Belongs within: Hyloidea.
Eleutherodactylus is a very diverse genus of directly developing, mostly small frogs found in Central and South America.
Eleutherodactylus Duméril & Bibron 1841 [incl. Pelorius Hedges 1989; Eleutherodactylini] FG06
|--+--+--E. counouspeus (Schwartz 1964) PW11, FG06 [=Euhyas counouspea FG06]
| | `--+--+--E. inoptatus Barbour 1914 PW11, FG06 [=Pelorius inoptatus FG06]
| | | `--+--E. chlorophenax Schwartz 1976 PW11, FG06 [=Pelorius chlorophenax FG06]
| | | `--E. nortoni Schwartz 1976 PW11, FG06 [=Pelorius nortoni FG06]
| | `--+--+--E. bothroboans PW11
| | | `--E. ruthae Noble 1923 PW11, FG06 [=Pelorius ruthae FG06]
| | `--+--E. hypostenor Schwartz 1965 PW11, FG06 [=Pelorius hypostenor FG06]
| | `--E. parapelates Hedges & Thomas 1987 PW11, FG06 [=Pelorius parapelates FG06]
| `--+--+--E. richmondi Stejneger 1904 PW11, FG06 [=Euhyas richmondi FG06]
| | `--E. unicolor PW11
| `--+--+--+--E. auriculatoides PW11
| | | `--E. patriciae PW11
| | `--+--+--+--E. fowleri PW11
| | | | `--E. lamprotes PW11
| | | `--+--E. sommeri PW11
| | | `--E. wetmorei PW11
| | `--+--+--E. leberi PW11
| | | `--E. melacara PW11
| | `--+--E. varians PW11
| | `--+--E. guantanamera PW11
| | `--E. ionthus PW11
| `--+--+--+--E. minutus Noble 1923 PW11, FG06 [=Euhyas minuta FG06]
| | | `--E. poolei PW11
| | `--+--+--E. pituinus PW11
| | | `--+--E. parabates Schwartz 1964 PW11, FG06 [=Euhyas parabates FG06]
| | | `--+--E. abbotti PW11
| | | `--+--E. audanti PW11
| | | `--E. haitianus PW11
| | `--+--E. eileenae PW11
| | `--+--E. glamyrus PW11
| | `--+--+--E. mariposa PW11
| | | `--E. ronaldi PW11
| | `--+--E. bartonsmithi PW11
| | `--+--E. auriculatus PW11
| | `--E. principalis PW11
| `--+--+--+--E. barlagnei PW11
| | | `--E. pinchoni PW11
| | `--+--E. johnstonei Barbour 1914 PW11, B-AC-O04
| | `--+--E. amplinympha PW11
| | `--E. locustus PW11
| `--+--+--E. antillensis PW11
| | `--+--E. brittoni PW11
| | `--+--E. cochranae PW11
| | `--E. hedricki PW11
| `--+--E. gryllus PW11
| `--+--+--E. coqui PW11
| | `--E. schwartzi PW11
| `--+--E. portoricensis PW11
| `--E. wightmanae PW11
`--+--+--+--E. symingtoni Schwartz 1957 PW11, FG06 [=Euhyas symingtoni FG06]
| | `--E. zeus Schwartz 1958 PW11, FG06 [=Euhyas zeus FG06]
| `--+--E. marnockii (Cope 1878) PW11, FG06 [=E. (Syrrhophus) marnockii FG06]
| `--+--E. nitidus Peters 1870 PW11, FG06 [=E. (Syrrhophus) nitidus FG06]
| `--E. pipilans (Taylor 1940) PW11, FG06 [=Syrrhophus pipilans FG06]
`--+--+--+--E. klinikowskii Schwartz 1959 PW11, FG06 [=Euhyas klinikowskii FG06]
| | `--E. zugi Schwartz 1958 PW11, FG06 [=Euhyas zugi FG06]
| `--+--+--+--E. brevirostris Shreve 1936 PW11, FG06 [=Craugastor brevirostris FG06]
| | | `--E. ventrilineatus Shreve 1936 PW11, FG06 [=Euhyas ventrilineata FG06]
| | `--+--E. glandulifer Cochran 1935 PW11, FG06 [=Euhyas glandulifer FG06]
