Hypoxestinae

Diagnostic features of Hypoxestus glaber, from Kauri (1985).

Belongs within: Assamiidae.

The Hypoxestinae are a group of assamiid harvestmen found in Africa and Asia (Santos & Prieto 2010).

Characters (from Santos & Prieto 2010): Anterior margin of carapace unarmed; spiracles visible, unprotected; femur of pedipalp with short ventral teeth; distitarsus I two-segmented; metatarsus II undivided; tarsi III and IV with smooth claws.

<==Hypoxestinae R35
    |--Scabrobunus Roewer 1912 R35
    |    `--S. filipes Roewer 1912 R35
    |--Passula Roewer 1927 R35
    |    `--P. scabricula Roewer 1927 R35
    |--Podaucheniellus Roewer 1927 R35
    |    `--*P. bipalaris Roewer 1927 S92
    |--Nkogoa Roewer 1927 R35
    |    `--*N. feai Roewer 1927 S92 [=N. feae R35]
    |--Findia Roewer 1915 R35
    |    `--*F. atrolutea Roewer 1915 S92
    |--Congonella Roewer 1935 R35
    |    `--*C. frontalis Roewer 1935 R35
    |--Randilea Roewer 1935 R35
    |    `--*R. scabricula Roewer 1935 R35 [=Hypoxestus scabricula S92]
    |--Talaspus Roewer 1935 R35
    |    `--*T. spinimanus Roewer 1935 R35
    |--Parasesostris Roewer 1915 R35
    |    `--*P. granulatus Roewer 1915 S92
    |--Metasesostris Roewer 1915 R35
    |    `--*M. armatus Roewer 1915 S92
    |--Tusipulla Roewer 1935 R35
    |    `--*T. coxalis Roewer 1935 R35
    |--Podauchenius Sørensen 1896 R35
    |    `--*P. longipes Sørensen 1896 S92
    |--Musola Roewer 1927 R35
    |    `--*M. longipes Roewer 1927 S92
    |--Adamauna Roewer 1935 R35
    |    `--*A. maculatipes Roewer 1935 R35
    |--Rhabdopygata Roewer 1954 SP10, S92
    |    `--*R. mossambica Roewer 1954 S92
    |--Lossida Roewer 1935 L62, R35
    |    `--*L. rugosa Roewer 1935 R35
    |--Lossidacola Roewer 1935 L62, R35
    |    `--*L. pachytarsus Roewer 1935 R35
    |--Bwitonatus Roewer 1950 L62, S92
    |    `--*B. marlieri Roewer 1950 S92
    |--Leleupiolus Roewer 1951 L62, S92
    |    `--*L. marmoratus Roewer 1951 S92
    |--Mecutina Roewer 1935 R35
    |    |--*M. filipes Roewer 1935 R35
    |    `--M. moshinia Roewer 1940 S92
    |--Sesostranus Roewer 1935 R35
    |    |--*S. niger Roewer 1935 R35
    |    `--S. longipes Roewer 1950 S92
    |--Bandona Roewer 1927 SG11, R35
    |    |--B. boninensis Suzuki 1974 S78
    |    `--B. palpalis Roewer 1927 R35
    |--Hypoxestinus Roewer 1927 R35
    |    |--*H. nkogoi Roewer 1927 S92
    |    `--H. frontalis Roewer 1935 R35
    |--Rhabdopygella Roewer 1935 R35
    |    |--*R. laevis Roewer 1935 S92
    |    `--R. ferruginea Roewer 1935 R35
    |--Dongila Roewer 1927 R35
    |    |--*D. spinosa Roewer 1927 S92
    |    `--D. silvatica Roewer 1927 R35
    |--Mabwella Roewer 1952 L62, S92
    |    |--*M. wittei Roewer 1952 S92
    |    `--M. trochanteralis Roewer 1954 S92
    |--Sesostrellus Roewer 1935 R35
    |    |--*S. robustus Roewer 1935 R35
    |    |--S. gibbosus Kauri 1985 S92
    |    `--S. umbonatus (Roewer 1935) [=Sesostris umbonatus] S92
    |--Doloressus Roewer 1935 [incl. Ealabius Roewer 1950] K85
    |    |--*D. cippatus Roewer 1935 R35
    |    |--D. filipes Lawrence 1949 S92
    |    |--D. ghesquierei (Roewer 1950) [=Ealabius ghesquierei] S92
    |    `--D. palmgreni Kauri 1985 S92
    |--Dicoryphus Loman 1902 R35 (see below for synonymy)
    |    |--*D. furvus Loman 1902 S92
    |    |--D. ater (Lawrence 1962) [=Caelobunus ater] S92
    |    |--D. jeanneli (Roewer 1913) [=Metarhabdopygus jeanneli, Caelobunus jeanneli] S92
    |    `--D. melanacanthus (Loman 1902) (see below for synonymy) S92
    |--Sesostris Sørensen 1910 R35
    |    |--*S. gracilis Sørensen 1910 S92
    |    |--S. brevipes Kauri 1985 S92
    |    |--S. insulanus Roewer 1912 R35
    |    |--S. maculatus Roewer 1915 R35
    |    `--S. maculifer Kauri 1985 [=S. maculiferus] S92
    |--Viglua Roewer 1940 SP10
    |    |--*V. brunnipes Roewer 1940 S92
    |    `--V. machadoi Lawrence 1949 S92
    |         |--V. m. machadoi S92
    |         |--V. m. granulosa Lawrence 1957 S92
    |         `--V. m. major Lawrence 1949 [=V. majora] S92
    |--Spinixestus Roewer 1952 L62, S92
    |    |--*S. armatus Roewer 1952 S92
    |    |--S. benderanus Kauri 1985 S92
    |    |--S. leleupi Kauri 1985 S92
    |    |--S. polycuspidatus Kauri 1985 S92
    |    |--S. rufus Roewer 1952 S92
    |    `--S. siteteus Roewer 1961 S92
    |--Rhabdopygus Roewer 1912 R35
    |    |--*R. fuscus Roewer 1912 S92
    |    |--R. benoiti Kauri 1985 S92
    |    |--R. funilignorum Santos & Prieto 2010 SP10
    |    |--R. maculatus Roewer 1935 R35
    |    |--R. puylaerti Bauer & Prieto 2009 SP10
    |    |--R. robustus Lawrence 1957 S92
    |    |--R. rugipalpis Roewer 1952 S92
    |    `--R. termitarum Roewer 1951 S92
    `--Hypoxestus Loman 1902 S92 [=Amasilla Strand 1911 R23, Amasis Sørensen 1910 R23]
         |  i. s.: H. bituberculatus Lawrence 1962 S92
         |         H. coxicornis Roewer 1940 S92
         |         H. patellaris (Sørensen 1910) [=Amasis patellaris] S92
         |         H. planus Goodnight & Goodnight 1959 S92
         |         H. roeweri Staręga 1992 [=Randilea obscura Roewer 1952 non H. obscurus Roewer 1912] S92
         |         H. trituberculatus Lawrence 1962 S92
         |--*H. (Hypoxestus) levis Loman 1902 (see below for synonymy) S92
         `--H. (Lomanus Kauri 1985) K85
              |--H. (*L.) glaber Kauri 1985 K85
              |--H. (L.) ealanus Kauri 1985 K85
              `--H. (L.) scaphoides Kauri 1985 K85

