Termitonannini

Termitonannus vagans, from Seevers (1957).

Belongs within: Staphylinidae.

The Termitonannini are a group of mostly limuloid staphylinid beetles found in association with nasutitermitine termites (Seevers 1957).

Characters (from Seevers 1957): Body usually non-physogastric and limuloid, very rarely physogastric. Head either prognathous or hypognathous; antennae 9-11-segmented, terminal antennal segments bearing one or more pairs of polytrichous coeloconic sensilla; mentum free except in certain derived genera. Mesocoxal cavities margined; hind coxae and metepimera generalised except in certain derived genera. Tarsi 4, 4, 4-segmented or 4, 4, 5-segmented.

<==Termitonannini S57
    |--Termitonannina S57
    |    |  i. s.: Termitocomes Seevers 1941 S57
    |    |           |--*T. wasmanni Seevers 1941 S57
    |    |           `--T. trinidadensis Seevers 1957 S57
    |    |--+--Eunannodes Silvestri 1946 S57
    |    |  |    `--*E. reconditi Silvestri 1946 S57
    |    |  `--Nannellus Silvestri 1946 S57
    |    |       `--*N. anoplotermitis Silvestri 1946 S57
    |    `--+--Termitonilla Borgmeier 1950 S57
    |       |    `--*T. luteola Borgmeier 1950 S57
    |       `--+--Macrotrichuris Silvestri 1946 S57
    |          |    |--*M. brasiliensis Silvestri 1946 S57
    |          |    `--M. notabilis Silvestri 1946 S57
    |          `--Termitonannus Wasmann 1902 [incl. Tetraphilus Silvestri 1946] S57
    |               |--*T. schmalzi Wasmann 1902 S57
    |               |--T. brachycerus Silvestri 1946 [=T. (*Tetraphilus) brachycerus] S57
    |               |--T. domunculi Silvestri 1946 S57
    |               |--T. echinoides Seevers 1946 S57
    |               |--T. gatuni Seevers 1941 S57
    |               |--T. major Wasmann 1902 S57
    |               |--T. microsomatis Silvestri 1946 S57
    |               |--T. parvulus Silvestri 1946 S57
    |               |--T. proximatus Silvestri 1946 S57
    |               |--T. setosus Seevers 1941 S57
    |               |--T. silvestrii Wasmann 1902 S57
    |               `--T. vagans Silvestri 1946 S57
    `--Perinthina [Perinthini] S57
         |--Termitopelta Borgmeier 1950 S57
         |    `--*T. fulgens Borgmeier 1950 S57
         `--+--Termitonicus Mann 1926 S57
            |    |--*T. mahout Mann 1926 S57
            |    `--T. uroclaviger Silvestri 1946 S57
            `--+--Macrognathellus Silvestri 1946 S57
               |    `--M. crassicornis (Wasmann 1902) (see below for synonymy) S57
               `--+--Poduroides Mann 1926 S57
                  |    `--*P. bovingi Mann 1926 S57
                  `--+--Perinthus Casey 1890 S57
                     |    |--*P. dudleyanus Casey 1890 S57
                     |    |    |--P. d. dudleyanus S57
                     |    |    |--P. d. major Seevers 1957 S57
                     |    |    `--P. d. wasmanni Mann 1923 S57
                     |    |--P. bolivari Seevers 1946 S57
                     |    |--P. guatemalae Seevers 1946 S57
                     |    |--P. hageni Seevers 1937 S57
                     |    |--P. mayae S57
                     |    |--P. tarsatus Mann 1923 S57
                     |    `--P. xenocostalis Seevers 1937 S57
                     `--+--Termitocola Seevers 1937 S57
                        |    |--*T. cylindricornis Seevers 1937 S57
                        |    `--T. silvestrii (Wasmann 1902) [=Perinthus silvestrii] S57
                        `--+--Lauella Mann 1921 S57
                           |    |--*L. vitiensis Mann 1921 [incl. Hetairotermes leai Cameron 1927] S57
                           |    |--L. javana Seevers 1957 S57
                           |    `--L. palauensis Seevers 1957 S57
                           `--+--Eutermitophila Cameron 1939 S57
                              |    `--*E. fletcheri Cameron 1939 S57
                              `--+--Perinthodes Seevers 1957 S57
                                 |    `--*P. africanus Seevers 1957 S57
                                 `--Paraperinthus Seevers 1957 S57
                                      `--*P. pauciseta Seevers 1957 S57

Macrognathellus crassicornis (Wasmann 1902) [=Perinthus crassicornis; incl. *M. paraguayensis Silvestri 1946] S57

*Type species of generic name indicated

REFERENCES

[S57] Seevers, C. H. 1957. A monograph on the termitophilous Staphylinidae (Coleoptera). Fieldiana Zoology 40: 1–334.