Beyrichioidea

Male (above) and female Kloedenia normalis, from Benson et al. (1961).

Belongs within: Palaeocopida.
Contains: Beyrichia.

The Beyrichioidea are a group of ostracods known from the mid-Ordovician to the early Permian characterised by the dimorphic development of a ventral to anteroventral lobe or pouch in females. The hinge of the carapace is straight but the Silurian Zygobolbidae have the lateral lobes L₂ and L₃ tending to project above the level of the dorsum (Benson et al. 1961).

Characters (from Benson et al. 1961): Straight-hinged ostracodes with subequal ends or forward swing, mostly with well-developed lobes and sulci and showing tendency to have carinal, velate, and marginal structures; dimorphism usually well defined.

Beyrichioidea [Beyrichiacea, Beyrichiomorphinina] L05
    |  i. s.: Gillatia Öpik 1953 BB61
    |           `--*G. trinacria Öpik 1953 BB61
    |--Treposellidae [Treposellinae] WSB93
    |    |--Loutriella jupiterensis Copeland 1974 WSB93
    |    |--Parabouckekius martinssoni Jones 1985 WSB93
    |    |--Katatona romei Jones 1985 WSB93
    |    `--Treposella Ulrich & Bassler 1908 BB61
    |         |--*T. lyoni (Ulrich 1891) [=Beyrichia lyoni] BB61
    |         `--T. stellata BB61
    |--Craspedobolbinidae WSB93
    |    |--Malnina spinosa Jones 1989 WSB93
    |    |--Armilla sibirica Bushmina 1975 WSB93
    |    |--Craspedobolbina Kummerow 1923 BB61
    |    |    `--*C. dietrichi Kummerow 1923 BB61
    |    |--Copelandella Bless & Jordan 1971 WSB93, G88
    |    |    `--C. novascotia (Jones & Kirby 1884) WSB93
    |    `--Bolbineossia Kesling, Heany et al. 1958 WSB93, BB61
    |         |--*B. didictyosa Kesling, Heany et al. 1958 BB61
    |         `--B. pineaulti Copeland 1974 WSB93
    |--Zygobolbidae [Zygobolbinae] BB61
    |    |--Bonnemaia Ulrich & Bassler 1923 BB61
    |    |    `--*B. celsa Ulrich & Bassler 1923 BB61
    |    |--Glymmatobolbina Harris 1957 BB61
    |    |    `--*G. quadrata Harrs 1957 BB61
    |    |--Mastigobolbina Ulrich & Bassler 1923 BB61
    |    |    `--*M. typa Ulrich & Bassler 1923 BB61
    |    |--Plethobolbina Ulrich & Bassler 1923 BB61
    |    |    `--*P. typicalis Ulrich & Bassler 1923 BB61
    |    `--Zygobolba Ulrich & Bassler 1923 BB61
    |         |--*Z. (Zygobolba) decora (Billings 1866) [=Beyrichia decora] BB61
    |         |--Z. (Zygobolbina Ulrich & Bassler 1923) BB61
    |         |    `--Z. (*Z.) conradi (Ulrich & Bassler 1923) [=*Zygobolbina conradi] BB61
    |         `--Z. (Zygosella Ulrich & Bassler 1923) BB61
    |              `--Z. (*Z.) vallata (Ulrich & Bassler 1923) [=*Zygosella vallata] BB61
    |--Kloedenia Jones & Holl 1886 BB61 [Kloedeniidae L05, Kloedeniinae, Kloedeninae]
    |    |--*K. wilckensiana (Jones 1855) [=Beyrichia wilckensiana] BB61
    |    |--K. apiculata Jones 1888 C01
    |    |--K. australis Chapman 1920 F71
    |    |--K. fifieldensis Chapman 1920 F71
    |    |--K. gotlandica Chapman 1901 C01
    |    |--K. normalis BB61
    |    `--K. sussexensis BB61
    |--Ochescaphidae L05
    `--Beyrichiidae [Beyrichiinae, Beyrichinae] BB61
         |--Fallaticella schaeferi Schallreuter 1984 WSB93
         |--Apatobolbina Ulrich & Bassler 1923 BB61
         |    `--*A. granifera Ulrich & Bassler 1923 BB61
         |--Bolbibollia Ulrich & Bassler 1923 BB61
         |    `--*B. labrosa Ulrich & Bassler 1923 BB61
         |--Bolbiprimitia Kay 1940 BB61
         |    `--*B. fissurella (Ulrich & Bassler 1923) [=Halliella fissurella] BB61
         |--Cornikloedenia Henningsmoen 1954 BB61
         |    `--*C. ventralis (Ulrich & Bassler 1923) [=Drepanellina ventralis] BB61
         |--Ctenobolbinella Kummerow 1953 BB61
         |    `--*C. carinata Kummerow 1953 BB61
         |--Dolichoscapha Kesling & Ehlers 1958 BB61
         |    `--*D. escharota Kesling & Ehlers 1958 BB61
         |--Drepanellina Ulrich & Bassler 1923 BB61
         |    `--*D. clarki Ulrich & Bassler 1923 BB61
         |--Halliella Ulrich 1891 BB61
         |    `--*H. retifera Ulrich 1891 BB61
         |--Hibbardia Kesling 1953 BB61
         |    `--*H. lacrimosa (Swartz & Oriel 1948) [=Amphissites lacrimosus] BB61
         |--Mesomphalus Ulrich & Bassler 1913 [=Mezomphalus Zaspelova 1952] BB61
         |    `--*M. hartleyi Ulrich & Bassler 1913 BB61
         |--Myomphalus Swartz & Whitmore 1956 BB61
         |    `--*M. dorsinodosus Swartz & Whitmore 1956 BB61
         |--Phlyctiscapha Kesling 1953 BB61
         |    `--*P. rockportensis Kesling 1953 BB61
         |--Pseudobeyrichia Swartz & Whitmore 1956 BB61
         |    `--*P. perornata Swartz & Whitmore 1956 BB61
         |--Saccarchites Swartz & Whitmore 1956 BB61
         |    `--*S. saccularis Swartz & Whitmore 1956 BB61
         |--Welleria Ulrich & Bassler 1923 BB61
         |    `--*W. obliqua Ulrich & Bassler 1923 BB61
         |--Bodenia Ivanova 1959 LH61
         |    `--*B. aspera Ivanova 1959 LH61
         |--Pseudoleperditia Schneider 1956 WSB93, BB61
         |    `--*P. tuberculifera Schneider 1956 BB61
         |--Dibolbina Ulrich & Bassler 1923 BB61
         |    |--*D. cristata Ulrich & Bassler 1923 BB61
         |    `--D. steusloffi BB61
         |--Lophokloedenia Swartz & Whitmore 1956 BB61
         |    |--*L. manliensis (Weller 1903) [=Beyrichia manliensis] BB61
         |    `--L. kummeli BB61
         |--Welleriopsis Swartz & Whitmore 1956 BB61
         |    |--*W. diplocystulis Swartz & Whitmore 1956 BB61
         |    `--W. jerseyensis BB61
         |--Zygobeyrichia Ulrich 1916 BB61
         |    |--*Z. apicalis Ulrich 1916 BB61
         |    `--Z. barretti BB61
         |--Kyamodes Jones 1888 [=Kyammodes Ulrich & Bassler 1908] BB61
         |    |--*K. whidbornei Jones 1888 BB61
         |    |--K. kiesowi BB61
         |    `--K. tricornis BB61
         `--Beyrichia BB61