*Hypoxestus (Hypoxestus) levis Loman 1902 [incl. Amasis accentuatus Sørensen 1910, H. accentuatus, A. mesoleucus Sørensen 1910, H. mesoleucus, H. obscurus Roewer 1912, Amasis quadricornis Sørensen 1910, H. quadricornis] S92

*Type species of generic name indicated

REFERENCES

[K85] Kauri, H. 1985. Opiliones from central Africa. Koninklijk Museum voor Midden-Afrika, Turveren, België, Zoologische Wetenschappen, Annalen 245: 1–167.

[L62] Lawrence, R. F. 1962. Resultats scientifiques des missions zoologiques de l'I.R.S.A.C. en Afrique orientale (P. Basilewsky et N. Leleup, 1957). LXXIV. Opiliones. Annales du Musée Royal de l'Afrique Centrale: Sciences Zoologiques 110: 9–89.

[R23] Roewer, C.-F. 1923. Die Weberknechte der Erde: Systematische Bearbeitung der bisher bekannnten Opiliones. Gustav Fischer: Jena.

[R35] Roewer, C.-F. 1935. Alte und neue Assamiidae. Weitere Weberknechte VIII. (8. Ergänzung der “Weberknechte der Erde” 1923). Veröffentlichungen aus dem Deutschen Kolonial- und Uebersee-Museum in Bremen 1: 1–168.

[SP10] Santos, R., & C. E. Prieto. 2010. Los Assamiidae (Opiliones: Assamiidae) de Río Muni (Guinea Ecuatorial), con la descripción de ocho nuevas especies. Revista de Biologia Tropical 58 (1): 203–243.

[SG11] Sharma, P. P., & G. Giribet. 2011. The evolutionary and biogeographic history of the armoured harvestmen—Laniatores phylogeny based on ten molecular markers, with the description of two new families of Opiliones (Arachnida). Invertebrate Systematics 25: 106–142.

[S92] Staręga, W. 1992. An annotated check-list of Afrotropical harvestmen, excluding the Phalangiidae (Opiliones). Annals of the Natal Museum 33 (2): 271–336.

[S78] Suzuki, S. 1978. Three harvestmen (Arachnida, Opiliones) from the Bonin Islands. Annotationes Zoologicae Japonenses 51 (3): 179–185.