Gastrocopta

Gastrocopta procera, from here.

Belongs within: Pupilloidea.

Gastrocopta is a cosmopolitan genus of small snails with the shell aperture partially closed by lamellae. A group of Australian species with particularly dense arrangements of lamellae has been recognised as a distinct genus Pumilicopta but the value of this distinction has subsequently been questioned (Köhler et al. 2012).

Characters (from Stanisic et al. 2010, as Gastrocopta + Pumilicopta): Shell tiny, cylindrically pupiform to conical, mainly dextral, rarely sinistral, smooth with occasional, very fine growth threads, opaque; aperture quadrate to subquadrate with varying numbers and arrangements of teeth and lamellae; lip strongly reflected; umbilicus extremely narrow. Colour silky drab white to brown.

<==Gastrocopta Wollaston 1878 WK13 (see below for synonymy)
    |--+--G. ashmuni WK13
    |  |    |--G. a. ashmuni WK13
    |  |    `--G. a. minor WK13
    |  `--+--G. contracta WK13
    |     `--+--+--G. quadridens WK13
    |        |  `--G. similis WK13
    |        `--+--+--G. tappaniana WK13
    |           |  `--+--G. holzingeri WK13
    |           |     `--G. pentodon WK13
    |           `--+--+--G. armigerella WK13
    |              |  `--G. corticaria WK13
    |              `--+--+--G. bannertonensis (Gabriel 1930) WK13
    |                 |  `--G. margaretae (Cox 1868) WK13 (see below for synonymy)
    |                 `--+--G. mussoni Pilsbry 1917 (see below for synonymy) WK13
    |                    `--+--G. hedleyi Pilsbry 1917 [=Australbinula hedleyi] WK13
    |                       `--G. larapinta (Tate 1896) [=Pupa larapinta, Australbinula larapinta] WK13
    `--+--+--G. pellucida (Pfeiffer 1841) WK13, V07
       |  |    |--G. p. pellucida BP90
       |  |    `--G. p. hordeacella BP90
       |  `--G. servilis (Gould 1843) (see below for synonymy) WK13
       |       |--G. s. servilis V07
       |       `--G. s. riisei (Pfeiffer 1852) V07
       `--+--G. (Immersidens) NC01
          |    |--G. (I.) bilamellata (Sterki & Clapp 1909) NC01
          |    `--G. (I.) dalliana (Sterki 1898) NC01, WK13, NC01
          `--G. (Gastrocopta) NC01
               |  i. s.: *G. (G.) acarus (Benson 1856) [=Pupa acarus] BR17
               |--G. (G.) procera (Gould 1840) WK13, NC01
               |    |--G. p. procera NC01
               |    |--G. p. mcclungi (Hanna & Johnston 1913) NC01
               |    |--G. p. riparia (Pilsbry 1912) NC01
               |    `--G. p. sterkiana (Pilsbry 1912) NC01
               `--+--G. (G.) cristata WK13
                  `--G. (G.) rogersensis Nekola & Coles 2001 WK13, NC01

Gastrocopta incertae sedis:
  G. armifera BP90
  G. bifurcata (Solem 1988) SC07, SS10 [=Pumilicopta bifurcata SS10]
  ‘Pumilicopta’ kamberae (Rensch 1932) S88
  G. kessneri (Solem 1988) KKW12 [=*Pumilicopta kessneri SS10]
  G. klunzingeri V07
    |--G. k. klunzingeri V07
    `--G. k. senegalensis (Maltzan 1890) V07
  G. macdonnelli (Brazier 1875) KKW12 (see below for synonymy)
  G. moellendorffiana Pilsbry 1917 S88
  ‘Pumilicopta’ orientalis (Rensch 1935) S88
  G. pediculus (Shuttleworth 1852) KKW12 (see below for synonymy)
  G. pontifex Rensch 1937 S88
  G. recondita (Tapparone-Canefri 1883) WK13 (see below for synonymy)
  G. seignaciana (Crosse & Fischer 1879) V07
  G. semiclausa Thiele 1928 S88
  G. strangei (Pfeiffer 1852) SS10 (see below for synonymy)
  G. stupefaciens Pokryszko 1996 WK13
  G. (Albinula) E99
    |--G. (A.) acuminata E99
    |    |--G. a. acuminata E99
    |    `--G. a. fossanensis E99
    |--G. (A.) dupuy E99
    |--G. (A.) edlaueri E99
    `--G. (A.) turgida E99
  G. (Sinalbinula Pilsbry 1916) E99, S88
    |--G. (S.) baudoni E99
    |--G. (S.) larteti E99
    |--G. (S.) nouletiana E99
    `--G. (S.) serotina E99
  G. (Vertigopsis) dehmi E99

