Thelyphoninae

Thelyphonus sp., copyright Arthur Anker.

Belongs within: Uropygi.

The Thelyphoninae is a group of whipscorpions found in south-east Asia and Melanesia (Rowland & Cooke 1973).

Characters (from Rowland & Cooke 1973): Tarsal segments of first leg of female usually modified (except in Minbosius manilanus and Abaliella rohdei), variously incrassate or apophysate; pedipalpal patellar apophysis of male unmodified, occasionally longer than in female; abdominal sternites II and III modified; two, four or no ommatoids present; keel present between lateral and median eyes; posterior abdominal tergites divided.

<==Thelyphoninae [Thelyphonini]
    |--Minbosius Speijer 1936 RC73
    |    `--*M. manilanus (Koch 1843) RC73
    |--Chajnus Speijer 1936 RC73
    |    `--*C. renschi Speijer 1936 RC73
    |--Glyptogluteus Rowland 1973 RC73
    |    `--*G. augustus Rowland 1973 RC73
    |--Ginosigma Speijer 1936 RC73
    |    |--*G. schimkewitschi (Tarnani 1894) RC73
    |    `--G. lombokensis Speijer 1936 RC73
    |--Tetrabalius Thorell 1888 RC73, H02
    |    |--*T. seticauda (Doleschall 1857) RC73
    |    `--T. nasutus Thorell 1889 RC73
    |--Abaliella Strand 1928 [=Abalius Kraepelin 1897 (preoc.)] RC73
    |    |--*A. rohdei (Kraepelin 1897) RC73
    |    |--A. dicranotarsalis Rowland 1973 RC73
    |    |--A. gertschi Rowland 1973 RC73
    |    |--A. manilana (Kraepelin 1901) RC73
    |    |--A. samoana (Kraepelin 1879) RC73
    |    `--A. willeyi (Pocock 1898) RC73
    `--Thelyphonus Latreille 1802 H02 [=Telyphonus (l. c.) KK84]
         |--*T. caudatus (Linnaeus 1758) RC73 [=Phalangium caudatum H02; incl. T. thorellii Tarnani 1890 T94]
         |--T. anthracinus Pocock 1894 RC73
         |--T. antillanus T94
         |--T. asperatus Thorell 1888 RC73
         |--T. assamensis Stoliczka 1869 [=Telyphonus assamensis] S69
         |--T. billitonensis Speijer 1931 RC73
         |--T. borneensis Kraepelin 1897 RC73
         |--T. burchardi Kraepelin 1912 RC73
         |--T. celebensis Kraepelin 1897 RC73
         |--T. doriae Thorell 1888 RC73
         |--T. grandis Speijer 1931 RC73
         |--T. hanseni Kraepelin 1897 RC73
         |--T. ignotus T94
         |--T. insulanus Koch & Keyserling 1884 RC73, KK84
         |--T. javanus T94
         |--T. kinabaluensis Speijer 1933 RC73
         |--T. klugi Kraepelin 1897 RC73
         |--T. lawrencei Rowland 1973 RC73
         |--T. leucurus Pocock 1898 RC73
         |--T. linganus Koch 1843 RC73
         |--T. pococki Tarnani 1901 RC73
         |--T. proscorpius T94
         |--T. rufipes [=Telyphonus rufipes] D57
         |--T. schnehageni Kraepelin 1897 RC73
         |--T. semperi Kraepelin 1897 RC73
         |--T. sepiaris Butler 1873 RC73 [incl. T. nigrescens Butler 1873 T89]
         |--T. seticauda Doleschall 1857 [=Telyphonus seticauda] D57
         |--T. spinimanus [=Telyphonus spinimanus] S69
         |--T. stimpsonii Wood 1861 [=Telyphonus stimpsonii] S69
         |--T. sucki Kraepelin 1897 RC73
         |--T. sumatranus Kraepelin 1897 RC73
         |--T. vanoorti Speijer 1936 RC73
         `--T. wayi Pocock 1900 RC73

*Type species of generic name indicated

REFERENCES

[D57] Doleschall, C. L. 1857. Bijdrage tot de kennis der Arachniden van den Indischen Archipel. Natuurkundig Tijdschrift voor Nederlandsch Indië, series 3, 3 (5–6): 399–434, plates 1–2.

[H02] Harvey, M. S. 2002. The neglected cousins: What do we know about the smaller arachnid orders? Journal of Arachnology 30 (2): 357–372.

[KK84] Koch, L., & E. Keyserling. 1884-1890. Die Arachniden Australiens nach der Natur beschrieben und abgebildet vol. 2. Bauer & Raspe: Nürnberg.

[RC73] Rowland, J. M., & J. A. L. Cooke. 1973. Systematics of the arachnid order Uropygida (=Thelyphonida). Journal of Arachnology 1: 55–71.

[S69] Stoliczka, F. 1869. Contribution to the knowledge of Indian Arachnoidea. Journal of the Asiatic Society of Bengal 38 pt 2 (4): 201–251, plates 18–20.

[T89] Thorell, T. 1889. Viaggio di Leonardo Fea in Birmania e regioni vicine. XXI.—Aracnidi Artrogastri Birmani raccolti da L. Fea nel 1885–1887. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a 7: 521–729.

[T94] Thorell, T. 1894. Förteckning öfver Arachnider från Java och närgränsande öar, insamlade af docenten D:r Carl Aurivillius; jemte beskrifingar å några sydasiatiska och sydamerikanska Spindlar. Bihang till K. Svenska Vetenskaps-Akademiens Handlingar 20 pt. 4 (4): 1–63.