*Type species of generic name indicated

REFERENCES

[BB61] Benson, R. H., J. M. Berdan, W. A. van den Bold, T. Hanai, I. Hessland, H. V. Howe, R. V. Kesling, S. A. Levinson, R. A. Reyment, R. C. Moore, H. W. Scott, R. H. Shaver, I. G. Sohn, L. E. Stover, F. M. Swain & P. C. Sylvester-Bradley. 1961. Systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q99–Q421. Geological Society of America and University of Kansas Press.

[C01] Chapman, F. 1901. On some fossils of Wenlock age from Mulde, near Klinteberg, Gotland; with notes by Prof. T. Rupert Jones and Dr. F. A. Bather. Annals and Magazine of Natural History, series 7, 7: 141–160, pl. 3.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[G88] Gray, J. 1988. Evolution of the freshwater ecosystem: the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology 62: 1–214.

[LH61] Levinson, S. A., & H. V. Howe. 1961. Addendum. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q421–Q422. Geological Society of America and University of Kansas Press.

[L05] Liebau, A. 2005. A revised classification of the higher taxa of the Ostracoda (Crustacea). Hydrobiologia 538: 115–137.

[WSB93] Whatley, R. C., D. J. Siveter & I. D. Boomer. 1993. Arthropoda (Crustacea: Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 343–356. Chapman & Hall: London.