| | `--E. sciagraphus Schwartz 1973 PW11, FG06 [=Euhyas sciagraphus FG06]
| `--+--E. paulsoni Schwartz 1964 PW11, FG06 [=Euhyas paulsoni FG06]
| `--+--+--E. furcyensis Shreve & Williams 1963 PW11, FG06 [=Euhyas furcyensis FG06]
| | `--E. rufifemoralis PW11
| `--+--+--E. apostates Schwartz 1973 PW11, FG06 [=Euhyas apostates FG06]
| | `--E. oxyrhyncus Duméril & Bibron 1841 PW11, FG06 [=Euhyas oxyrhyncha FG06]
| `--+--E. jugans Cochran 1937 PW11, FG06 [=Euhyas jugans FG06]
| `--+--E. glanduliferoides Shreve 1936 PW11, FG06 [=Euhyas glanduliferoides FG06]
| `--+--E. thorectes Hedges 1988 PW11, FG06 [=Euhyas thorectes FG06]
| `--+--+--E. bakeri PW11
| | `--+--E. dolomedes Hedges & Thomas 1992 PW11, FG06 [=Euhyas dolomedes FG06]
| | `--E. glaphycompus Schwartz 1973 PW11, FG06 [=Euhyas glaphycompus FG06]
| `--+--E. heminota PW11
| `--+--E. corona Hedges & Thomas 1992 PW11, FG06 [=Euhuyas corona FG06]
| `--+--E. amadeus Hedges, Thomas & Franz 1987 PW11, FG06 (see below for synonymy)
| `--+--E. caribe Hedges & Thomas 1992 PW11, FG06 [=Euhyas caribe FG06]
| `--E. eunaster PW11
`--+--+--E. schmidti Noble 1923 PW11, FG06 [=Euhyas schmidti FG06]
| `--+--E. albipes Barbour & Shreve 1937 PW11, FG06 [=Euhyas albipes FG06]
| `--+--E. maestrensis Diaz, Cádiz & Navarro 2005 PW11, FG06 [=Euhyas maestrensis FG06]
| `--+--E. dimidiatus Cope 1862 PW11, FG06 [=Euhyas dimidiatus FG06]
| `--E. emiliae Dunn 1926 PW11, FG06 [=Euhyas emiliae FG06]
`--+--E. greyi Dunn 1926 PW11, FG06 [=Euhyas greyi FG06]
`--+--+--E. atkinsi Dunn 1925 PW11, FG06 [=Euhyas atkinsi FG06]
| |--+--+--E. guanahacabibes Estrada & Rodriguez 1985 PW11, FG06 [=Euhyas guanahacabibes FG06]
| | | `--+--E. casparii Dunn 1926 PW11, FG06 [=Euhyas casparii FG06]
| | | `--E. planirostris Cope 1862 PW11, FG06 [=E. (Euhyas) planirostris FG06]
| | `--+--+--E. goini Schwartz 1960 PW11, FG06 [=Euhyas goini FG06]
| | | `--+--E. rogersi PW11
| | | `--E. tonyi Estrada & Hedges 1997 PW11, FG06 [=Euhyas tonyi FG06]
| | `--+--E. pezopetrus PW11
| | `--+--E. pinarensis PW11
| | `--+--E. blairhedgesi PW11
| | `--E. thomasi Schwartz 1959 PW11, FG06 [=Euhyas thomasi FG06]
| `--+--+--E. gundlachi Schmidt 1920 PW11, FG06 [=Euhyas gundlachi FG06]
| | `--+--E. intermedius Barbour & Shreve 1937 PW11, FG06 [=Euhyas intermedia FG06]
| | `--E. varleyi Dunn 1925 PW11, FG06 [=Euhyas varleyi FG06]
| `--+--+--E. cubanus Barbour 1942 PW11, FG06 [=Euhyas cubana FG06]
| | `--E. orientalis Barbour & Shreve 1937 PW11, FG06 [=Euhyas orientalis FG06]
| `--+--E. etheridgei Schwartz 1958 PW11, FG06 [=Euhyas etheridgei FG06]
| `--+--E. iberia Estrada & Hedges 1996 PW11, FG06 [=Euhyas iberia FG06]
| `--+--E. jaumei Estrada & Alonso 1997 PW11, FG06 [=Euhyas jaumei FG06]
| `--E. limbatus Cope 1862 PW11, FG06 [=Euhyas limbata FG06]
