Coccophagus

Ovipositing female of Coccophagus gurneyi, photographed by Max Badgley.

Belongs within: Coccophaginae.

Coccophagus is a cosmopolitan genus of small wasps including parasitoids of scale insects. It is distinguished from many related genera by its relatively setose thorax, while it and the antennae are usually not markedly (Hayat 1983).

Characters (from Hayat 1983): Body moderately setose; antenna usually 8-segmented (1, 1, 3, 3), sometimes 7-segmented with 2-segmented clava; antennal scape not or only slightly flattened and not or hardly expanded beneath; scutellum with at least six setae, each axillae usually with at least two setae (rarely with just one); axillae large, subtriangular, strongly projecting forwards; separated mesally by about the maximum length of an axilla; scutellum not extending over base of gaster and metanotum without a membranous extension overlapping tergum I of gaster (several species have a thin transparent extension of the metanotum, which partly overlaps the propodeum in the middle); submarginal vein not enlarged; linea calva absent; hypopygium not pointed at apex and not extending to apex of gaster.
Coccophagus Westwood 1833 GM79 [incl. Ataneostigma B88; Coccophagini]
    |--C. aethiopis Girault 1935 [=Prospaltella berlesei Silvestri 1930 non (Howard 1906)] G35
    |--C. antiopa Girault 1915 GM79
    |--C. argentifascia Girault 1915 GM79
    |--C. auricaput Girault 1915 GM79
    |--C. basalis A71
    |--C. bivittatus A71
    |--C. brahmsi Girault 1933 G33b
    |--C. brimblecombei Girault 1933 G33a
    |--C. capensis A71
    |--C. caridei A71
    |--C. coccidis Girault 1917 G17
    |--C. cowperi Girault 1917 G17
    |--C. crucigerus Girault 1931 G31
    |--C. exiguiventris Girault 1929 G29
    |--C. flavoscutellum Ashmead 1881 GM79
    |--C. grotei Girault 1931 G31
    |--C. gurneyi A71
    |--C. herndoni Girault 1935 [=Prospaltella aurantii Silvestri 1930 non (Howard 1894)] G35
    |--C. immaculatus Howard 1881 GM79
    |--C. inkermani Girault 1926 G26b
    |--C. inlacertus Girault 1931 G31
    |--C. insidiator A71
    |--C. iris Girault 1930 G30b
    |--C. lecanii Fitch 1858 GM79
    |--C. leucippi Girault 1936 G36b
    |--C. longifasciatus A71
    |--C. lucani Girault 1922 GM79
    |--C. lunulatus Howard 1894 GM79
    |--C. lycimnia A71
    |--C. malthusi Girault 1917 G17
    |--C. maria Girault 1931 G31
    |--C. marxi Girault 1936 G36a
    |--C. mexicensis Girault 1917 G17
    |--C. modesta Silvestri 1914 GM79
    |--C. multisetae Girault 1931 G31
    |--C. nigritus A71
    |--C. nigriventris (Girault 1913) GM79
    |--C. nigropleurum Girault 1917 G17
    |--C. ochraceus A71
    |--C. perhispidus Westwood 1833 GM79
    |--C. pulcini Girault 1926 G26b
    |--C. quaestor Girault 1917 G17
    |--C. redini Girault 1924 G24
    |--C. rusti MH11
    |--C. saintebeauvei Girault 1917 G17
    |--C. sanctus Girault 1928 G28
    |--C. scutellaris A71
    |--C. seminigriclavus Girault 1913 GM79
    |--C. shakespearella Girault 1929 G29
    |--C. signus Girault 1920 GM79
    |--C. silvifilia Girault 1924 G24
    |--C. socratis Girault 1931 G31
    |--C. sophia Girault & Dodd 1915 GM79
    |--C. submetallicus Girault 1930 G30b
    |--C. subsignus Girault 1932 G32b
    |--C. taciti Girault 1930 G30a
    |--C. tennysoni Girault 1931 G31
    |--C. thoreauini Girault 1915 GM79
    |--C. totiaureus Girault 1930 G30a
    |--C. triangulatinotus Girault 1926 G26b
    |--C. triguttatus Girault 1915 GM79
    |--C. tristis G26a
    |--C. unifasciata Girault 1915 GM79
    `--C. vegai Girault 1932 G32a

*Type species of generic name indicated

REFERENCES

[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[G17] Girault, A. A. 1917. Descriptiones stellarum novarum. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 80-101).

[G24] Girault, A. A. 1924. Homo perniciosus and new Hymenoptera. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 178-181).

[G26a] Girault, A. A. 1926a. New pests from Australia. II. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 200-202).

[G26b] Girault, A. A. 1926b. New pests from Australia. III. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 203-204).

[G28] Girault, A. A. 1928. Some Insecta and a new All Highness. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 225-228).

[G29] Girault, A. A. 1929. Description of a case of lunacy in Homo and of six-legged articulates. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 262-265).

[G30a] Girault, A. A. 1930a. New pests from Australia. VII. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 270-272).

[G30b] Girault, A. A. 1930b. New pests from Australia. VIII. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 273-277).

[G31] Girault, A. A. 1931. A new habit in an old insect, Homo pudicus and new Eurytomidae. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 280-283).

[G32a] Girault, A. A. 1932a. New pests from Australia, X. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 286-291).

[G32b] Girault, A. A. 1932b. New lower Hymenoptera from Australia and India. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 293-298).

[G33a] Girault, A. A. 1933a. Some beauties inhabitant not of commercial boudoirs but of Nature’s bosom, notably new insects. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 299-303).

[G33b] Girault, A. A. 1933b. Some beauties inhabitant not of the boudoirs of commerce but of Nature’s bosom—new insects. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 304-305).

[G35] Girault, A. A. 1935. Microhymenoptera Australiensis nova, mostly Chalcididae. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 315-318).

[G36a] Girault, A. A. (as ‘L. de St. Fargeau’). 1936a. Chalcididae, Capsidae species nova Australiensis Giraulti. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 319-321).

[G36b] Girault, A. A. 1936b. Terror-errors; and novitates of Pterygota (or earth realities not state-bound). Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 322-325).

[GM79] Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 1-400.

Hayat, M. 1983. The genera of Aphelinidae (Hymenoptera) of the world. Systematic Entomology 8: 63-102.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.