Doryctinae

Siragra sp., copyright Stephane Hanot.

Belongs within: Ichneumonoidea.

The Doryctinae are a cosmopolitan group of braconid wasps with ectoparasitic larvae, most commonly on concealed Coleoptera larvae. The Neotropical Ypsistocerini, distinguished from other tribes by an enlarged telotarsus, are known from termite nests but their biology is otherwise unknown (Achterberg 1993).

Characters (from Achterberg 1993): Maxillary palp with six segments; hypoclypeal depression deep and wide, middle of apparent ventral margin of clypeus distinctly above upper level of mandibular bases, labrum concave; eyes wider than base of mandible; posterior flange of propleuron present, largely dorsally situated; fore tibia often with row of stout pegs or spines, at most as long as about six times their length; hind coxa often angulate anteroventrally, frequently with a ventral tubercle; ovipositor dorsally nearly always with double nodus subapically.

<==Doryctinae A93
    |  i. s.: Sericobracon R00
    |         Stenocorse Marsh 1968 A93
    |         Monolexis N91
    |         ‘Atanycolus’ tomentosus QI93
    |--Termitobracon [Ypsistocerinae, Ypsistocerini] A93
    |    `--T. emersoni A93
    |--Evaniodes [Evaniodini] A93
    |    `--E. areolatus A93
    |--Pareucorystes [Hecabolini] A93
    |    `--P. varinervis A93
    |--Leptorhaconotus [Leptorhaconotini] A93
    |    `--L. brunneus A93
    |--Spathiini A93
    |    |--Spathioplites phreneticus A93
    |    `--Spathius clavatus [=Bracon (Spathius) clavatus] G20
    |--Syngastrini A93
    |    |--Syngaster A93
    |    `--Siragra nitida A93
    `--Doryctini A93
         |--Priosphys biannulatus A93
         |--Doryctes erythromelas HR11
         `--Dendrosoter Wesmael 1838 A93, M38 [incl. Caenopachys Foerster 1862 M38, Eurybolus Ratzeburg 1848 M38]
              |--*D. protuberans [=Bracon protuberans; incl. B. curtisii, *Eurybolus curtisii] M38
              |--D. hartigii [=Bracon hartigii, *Caenopachys hartigii] M38
              |--D. integer Muesebeck 1938 M38
              |--D. middendorffii M38
              |--D. scaber Muesebeck 1938 M38
              |--D. scolytivorus (Viereck & Rohwer 1913) [=Caenopachys scolytivora] M38
              `--D. sulcatus Muesebeck 1938 M38

*Type species of generic name indicated

REFERENCES

[A93] Achterberg, C. van. 1993. Illustrated key to the subfamilies of the Braconidae (Hymenoptera: Ichneumonoidea). Zoologische Verhandelingen 283: 1–189.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[HR11] Heraty, J., F. Ronquist, J. M. Carpenter, D. Hawks, S. Schulmeister, A. P. Dowling, D. Murray, J. Munro, W. C. Wheeler, N. Schiff & M. Sharkey. 2011. Evolution of the hymenopteran megaradiation. Molecular Phylogenetics and Evolution 60: 73–88.

[M38] Muesebeck, C. F. W. 1938. The genus Dendrosoter Wesmael in the United States (Hymenoptera: Braconidae). Proceedings of the Entomological Society of Washington 40 (9): 281–287.

[N91] Naumann, I. D. 1991. Hymenoptera (wasps, bees, ants, sawflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2^nd ed. vol. 2 pp. 916–1000. Melbourne University Press: Carlton (Victoria).

[QI93] Quicke, D. L. J., & S. N. Ingram. 1993. Braconine wasps of Australia. Memoirs of the Queensland Museum 33 (1): 299–336.

[R00] Ross, E. S. 2000. Embia: Contributions to the biosystematics of the insect order Embiidina. Occasional Papers of the California Academy of Sciences 149: 1–53, 1–36.

Last updated: 7 March 2021.