`--+--+--+--E. alcoae Schwartz 1971 PW11, FG06 [=Euhyas alcoae FG06]
| | `--E. armstrongi Noble & Hassler 1933 PW11, FG06 [=Euhyas armstrongi FG06]
| `--+--E. darlingtoni PW11
| `--E. leoncei Shreve & Williams 1963 PW11, FG06 [=Euhyas leoncei FG06]
`--+--+--+--E. bresslerae PW11
| | `--+--E. acmonis Schwartz 1960 PW11, FG06 [=Euhyas acmonis FG06]
| | `--E. ricordii Duméril & Bibron 1841 PW11, FG06 [=E. (Euhyas) ricordii FG06]
| `--+--E. probolaeus PW11
| `--+--E. monensis Meerwarth 1901 PW11, FG06 [=Euhyas monensis FG06]
| `--+--E. grahami Dunn 1926 PW11, FG06 [=Euhyas grahami FG06]
| `--+--E. lentus PW11
| `--+--E. rhodesi PW11
| `--E. weinlandi Barbour 1914 PW11, FG06 [=Euhyas weinlandi FG06]
`--+--+--E. cuneatus Cope 1862 PW11, FG06 (see below for synonymy)
| `--E. turquinensis Barbour & Shreve 1937 PW11, FG06 [=Euhyas turquinensis FG06]
`--+--+--E. riparius PW11
| `--E. rivularis Díaz, Estrada & Hedges 2001 PW11, DEH01 [=E. (Euhyas) rivularis DEH01]
`--+--E. toa Estrada & Hedges 1991 PW11, FG06 [=Euhyas toa FG06]
`--+--E. luteolus Gosse 1851 PW11, FG06 [=Euhyas luteola FG06]
`--+--+--+--E. sisyphodemus Crombie 1977 PW11, FG06 [=Euhyas sisyphodemus FG06]
| | `--+--E. cavernicola Lynn 1954 PW11, FG06 [=Euhyas cavernicola FG06]
| | `--E. grabhami Dunn 1926 PW11, FG06 [=Euhyas grabhami FG06]
| `--+--E. jamaicensis Barbour 1910 PW11, FG06 [=Euhyas jamaicensis FG06]
| `--+--E. nubicola Dunn 1926 PW11, FG06 [=E. (Euhyas) nubicola FG06]
| `--+--E. andrewsi Lynn 1937 PW11, FG06 [=Euhyas andrewsi FG06]
| `--E. orcutti Dunn 1928 PW11, FG06 [=Euhyas orcutti FG06]
`--+--E. alticola Lynn 1937 PW11, FG06 [=Euhyas alticola FG06]
`--+--E. fuscus Lynn & Dent 1943 PW11, FG06 [=Euhyas fusca FG06]
`--+--E. junori Dunn 1926 PW11, FG06 [=Euhyas junori FG06]
`--+--E. gossei Dunn 1926 PW11, FG06 [=Euhyas gossei FG06]
`--+--E. griphus Crombie 1986 PW11, FG06 [=Euhyas griphus FG06]
`--+--+--E. cundalli PW11
| `--E. glaucoreius Schwartz & Fowler 1973 PW11, FG06 (see below for synonymy)
`--+--E. pantoni Dunn 1926 PW11, FG06 [=Euhyas pantoni FG06]
`--E. pentasyringos Schwartz & Fowler 1973 PW11, FG06 (see below for synonymy)
Eleutherodactylus incertae sedis:
E. angustidigitorum Taylor 1940 NW07, FG06 [=Syrrhophus angustidigitorum FG06]
E. anolirex Lynch 1983 B-AC-O04
E. (Eleutherodactylus) binotatus FG06
E. boconoensis Rivero & Mayorga 1973 B-AC-O04
E. chloronotus FG06
E. chlorosoma Rivero 1984 B-AC-O04
E. colostichos La Marca & Smith 1982 B-AC-O04
E. conspicillatus B-AC-O04
E. costaricensis M83
E. diastema FG06
E. discoidalis FG06
E. duellmani FG06
E. erythropleurus NW07
E. ginesi (Rivero 1964) B-AC-O04
E. guentheri NW07
E. hedriki SWK87
E. jasperi FG06
E. karlschmidti Grant 1931 NW07, FG06 [=Euhyas karlschmidti FG06]
E. lancinii Donoso-Barros 1968 B-AC-O04