Gastrocopta Wollaston 1878 WK13 [incl. Australbinula Pilsbry 1916 SS10, Gyrodaria Iredale 1940 SS10, Pumilicopta Solem 1988 KKW12]

Gastrocopta macdonnelli (Brazier 1875) KKW12 [=Pupa (Vertigo) macdonnelli KKW12, Australbinula macdonnelli KKW12, Vertigo macdonelli KKW12; incl. Pupa (Vertigo) macleayi Brazier 1876 KKW12, Bifidaria macleayi KKW12, Gastrocopta macdonnelli macrodon Pilsbry 1917 KKW12, Pupa pedicula Hedley 1891 non P. pediculus Shuttleworth 1852 S88]

Gastrocopta margaretae (Cox 1868) WK13 [=Pupa margaretae WK13, Australbinula margaretae WK13; incl. P. mooreana Tate 1896 non Smith 1894 S88, Gastrocopta pilbarana Solem 1986 WK13, G. tatei Pilsbry 1917 WK13, Australbinula tatei WK13, Pupa wallabyensis Smith 1894 WK13, A. wallabyensis WK13, Gastrocopta wallabyensis WK13]

Gastrocopta mussoni Pilsbry 1917 [=Australbinula mussoni; incl. G. larapinta deserti Pilsbry 1917, A. helmsiana Iredale 1939] WK13

Gastrocopta pediculus (Shuttleworth 1852) KKW12 [=Pupa pediculus KKW12, G. (Sinalbinula) pediculus S88; incl. P. artensis Montrouzier 1859 KKW12, P. moretonensis Cox 1868 KKW12, Gastrocopta pediculus ovatula Moellendorff 1890 KKW12, G. pediculus queenslandica Pilsbry 1917 KKW12, Pupa (Vertigo) rossiteri Brazier 1875 KKW12, G. rossiteri S88, G. simplex Solem 1988 KKW12]

Gastrocopta recondita (Tapparone-Canefri 1883) WK13 [=Pupa recondita S88, Leucochilus reconditus S88; incl. P. (Leucochilus) niobe Fulton 1899 S88, Gastrocopta niobe S88]

Gastrocopta servilis (Gould 1843) [=Pupa servilis; incl. P. lyonsiana Ancey 1892, Gastrocopta lyonsiana, P. microsoma Tapparone-Canefri 1883, G. microsoma] WK13

Gastrocopta strangei (Pfeiffer 1852) SS10 [=*Gyrodaria strangei S88; incl. Australbinula strangeana Iredale 1937 SS10, Gastrocopta strangeana SS10, Gyrodaria strangeana trita Iredale 1940 SS10]

*Type species of generic name indicated

REFERENCES

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BP90] Burch, J. B., & T. A. Pearce. 1990. Terrestrial Gastropoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 201–309. John Wiley & Sones: New York.

[E99] Esu, D. 1999. Contribution to the knowledge of Neogene climate changes in western and central Europe by means of non-marine molluscs. In: J. Agustí, L. Rook & P. Andrews (eds) The Evolution of Neogene Terrestrial Ecosystems in Europe pp. 328–354. Cambridge University Press.

[KKW12] Köhler, F., V. Kessner & C. Whisson. 2012. New records of non-marine, non-camaenid gastropods (Mollusca: Gastropoda) from islands off the Kimberley coast, Western Australia. Records of the Western Australian Museum 27 (1): 21–39.

[NC01] Nekola, J. C., & B. F. Coles. 2001. Systematics and ecology of Gastrocopta (Gastrocopta) rogersensis (Gastropoda: Pupillidae), a new species of land snail from the Midwest of the United States of America. Nautilus 115 (3): 105–114.

[S88] Solem, A. 1988. Non-camaenid land snails of the Kimberley and Northern Territory, Australia. I. Systematics, affinities and ranges. Invertebrate Taxonomy 4: 455–604.

[SC07] Stanisic, J., R. A. D. Cameron, B. M. Pokryszko & J. C. Nekola. 2007. Forest snail faunas from S. E. Queensland and N. E. New South Wales (Australia): patterns of local and regional richness and differentiation. Malacologia 49 (2): 445–462.

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

[V07] Verdcourt, B. 2007. Miscellaneous notes on tropical African non-marine Mollusca. Annales Historico-Naturales Musei Nationalis Hungarici 99: 189–197.

[WK13] Whisson, C. S. & F. Köhler. 2013. Gastrocopta (Mollusca, Gastropoda, Pupillidae) in the Pilbara region of Western Australia. ZooKeys 261: 15–39.

Last updated: 20 January 2020.