piatra Craiului), die Perle der Burzenländer Gebirge. Mauritiana 20 (3): 515–527.
[H93] Hickman, J. C. (ed.) 1993. The Jepson Manual: Higher Plants of California. University of California Press: Berkeley (California).
[LC02] Långström, E., & M. W. Chase. 2002. Tribes of Boraginoideae (Boraginaceae) and placement of Antiphytum, Echiochilon, Ogastemma and Sericostoma: a phylogenetic analysis based on atpB plastid DNA sequence data. Plant Systematics and Evolution 234: 137–153.
[O88] Ohba, H. 1988. The alpine flora of the Nepal Himalayas: an introductory note. In: Ohba, H., & S. B. Malla (eds) The Himalayan Plants vol. 1. The University Museum, University of Tokyo, Bulletin 31: 19–46.
Last updated: 27 October 2019.
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