Raninoida

Spanner crab Ranina ranina, photographed by Massimo Boyer.

Belongs within: Brachyura.
Contains: Etyidae.

The Raninoida or Archaeobrachyura are a group of crabs first recorded from the Cretaceous. They are characterised by the complete loss of the uropods, without even a modified remnant (Guinot & Tavares 2001). The Raninoidea camouflage themselves by burrowing into sediment, whereas the Cyclodorippoidea may bury themselves or carry debris for concealment.

<==Raninoida [Archaeobrachyura, Gymnopleura]
    |  i. s.: Etyidae GT01
    |--Dakoticancridae [Dakoticancroidea] GT01
    |    |--Avitelmessus grapsoideus Rathbun 1923 GT01
    |    |--Tetracarcinus Weller 1905 CF06
    |    |--Seorsus Bishop 1988 GT01
    |    `--Dakoticancer Rathbun 1917 CF06
    |         `--D. overanus Rathbun 1917 GT01
    |--Cyclodorippoidea [Tymoloidea] MD01
    |    |--Phyllotymolinum GT01 [Phyllotymolinidae MD01]
    |    |    `--P. crosnieri Tavares 1993 GT01
    |    |--Cymonomidae MD01
    |    |    |--Elassopodus stellatus Tavares 1993 GT01
    |    |    `--Cymonomus MG-H11
    |    |         |--C. andamanicus Alcock 1905 MG-H11
    |    |         `--C. leblondi Tavares 1994 GT01
    |    `--Cyclodorippidae MD01
    |         |--Krangalangia spinosa (Zarenkov 1970) MG-H11
    |         |--Tymolus [Tymolidae] MG-H11
    |         |    |--T. brucei Tavares 1991 MG-H11
    |         |    `--T. similis (Grant 1905) MG-H11
    |         `--Corycodus GT01
    |              |--C. decorus Tavares 1993 GT01
    |              `--C. mervae Tavares 1993 GT01
    `--Raninoidea MD01
         |--Symethis [Symethidae, Symethinae] MD01
         `--Raninidae CF06
              |--Notosceles serratifrons (Henderson 1893) MG-H11
              |--Umalia trirufomaculata (Davie & Short 1989) MG-H11
              |--Cosmonotus Adams & White 1847 B64
              |    |--C. eocaenicus Beschin et al. 1988 B04
              |    `--C. grayi Adams & White 1848 MG-H11
              |--Ranina Lamarck 1801 B02
              |    `--*R. ranina (Linnaeus 1758) (see below for synonymy) B02
              |--Notopus de Haan 1841 B02
              |    `--N. dorsipes (Fabricius 1787) (see below for synonymy) B02
              |--Raninella Milne Edwards 1862 CF06
              |    |--*R. trigeri Milne Edwards 1862 CF06
              |    |--R. carlilensis Feldmann & Maxey 1980 CF06
              |    |--R. eocenica Rathbun 1935 CF06
              |    |--R. libyca Van Straelen 1935 CF06
              |    |--R. mucronata Rathbun 1935 CF06
              |    |--R. oaheensis Bishop 1978 CF06
              |    |--R. quadrispinosus (Collins, Fraaye & Jagt 1995) [=Raninoides quadrispinosus] CF06
              |    |--R. trechmanni (Withers 1927) [=Ranina trechmanni] CF06
              |    `--R. tridens Roberts 1962 CF06
              |--Lyreidus de Haan 1841 B64
              |    |--L. alseanus B64
              |    |--L. australiensis Ward 1933 B64
              |    |--L. bairdi Smith 1888 B64
              |    |--L. elegans B64
              |    |--L. elongatus Miers 1879 B64
              |    |--L. fastigiatus Rathbun 1918 B64
              |    |--L. fossor Bennett 1964 B64
              |    |--L. channeri Wood-Mason 1885 [incl. L. gracilis Wood-Mason 1887] B64
              |    |--L. integra B64
              |    |--L. politus Parisi 1914 B64
              |    |--L. stenops Wood-Mason 1887 MG-H11
              |    `--L. tridentatus De Haan 1841 MG-H11
              |--Erroranilia Boyko 2004 [=Paralbunea Hu & Tao 1996 non Serène 1977] B04
              |    `--*E. taipeiensis (Hu & Tao 1996) [=*Paralbunea taipeiensis, P. taipeinensis] B04
              `--Ranilia Milne Edwards 1837 B04
                   `--R. punctulata Beschin et al. 1988 B04

Notopus dorsipes (Fabricius 1787) [=Hippa dorsipes, Albunea dorsipes, A. dorsipede (l. c.), Cancer dorsipes] B02

*Ranina ranina (Linnaeus 1758) [=Cancer raninus; incl. Hippa scabra Fabricius 1787, Albunea scabra, Cancer scaber] B02

*Type species of generic name indicated

REFERENCES

[B64] Bennett, E. W. 1964. The marine fauna of New Zealand: Crustacea Brachyura. New Zealand Department of Scientific and Industrial Research Bulletin 153: 1–120.

[B02] Boyko, C. B. 2002. A worldwide revision of the recent and fossil sand crabs of the Albuneidae Stimpson and Blepharipodidae, new family (Crustacea: Decapoda: Anomura: Hippoidea). Bulletin of the American Museum of Natural History 272: 1–396.

[B04] Boyko, C. B. 2004. The problematic Paralbunea Hu and Tao, 1996: homonymy, generic nom. nov., and correct taxonomic placement. Palaeontology 47 (4): 929–932.

[CF06] Crawford, R. S., R. F. Feldmann, D. A. Waugh, B. M. Kelley & J. G. Allen. 2006. Decapod crustaceans from the Maastrichtian Fox Hills formation. Bulletin of the Peabody Museum of Natural History 47 (1–2): 3–28.

[GT01] Guinot, D., & M. Tavares. 2001. Une nouvelle famille de crabes du Crétacé, et la notion de Podotremata Guinot, 1977 (Crustacea, Decapoda, Brachyura). Zoosystema 23 (3): 507–546.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1–124.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O'Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.