E. lentiginosus Rivero 1984 B-AC-O04
E. malkini B-AC-O04
E. megalops B-AC-O04
E. melanoproctus Rivero 1984 B-AC-O04
E. mercedesae FG06
E. mondolfii Rivero 1984 B-AC-O04
E. nicefori Cochran & Goin 1971 B-AC-O04
E. nigrovittatus FG06
E. paramerus Rivero 1984 B-AC-O04
E. parvus NW07
E. pedimontanus La Marca 2004 B-AC-O04
E. pleurostriatus Rivero 1984 B-AC-O04
E. prolixodiscus Lynch 1978 B-AC-O04
E. pulidoi Rivero 1984 B-AC-O04
E. rearki M83
E. ridens M81
E. supernatis FG06
E. thymelensis FG06
E. tubernasus Rivero 1984 B-AC-O04
E. tubolus KF69
E. underwoodi M83
E. unistrigatus B-AC-O04
E. urichi (Boettger 1894) B-AC-O04
E. vanadisae La Marca 1984 B-AC-O04
E. w-nigrum B-AC-O04
E. yustizi Barrio-Amorós & Chacón-Ortiz 2004 B-AC-O04
Eleutherodactylus amadeus Hedges, Thomas & Franz 1987 PW11, FG06 [=Euhyas amadeus FG06]
Eleutherodactylus cuneatus Cope 1862 PW11, FG06 [=Euhyas cuneata FG06; incl. El. sierramaestrae DEH01]
Eleutherodactylus glaucoreius Schwartz & Fowler 1973 PW11, FG06 [=Euhyas glaucoreia FG06]
Eleutherodactylus pentasyringos Schwartz & Fowler 1973 PW11, FG06 [=Euhyas pentasyringos FG06]
*Type species of generic name indicated
REFERENCES
[B-AC-O04] Barrio-Amorós, C. L., & A. Chacón-Ortiz. 2004. Un nuevo Eleutherodactylus (Anura, Leptodactylidae) de la Cordillera de Mérida, Andes de Venezuela. Graellsia 60 (1): 3–11.
[DEH01] Díaz, L. M., A. R. Estrada & S. B. Hedges. 2001. A new riparial frog of the genus Eleutherodactylus (Anura: Leptodactylidae) from eastern Cuba. Caribbean Journal of Science 37 (1–2): 63–71.
[FG06] Frost, D. R., T. Grant, J. Faivovich, R. H. Bain, A. Haas, C. F. B. Haddad, R. O. de Sá, A. Channing, M. Wilkinson, S. C. Donnellan, C. J. Raxworthy, J. A. Campbell, B. L. Blotto, P. Moler, R. C. Drewes, R. A. Nussbaum, J. D. Lynch, D. M. Green & W. C. Wheeler. 2006. The amphibian tree of life. Bulletin of the American Museum of Natural History 297: 1–370.
[KF69] Kluge, A. G., & J. S. Farris. 1969. Quantitative phyletics and the evolution of anurans. Systematic Zoology 18 (1): 1–32.
[M81] Miyamoto, M. M. 1981. Congruence among character sets in phylogenetic studies of the frog genus Leptodactylus. Systematic Zoology 30 (3): 281–290.
[M83] Miyamoto, M. M. 1983. Biochemical variation in the frog Eleutherodactylus bransfordii: geographic patterns and cryptic species. Systematic Zoology 32 (1): 43–51.
[NW07] Nussbaum, R. A., & S.-H. Wu. 2007. Morphological assessments and phylogenetic relationships of the Seychellean frogs of the family Sooglossidae (Amphibia: Anura). Zoological Studies 46 (3): 322–335.
[SWK87] Snyder, N. F. R., J. W. Wiley & C. B. Kepler. 1987. The Parrots of Luquillo: Natural history and conservation of the Puerto Rican parrot. Western Foundation of Vertebrate Zoology: Los Angeles.
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