Hoplocarida

Heterosquilla tricarinata, copyright Svamp.

Belongs within: Eumalacostraca.
Contains: Squillidae, Lysiosquillidae, Gonodactyloidea.

The Hoplocarida, including the mantis shrimps and their fossil relatives, are a group of malacostracan crustaceans bearing a small carapace and at least four free thoracic segments, together with a large abdomen ending in a tail fan.

<==Hoplocarida
    |  i. s.: Neogonodactylus RS10
    |           |--N. bredini SC17
    |           `--N. oerstedii RS10
    |--Perimecturidae [Palaeostomatopoda] BWW93
    |    |--Archaeocaris graffhami Brooks 1962 BWW93
    |    |--Eopteridium striatum Rohdendorf 1970 BWW93
    |    |--Bairdops beargulchensis Schram & Horner 1978 BWW93
    |    `--Perimecturus [incl. Eopterum; Archaeoptera] S77
    |         |--‘Eopterum’ devonicum Rohdendorf 1961 BWW93
    |         `--P. rapax Schram & Horner 1978 BWW93
    |--Aeschronectida G88
    |    |--Kallidechtes [Kallidecthidae] BWW93
    |    |    |--K. eageri Schram 1979 BWW93
    |    |    `--K. richardsoni Schram 1969 BWW93
    |    |--Aratidecthidae BWW93
    |    |    |--Crangopsis BWW93
    |    |    |    |--C. cooperi [=Pygocephalus cooperi] S74
    |    |    |    `--C. eskdalensis (Peach 1882) BWW93
    |    |    `--Aratidecthes johnsoni Schram 1969 BWW93
    |    `--Aenigmacarididae BWW93
    |         |--Joanellia BWW93
    |         |    |--J. elegans (Peach 1883) BWW93
    |         |    `--J. lundi G88
    |         `--Aenigmacaris G88
    |              |--A. cornigerum G88
    |              `--A. minima Schram & Schram 1979 BWW93
    `--Stomatopoda MD01
         |  i. s.: Alima hyalina Leach 1817 C73
         |         Pseudosquillopsis cerisii (Roux 1828) C73
         |         Eurysquilloides [Eurysquilloididae, Eurysquilloidoidea] MD01
         |         Erythrosquillidae [Erythrosquilloidea] MD01
         |--Tyrannophontidae [Archaeostomatopoda] BWW93
         |    |  i. s.: ‘Perimecturus’ pattoni Peach 1908 BWW93
         |    |--Tyrannophontes theridion BWW93
         |    `--Gorgonophontes peleron Schram 1984 BWW93
         `--Unipeltata BWW93
              |--Squillidae BWW93
              |--Bathysquilloidea BWW93
              |    |--Indosquillidae MD01
              |    |--Bathysquilla C73 [Bathysquillidae MD01]
              |    |    |--B. microps (Manning 1961) C73
              |    |    `--B. wetherelli (Woodward 1879) BWW93
              |    `--Sculdidae BWW93
              |         |--Pseudosculda laevis Schluter 1872 BWW93
              |         `--Sculda BWW93
              |              |--S. pennata Münster 1840 BWW93
              |              `--S. syriaca Dames 1886 BWW93
              |--Lysiosquilloidea BWW93
              |    |--Lysiosquillidae C73
              |    |--Coronis C73 [Coronididae MD01]
              |    |    `--C. excavatrix Brooks 1886 C73
              |    |--Nannosquilla C73 [Nannosquillidae MD01]
              |    |    |--N. occulta [=Lysiosquilla occulta] C73
              |    |    |--N. schmitti (Manning 1962) C73
              |    |    `--N. taylori Manning 1969 C73
              |    `--Heterosquilla K-MM02 [Heterosquillidae, Tetrasquillidae MD01]
              |         |--H. armata K-MM02
              |         |--H. brazieri HS01
              |         `--H. tricarinata HS01
              `--Gonodactyloidea BWW93

*Type species of generic name indicated

REFERENCES

[BWW93] Briggs, D. E. G., M. J. Weedon & M. A. Whyte. 1993. Arthropoda (Crustacea excluding Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 321–342. Chapman & Hall: London.

[C73] Camp, D. K. 1973. Stomatopod Crustacea. Memoirs of the Hourglass Cruises 3 (2): 1–100.

[G88] Gray, J. 1988. Evolution of the freshwater ecosystem: the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology 62: 1–214.

[HS01] Hayward, B. W., A. B. Stephenson, M. S. Morley, W. M. Blom, H. R. Grenfell, F. J. Brook, J. L. Riley, F. Thompson & J. J. Hayward. 2001. Marine biota of Parengarenga Harbour, Northland, New Zealand. Records of the Auckland Museum 37: 45–80.

[K-MM02] Klein-MacPhee, G., & R. S. McBride. 2002. Sea robins. Family Triglidae. In: Collette, B. B., & G. Klein-MacPhee (eds) Bigelow and Schroeder’s Fishes of the Gulf of Maine 3^rd ed. pp. 338–345. Smithsonian Institute Press: Washington.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1–124.

[RS10] Regier, J. C., J. W. Shultz, A. Zwick, A. Hussey, B. Ball, R. Wetzer, J. W. Martin & C. W. Cunningham. 2010. Arthropod relationships revealed by phylogenomic analysis of nuclear protein-coding sequences. Nature 463: 1079–1083.

[S74] Schram, F. R. 1974. Convergences between Late Paleozoic and modern caridoid Malacostraca. Systematic Zoology 23 (3): 323–332.

[S77] Schram, F. R. 1977. Paleozoogeography of Late Paleozoic and Triassic Malacostraca. Systematic Zoology 26 (4): 367–379.

[SC17] Schwentner, M., D. J. Combosch, J. P. Nelson & G. Giribet. 2017. A phylogenomic solution to the origin of insects by resolving crustacean-hexapod relationships. Current Biology 27: 1818–1824.